280 lines
38 KiB
XML
280 lines
38 KiB
XML
<document id="27F65245B64C3D4A7E04303C17A37012" ID-ISSN="0024-4082" ID-ZooBank="74C07292-2BD6-4E3E-B68D-B144B81BBD83" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629585783401" checkinUser="felipe" docAuthor="Bonifácio, Paulo & Menot, Lénaïck" docDate="2019" docId="0B1B8791FFC40668F8CCED197DDA5855" docLanguage="en" docName="ZoolJLinnSoc.185.555–635.pdf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Polaruschakov omnesae Bonifácio & Menot 2019, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="619" masterDocId="F722FFE9FFFA0628F820E81A7D025E58" masterDocTitle="New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)" masterLastPageNumber="635" masterPageNumber="555" pageNumber="617" updateTime="1699081480826" updateUser="plazi">
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<mods:titleInfo id="0ACA6628846AB4EC94E00DB60687FBE9">
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<mods:title id="1CF1FA48035E98B0C26E9983BDB420AA">New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)</mods:title>
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<mods:name id="ABC8435044DFB570E97D9AA00E721E05" type="personal">
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<mods:roleTerm id="DD2AF19C69D37B9A8FB44B40BB970D87">Author</mods:roleTerm>
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<mods:namePart id="F2276A027773E7E3A47F477228865366">Bonifácio, Paulo</mods:namePart>
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<mods:name id="70156B6B0A991E515A538BA8A5E07C20" type="personal">
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<mods:roleTerm id="38EDF5238D449C6B77B8516BE7626BF7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D1F8A074733A906E377425AE62C8BC9F">Menot, Lénaïck</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FACA9DD62C97323AB4EAC4B760698591">text</mods:typeOfResource>
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<mods:title id="32392DC53A0EEA90E3FE4453E131EAEB">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="DEF669C525A3C96A63A7340038A3237F">
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<mods:date id="AD9F3450E4ED63665864D98FA58366BD">2019</mods:date>
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<mods:detail id="015C7A686257999C22810ABA9E09B20A" type="volume">
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<mods:number id="F4D3F709B036E9CFC7379F5BA4046807">185</mods:number>
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<mods:extent id="165B3622E3D4D0A27F18CA65BE7DBAB3" unit="page">
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<mods:start id="7E47CC2859AE3723AA1AD9A2F4ECF425">555</mods:start>
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<mods:end id="712A712BB2CCC7646DADB95BFF5124EF">635</mods:end>
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<mods:classification id="6F0DCCC43C3DEC46A48A5B439412EB42">journal article</mods:classification>
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<mods:identifier id="E7F0664FBB23F0B7531FEA5959787A52" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="CD0B5B41C8DA2B1086C5878D8618C5B5" type="ZooBank">74C07292-2BD6-4E3E-B68D-B144B81BBD83</mods:identifier>
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<treatment id="0B1B8791FFC40668F8CCED197DDA5855" ID-DOI="http://doi.org/10.5281/zenodo.5240841" ID-Zenodo-Dep="5240841" LSID="urn:lsid:plazi:treatment:0B1B8791FFC40668F8CCED197DDA5855" httpUri="http://treatment.plazi.org/id/0B1B8791FFC40668F8CCED197DDA5855" lastPageId="64" lastPageNumber="619" pageId="62" pageNumber="617">
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<subSubSection id="CBA8650CFFC40616F8CCED197F9F5B43" box="[236,669,1283,1307]" pageId="62" pageNumber="617" type="nomenclature">
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<paragraph id="830D3687FFC40616F8CCED197F9F5B43" blockId="62.[236,669,1283,1307]" box="[236,669,1283,1307]" pageId="62" pageNumber="617">
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<heading id="D84581EBFFC40616F8CCED197F9F5B43" box="[236,669,1283,1307]" centered="true" fontSize="9" level="2" pageId="62" pageNumber="617" reason="2">
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<taxonomicName id="44B24D04FFC40616F8CCED197F315B41" ID-CoL="4KXCK" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[236,563,1283,1306]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="62" pageNumber="628" phylum="Annelida" rank="species" species="omnesae" status="sp. nov.">POLARUSCHAKOV OMNESAE</taxonomicName>
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<emphasis id="B1C6EA95FFC40616FA1AED1C7F9F5B43" bold="true" box="[570,669,1283,1307]" pageId="62" pageNumber="617">
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<taxonomicNameLabel id="AAF557EEFFC40616FA1AED1C7F9F5B43" box="[570,669,1283,1307]" pageId="62" pageNumber="617" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CBA8650CFFC40668F902ED337DDA5855" lastPageId="64" lastPageNumber="619" pageId="62" pageNumber="617" type="description">
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<paragraph id="830D3687FFC40616F902ED337F6A5B1A" blockId="62.[290,616,1321,1346]" box="[290,616,1321,1346]" pageId="62" pageNumber="617">
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(
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<figureCitation id="1B892A02FFC40616F90AED337F165B19" box="[298,532,1321,1346]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="62" pageNumber="617">FIG. 20A–G; TABLES</figureCitation>
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1, 2, 5)
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||
</paragraph>
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||
<paragraph id="830D3687FFC40616F8B1ED687C79590B" blockId="62.[145,761,1394,1875]" pageId="62" pageNumber="617">
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<emphasis id="B1C6EA95FFC40616F8B1ED687C455BDF" box="[145,327,1394,1415]" italics="true" pageId="62" pageNumber="617">Type material:</emphasis>
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||
<materialsCitation id="33DA3CDAFFC40616F97FED687F795806" collectingDate="2015-04-04" collectionCode="IOM" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="11.043533" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-119.65853" municipality="Eastern Pacific Ocean" pageId="62" pageNumber="617" specimenCode="IFR424" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="5C098825FFC40616F97FED687CD35BD0" box="[351,465,1394,1416]" pageId="62" pageNumber="617" type="holotype">Holotype</typeStatus>
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||
, MNHN-IA-TYPE 1841 (
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||
<specimenCode id="D3149EFCFFC40616F8B8ED8A7DF35BFE" box="[152,241,1424,1446]" collectionCode="IFR" pageId="62" pageNumber="617">IFR424</specimenCode>
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||
), complete, length
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||
<quantity id="444A9B62FFC40616F99FED8A7F205BFE" box="[447,546,1424,1446]" metricMagnitude="-3" metricUnit="m" metricValue="4.43" pageId="62" pageNumber="617" unit="mm" value="4.43">4.43 mm</quantity>
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||
, width
|
||
<quantity id="444A9B62FFC40616FA52ED8A7FD75BFE" box="[626,725,1424,1446]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="62" pageNumber="617" unit="mm" value="0.83">0.83 mm</quantity>
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||
, 19 segments (including tentacular segment),
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<location id="866D605CFFC40616FA5CEDB57C995BBB" LSID="urn:lsid:plazi:treatment:0B1B8791FFC40668F8CCED197DDA5855:866D605CFFC40616FA5CEDB57C995BBB" county="Clarion-Clipperton Fracture Zone" latitude="11.043533" longLatPrecision="1" longitude="-119.65853" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="62" pageNumber="617">
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||
Equatorial
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||
<collectingMunicipality id="6369ACFDFFC40616F8B1EDD47C995BBB" box="[145,411,1486,1508]" pageId="62" pageNumber="617">Eastern Pacific Ocean</collectingMunicipality>
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||
</location>
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||
,
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||
<collectingCounty id="6A6C4E0BFFC40616F988EDD47DC9585A" pageId="62" pageNumber="617">Clarion-Clipperton Fracture Zone</collectingCounty>
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||
,
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||
<collectionCode id="E5A3AE42FFC40616F8F8EDF67C0C5859" box="[216,270,1516,1537]" country="Russia" httpUri="http://grbio.org/cool/9zah-8jyz" name="Institute of Oceanology, Academy of Sciences" pageId="62" pageNumber="617">IOM</collectionCode>
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||
license area, station 99, collected
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||
<date id="F70C1047FFC40616FA87EDF67DCC5879" pageId="62" pageNumber="617" value="2015-04-04">
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||
<collectingDate id="E748E9AFFFC40616FA87EDF67DCC5879" pageId="62" pageNumber="617" value="2015-04-04">4 April 2015</collectingDate>
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||
</date>
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||
, epibenthic sledge epi-net, start
|
||
<geoCoordinate id="E6865040FFC40616FA4CEE117FF75878" box="[620,757,1546,1569]" degrees="11" direction="north" minutes="2.296" orientation="latitude" pageId="62" pageNumber="617" precision="1" value="11.038267">11°2.296′N</geoCoordinate>
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||
,
|
||
<geoCoordinate id="E6865040FFC40616F8B1EE337C315867" box="[145,307,1576,1599]" degrees="119" direction="west" minutes="40.825" orientation="longitude" pageId="62" pageNumber="617" precision="1" value="-119.68042">119°40.825′W</geoCoordinate>
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||
, end
|
||
<geoCoordinate id="E6865040FFC40616F953EE307CF45867" box="[371,502,1576,1600]" degrees="11" direction="north" minutes="2.612" orientation="latitude" pageId="62" pageNumber="617" precision="1" value="11.043533">11°2.612′N</geoCoordinate>
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||
,
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||
<geoCoordinate id="E6865040FFC40616FA23EE337FA45867" box="[515,678,1576,1599]" degrees="119" direction="west" minutes="39.512" orientation="longitude" pageId="62" pageNumber="617" precision="1" value="-119.65853">119°39.512′W</geoCoordinate>
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,
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||
<elevation id="089FD1B4FFC40616FA92EE307C9F5806" isEnumeration="true" pageId="62" pageNumber="617">
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<quantity id="444A9B62FFC40616FA92EE307DEB5806" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.401999999999999" metricValueMin="4.398" pageId="62" pageNumber="617" unit="m" value="4400.0" valueMax="4402.0" valueMin="4398.0">4398– 4402 m</quantity>
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||
depth,
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||
<quantity id="444A9B62FFC40616F965EE527C9F5806" box="[325,413,1608,1630]" metricMagnitude="3" metricUnit="m" metricValue="2.529" pageId="62" pageNumber="617" unit="m" value="2529.0">2529 m</quantity>
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||
</elevation>
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||
trawling distance
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="33DA3CDAFFC40616FAA7EE537C75590B" collectingDate="2015-04-07" collectionCode="GSR" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="13.877033" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" pageId="62" pageNumber="617" specimenCode="IFR530-1" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="5C098825FFC40616FAA7EE537FF65806" box="[647,756,1609,1630]" pageId="62" pageNumber="617" type="paratype">Paratype</typeStatus>
|
||
, MNHN-IA-TYPE 1842 (
|
||
<specimenCode id="D3149EFCFFC40616F98EEE7D7F1D5825" box="[430,543,1639,1661]" collectionCode="IFR" pageId="62" pageNumber="617">IFR530-1</specimenCode>
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||
), complete, length
|
||
<quantity id="444A9B62FFC40616F8B1EE9F7DFC58C3" box="[145,254,1669,1691]" metricMagnitude="-3" metricUnit="m" metricValue="4.16" pageId="62" pageNumber="617" unit="mm" value="4.16">4.16 mm</quantity>
|
||
, width
|
||
<quantity id="444A9B62FFC40616F97DEE9C7CC958C3" box="[349,459,1670,1691]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="62" pageNumber="617" unit="mm" value="0.81">0.81 mm</quantity>
|
||
, 20 segments (including tentacular segment),
|
||
<location id="866D605CFFC40616F984EEBE7DD85880" LSID="urn:lsid:plazi:treatment:0B1B8791FFC40668F8CCED197DDA5855:866D605CFFC40616F984EEBE7DD85880" county="Clarion-Clipperton Fracture Zone" latitude="13.877033" longLatPrecision="1" longitude="-123.23772" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="62" pageNumber="617">
|
||
Equatorial
|
||
<collectingMunicipality id="6369ACFDFFC40616FA17EEBF7DD85880" pageId="62" pageNumber="617">Eastern Pacific Ocean</collectingMunicipality>
|
||
</location>
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||
,
|
||
<collectingCounty id="6A6C4E0BFFC40616F8C5EED97F675880" box="[229,613,1731,1753]" pageId="62" pageNumber="617">Clarion-Clipperton Fracture Zone</collectingCounty>
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||
,
|
||
<collectionCode id="E5A3AE42FFC40616FA50EED97FA75880" box="[624,677,1731,1752]" pageId="62" pageNumber="617">GSR</collectionCode>
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||
license area, station 117, collected
|
||
<date id="F70C1047FFC40616F9FAEEF87F7258AF" box="[474,624,1761,1784]" pageId="62" pageNumber="617" value="2015-04-07">
|
||
<collectingDate id="E748E9AFFFC40616F9FAEEF87F7258AF" box="[474,624,1761,1784]" pageId="62" pageNumber="617" value="2015-04-07">7 April 2015</collectingDate>
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||
</date>
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||
, epibenthic sledge epi-net, start
|
||
<geoCoordinate id="E6865040FFC40616F9A5EF1A7F17594E" box="[389,533,1791,1814]" degrees="13" direction="north" minutes="52.317" orientation="latitude" pageId="62" pageNumber="617" precision="1" value="13.87195">13°52.317′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6865040FFC40616FA01EF1A7FC1594E" box="[545,707,1791,1814]" degrees="123" direction="west" minutes="15.442" orientation="longitude" pageId="62" pageNumber="617" precision="1" value="-123.25737">123°15.442′W</geoCoordinate>
|
||
, end
|
||
<geoCoordinate id="E6865040FFC40616F8B1EF057C1C596C" box="[145,286,1822,1845]" degrees="13" direction="north" minutes="52.622" orientation="latitude" pageId="62" pageNumber="617" precision="1" value="13.877033">13°52.622′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6865040FFC40616F909EF057CC5596C" box="[297,455,1822,1845]" degrees="123" direction="west" minutes="14.263" orientation="longitude" pageId="62" pageNumber="617" precision="1" value="-123.23772">123°14.263′W</geoCoordinate>
|
||
,
|
||
<elevation id="089FD1B4FFC40616F9F1EF057DA5590B" isEnumeration="true" pageId="62" pageNumber="617">
|
||
<quantity id="444A9B62FFC40616F9F1EF057F69596C" box="[465,619,1823,1845]" metricMagnitude="3" metricUnit="m" metricValue="4.5095" metricValueMax="4.521" metricValueMin="4.498" pageId="62" pageNumber="617" unit="m" value="4509.5" valueMax="4521.0" valueMin="4498.0">4498–4521 m</quantity>
|
||
depth,
|
||
<quantity id="444A9B62FFC40616FAE2EF057DA5590B" metricMagnitude="3" metricUnit="m" metricValue="3.129" pageId="62" pageNumber="617" unit="m" value="3129.0">3129 m</quantity>
|
||
</elevation>
|
||
trawling distance
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFC40616FB09ED1E79965BEB" blockId="62.[809,1426,1284,1888]" pageId="62" pageNumber="617">
|
||
<emphasis id="B1C6EA95FFC40616FB09ED1E78905B41" box="[809,1426,1284,1305]" italics="true" pageId="62" pageNumber="617">
|
||
Description (based on
|
||
<typeStatus id="5C098825FFC40616FC77ED1E79C05B41" box="[1111,1218,1284,1305]" pageId="62" pageNumber="617" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="5C098825FFC40616FD2EED1E78805B41" box="[1294,1410,1284,1305]" pageId="62" pageNumber="617" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
<typeStatus id="5C098825FFC40616FB09ED387E905B60" box="[809,914,1314,1336]" pageId="62" pageNumber="617" type="holotype">Holotype</typeStatus>
|
||
complete,
|
||
<quantity id="444A9B62FFC40616FC30ED3879745B60" box="[1040,1142,1314,1336]" metricMagnitude="-3" metricUnit="m" metricValue="4.43" pageId="62" pageNumber="617" unit="mm" value="4.43">4.43 mm</quantity>
|
||
long and
|
||
<quantity id="444A9B62FFC40616FCCCED3878505B60" box="[1260,1362,1314,1336]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="62" pageNumber="617" unit="mm" value="0.83">0.83 mm</quantity>
|
||
wide for 19 segments (including tentacular segment), dorsoventrally flattened, posteriorly tapering; colour of live animal not known; ethanol-preserved specimen pale white, slightly translucent.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFC40616FB61EDA67EC4589F" blockId="62.[809,1426,1284,1888]" pageId="62" pageNumber="617">
|
||
Prostomium bilobed, wider than long, anterior lobes not developed, conical; frontal filaments absent; median notch between prostomial lobes moderately narrow and moderately deep (
|
||
<figureCitation id="1B892A02FFC40616FD34EE02787D5875" box="[1300,1407,1560,1582]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="62" pageNumber="617">Fig. 20A</figureCitation>
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||
); eyes absent. Median and lateral antennae absent. Palps smooth, tapering into thin tips, short (reaching segment 3); palpophores not enlarged (
|
||
<figureCitation id="1B892A02FFC40616FD00EE6E788058D1" box="[1312,1410,1652,1674]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="62" pageNumber="617">Fig. 20A</figureCitation>
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||
). Facial tubercle absent. Upper lip with multiple minute folds.
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||
</paragraph>
|
||
<paragraph id="830D3687FFC40617FB61EECA7C365817" blockId="62.[809,1426,1284,1888]" lastBlockId="63.[163,779,1287,1891]" lastPageId="63" lastPageNumber="618" pageId="62" pageNumber="617">
|
||
Tentacular segment fused to prostomium, with a pair of short lobes, inserted laterally and slightly below prostomium; without acicula or chaetae; tentaculophores distinct, bulbous, equal sized; styles smooth, tapering into thin tips, short (reaching segment 3), dorsal tentacular style slightly shorter than ventral tentacular style (
|
||
<figureCitation id="1B892A02FFC50617FA29ED3C7F685B63" box="[521,618,1318,1340]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20A</figureCitation>
|
||
); ventrally to the tentaculophores is a distinct globular pad, located laterally to the mouth. Pharynx not everted in
|
||
<typeStatus id="5C098825FFC50617FAF1ED797DDA5BCF" pageId="63" pageNumber="618" type="holotype">holotype</typeStatus>
|
||
; dissected in
|
||
<typeStatus id="5C098825FFC50617F958ED987CE35BCF" box="[376,481,1410,1431]" pageId="63" pageNumber="618" type="paratype">paratype</typeStatus>
|
||
(MNHN-IA-TYPE 1842), with seven pairs of distal papillae, subtriangular, equal sized; two pairs of jaws, each one with one main fang, outer margin with a very small, secondary tooth (small elevation;
|
||
<figureCitation id="1B892A02FFC50617F95DEDE67CE45849" box="[381,486,1532,1554]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20B</figureCitation>
|
||
). Second segment with elytrophores, subbiramous parapodia, chaetae and ventral cirri.
|
||
</paragraph>
|
||
<caption id="D7CD660FFFC50617F883EC487E8A5A85" pageId="63" pageNumber="618" startId="63.[163,242,1106,1128]" targetBox="[163,1443,200,1067]" targetPageId="63">
|
||
<paragraph id="830D3687FFC50617F883EC487E8A5A85" blockId="63.[163,1443,1106,1245]" pageId="63" pageNumber="618">
|
||
<emphasis id="B1C6EA95FFC50617F883EC487F985A3F" bold="true" box="[163,666,1106,1128]" pageId="63" pageNumber="618">
|
||
Figure 20.
|
||
<taxonomicName id="44B24D04FFC50617F904EC487F3E5A3F" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[292,572,1106,1128]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="63" pageNumber="618" phylum="Annelida" rank="species" species="omnesae" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFC50617F904EC487F3E5A3F" bold="true" box="[292,572,1106,1128]" italics="true" pageId="63" pageNumber="618">Polaruschakov omnesae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFC50617FA63EC487F985A3F" box="[579,666,1106,1127]" pageId="63" pageNumber="618" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="830D3687FFC50617F89BEE427C0958F3" blockId="63.[163,779,1287,1891]" pageId="63" pageNumber="618">Nine pairs of distinct, knob-like elytrophores present on segments 2, 4, 5, 7, 9, 11, 13, 15 and 17 (all elytra missing).</paragraph>
|
||
<paragraph id="830D3687FFC50617F89BEEAE7DF4591D" blockId="63.[163,779,1287,1891]" pageId="63" pageNumber="618">
|
||
Cirrigerous segments with distinct, small dorsal cirrophores (
|
||
<figureCitation id="1B892A02FFC50617F935EEC97C7758B0" box="[277,373,1747,1769]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20C</figureCitation>
|
||
), inserted subdistally on notopodia; styles smooth, tapering into thin tips, long (slightly longer than tip of neuroacicular lobe); dorsal tubercles absent.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFC50617F89BEF547F32593B" blockId="63.[163,779,1287,1891]" box="[187,560,1869,1891]" pageId="63" pageNumber="618">Segment 6 without modification.</paragraph>
|
||
<paragraph id="830D3687FFC50617FB73ED1D79FE5BEE" blockId="63.[827,1449,1287,1891]" pageId="63" pageNumber="618">
|
||
Ventral cirri smooth, tapering into thin tips, present from segment 2 to last segment; inserted basally on neuropodia of segment 2, style short (shorter than tip of neuroacicular lobe); in subsequent segments inserted medially on neuropodia (
|
||
<figureCitation id="1B892A02FFC50617FC98ED9B78145BCF" box="[1208,1302,1409,1431]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20C</figureCitation>
|
||
), style short (shorter than tip of neuroacicular lobe).
|
||
</paragraph>
|
||
<paragraph id="830D3687FFC50617FB73EDA57E97593B" blockId="63.[827,1449,1287,1891]" pageId="63" pageNumber="618">
|
||
Parapodia subbiramous; notopodia much shorter than neuropodia (
|
||
<figureCitation id="1B892A02FFC50617FC09EDC779945BAB" box="[1065,1174,1501,1523]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20C</figureCitation>
|
||
). Notopodia narrow, subtriangular, tapering into very short acicular lobe, tip of notoacicula not penetrating epidermis. Neuropodia large, subtriangular, tapering into long acicular lobe, tip of neuroacicula not penetrating epidermis. Notochaetae very few (one to three observed), short, slender, slightly curved, with distinct spinous rows on convex side, with blunt tips (
|
||
<figureCitation id="1B892A02FFC50617FCB7EEAE79FE5891" box="[1175,1276,1716,1738]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20D</figureCitation>
|
||
);notochaetae more slender than neurochaetae. Neurochaetae moderate in number (ten to 15 observed), long, distally flattened to concave, serrated along both margins, with pointed tips (
|
||
<figureCitation id="1B892A02FFC50617FBF2EF35794C591C" box="[978,1102,1839,1861]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="63" pageNumber="618">Fig. 20E, F</figureCitation>
|
||
); slightly stouter in middle of fascicle.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F889E8DF7C6C5F6F" blockId="64.[145,761,197,312]" pageId="64" pageNumber="619">
|
||
Nephridial papillae absent. Pygidium rounded, not enclosed by last segment; with terminal anus; with ventral papilla, rounded to ovoid (
|
||
<figureCitation id="1B892A02FFBA0668FA35E9197F745F40" box="[533,630,259,281]" captionStart="Figure 20" captionStartId="63.[163,242,1106,1128]" captionTargetBox="[163,1443,200,1067]" captionTargetId="graphics-1266@63.[798,1024,480,711]" captionTargetPageId="63" captionText="Figure 20. Polaruschakov omnesae sp. nov., holotype MNHN-IA-TYPE 1841 (A, C–G) and paratype MNHN-IA-TYPE 1842 (B). A, anterior end, dorsolateral view, with chaetae omitted. B, inner view of half side of dissected pharynx with few papillae. C, right parapodia, anterior view, segment 10. D, notochaeta with faint spinous rows, segment 10. E, upper neurochaeta, frontolateral view, segment 3. F, middle neurochaeta, frontal view, segment 10. G, last two segments with pygidial papilla, ventrolateral view. Abbreviations: mf, minute folds; pa, papilla." pageId="64" pageNumber="619">Fig. 20G</figureCitation>
|
||
). Anal cirri lost, scars not seen.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F8B1E97A7DE55C16" blockId="64.[145,761,352,590]" pageId="64" pageNumber="619">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1E97A7CC05F2E" box="[145,450,352,374]" italics="true" pageId="64" pageNumber="619">Morphological variation:</emphasis>
|
||
Specimens with 19 and 20 segments were found, which share most of the morphological characters given in the species description. However, the
|
||
<typeStatus id="5C098825FFBA0668F9ECE9A77F305F8B" box="[460,562,445,467]" pageId="64" pageNumber="619" type="holotype">holotype</typeStatus>
|
||
shows a minute prostomial peak and a minute ventral papilla on the pygidium, whereas the
|
||
<typeStatus id="5C098825FFBA0668F981E9E17F0B5C48" box="[417,521,507,528]" pageId="64" pageNumber="619" type="paratype">paratype</typeStatus>
|
||
does not show those peaks but presents a more rounded, small, ventral papilla.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F8B1EA6D7F255A45" blockId="64.[145,760,631,1053]" pageId="64" pageNumber="619">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1EA6D7F325CD5" box="[145,560,631,653]" italics="true" pageId="64" pageNumber="619">
|
||
Remarks:
|
||
<taxonomicName id="44B24D04FFBA0668F930EA6D7F325CD5" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[272,560,631,653]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="omnesae" status="sp. nov.">Polaruschakov omnesae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AAF557EEFFBA0668FA1AEA6D7F965CD4" box="[570,660,631,652]" pageId="64" pageNumber="619" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is more similar to
|
||
<taxonomicName id="44B24D04FFBA0668F930EA8C7F185CF3" baseAuthorityName="Uschakov" baseAuthorityYear="1957" box="[272,538,662,683]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="polaris">
|
||
<emphasis id="B1C6EA95FFBA0668F930EA8C7F185CF3" box="[272,538,662,683]" italics="true" pageId="64" pageNumber="619">Polaruschakov polaris</emphasis>
|
||
</taxonomicName>
|
||
, with both having notochaetae with blunt tips, a wide notch and fewer chaetae than
|
||
<taxonomicName id="44B24D04FFBA0668F963EAC97F4C5CB1" authorityName="Pettibone" authorityYear="1976" box="[323,590,723,745]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="reyssi">
|
||
<emphasis id="B1C6EA95FFBA0668F963EAC97F4C5CB1" box="[323,590,723,745]" italics="true" pageId="64" pageNumber="619">Polaruschakov reyssi</emphasis>
|
||
</taxonomicName>
|
||
. However, in
|
||
<taxonomicName id="44B24D04FFBA0668F8B1EAE87CB15D50" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[145,435,754,776]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="omnesae" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1EAE87CB15D50" box="[145,435,754,776]" italics="true" pageId="64" pageNumber="619">Polaruschakov omnesae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFBA0668F99EEAE97F1B5D50" box="[446,537,755,776]" pageId="64" pageNumber="619" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the neurochaetae tips are pointed and the palps are short (reaching segment 3), whereas in
|
||
<taxonomicName id="44B24D04FFBA0668F999EB2A7FC55D1D" baseAuthorityName="Uschakov" baseAuthorityYear="1957" box="[441,711,816,837]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="polaris">
|
||
<emphasis id="B1C6EA95FFBA0668F999EB2A7FC55D1D" box="[441,711,816,837]" italics="true" pageId="64" pageNumber="619">Polaruschakov polaris</emphasis>
|
||
</taxonomicName>
|
||
the neurochaetae tips are rounded and the palps are longer (
|
||
<tableCitation id="CE30033CFFBA0668F8CFEB777C4A5DDB" box="[239,328,877,899]" captionStart="Table 5" captionStartId="68.[145,209,196,217]" captionText="Table 5. Diagnostic characters for all valid species in the genus Polaruschakov" pageId="64" pageNumber="619">Table 5</tableCitation>
|
||
). The average K2P distance among
|
||
<taxonomicName id="44B24D04FFBA0668F8B1EB967CAE5DF9" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" box="[145,428,908,929]" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="lamellae" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1EB967CAE5DF9" box="[145,428,908,929]" italics="true" pageId="64" pageNumber="619">Polaruschakov lamellae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFBA0668F996EB977F0F5DFA" box="[438,525,909,930]" pageId="64" pageNumber="619" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="44B24D04FFBA0668FA6BEB967DF35D99" authority="Bonifácio & Menot, 2019" authorityName="Bonifácio & Menot" authorityYear="2019" class="Polychaeta" family="Polynoidae" genus="Polaruschakov" kingdom="Animalia" order="Phyllodocida" pageId="64" pageNumber="619" phylum="Annelida" rank="species" species="omnesae" status="sp. nov.">
|
||
<emphasis id="B1C6EA95FFBA0668FA6BEB967DF35D99" italics="true" pageId="64" pageNumber="619">Polaruschakov omnesae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAF557EEFFBA0668F8D6EBB17C455D98" box="[246,327,939,960]" pageId="64" pageNumber="619" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was high (23.3% for
|
||
<emphasis id="B1C6EA95FFBA0668FA0BEBB17F585D98" box="[555,602,939,960]" italics="true" pageId="64" pageNumber="619">COI</emphasis>
|
||
and 24.4% for 16S). The presence of minute prostomial peaks could be an artefact of preservation, because its presence cannot be observed in the
|
||
<typeStatus id="5C098825FFBA0668F99AEC127F215A45" box="[442,547,1032,1053]" pageId="64" pageNumber="619" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F8B1EC5C7FD85A23" blockId="64.[145,760,1094,1147]" pageId="64" pageNumber="619">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1EC5C7C135A03" box="[145,273,1094,1115]" italics="true" pageId="64" pageNumber="619">Etymology:</emphasis>
|
||
This species is dedicated to Emmanuelle Omnes (Ifremer) for her help with laboratory work.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F8B1ECBE7CC55B0C" blockId="64.[145,761,1188,1364]" pageId="64" pageNumber="619">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1ECBE7C325AE1" box="[145,304,1188,1209]" italics="true" pageId="64" pageNumber="619">Genetic data:</emphasis>
|
||
DNA sequencing for this species was successful for
|
||
<emphasis id="B1C6EA95FFBA0668F965ECD97C745A80" box="[325,374,1219,1240]" italics="true" pageId="64" pageNumber="619">COI</emphasis>
|
||
, 16S and 18S. Both specimens shared 100% of genetic material in
|
||
<emphasis id="B1C6EA95FFBA0668FA0CECF87F595AAF" box="[556,603,1250,1271]" italics="true" pageId="64" pageNumber="619">COI</emphasis>
|
||
and 16S. 18S was not successfully sequenced for the
|
||
<typeStatus id="5C098825FFBA0668FA78ED1B7FC05B4E" box="[600,706,1281,1302]" pageId="64" pageNumber="619" type="paratype">paratype</typeStatus>
|
||
. The average K2P distance for intraspecific variation was 0.0% for both
|
||
<emphasis id="B1C6EA95FFBA0668F90EED247C5F5B0B" box="[302,349,1342,1363]" italics="true" pageId="64" pageNumber="619">COI</emphasis>
|
||
and 16S.
|
||
</paragraph>
|
||
<paragraph id="830D3687FFBA0668F8B1ED677DDA5855" blockId="64.[145,761,1405,1549]" pageId="64" pageNumber="619">
|
||
<emphasis id="B1C6EA95FFBA0668F8B1ED677C2B5BCA" box="[145,297,1405,1426]" italics="true" pageId="64" pageNumber="619">Distribution:</emphasis>
|
||
Based on the material examined (
|
||
<specimenCount id="95B4FD0EFFBA0668FAEEED677C0F5BE9" pageId="64" pageNumber="619" type="generic">two specimens</specimenCount>
|
||
), this species has a restricted distribution within the Clarion-Clipperton Fracture Zone, being sampled in IOM (
|
||
<typeStatus id="5C098825FFBA0668F953EDC37CA85BB6" box="[371,426,1497,1518]" pageId="64" pageNumber="619">type</typeStatus>
|
||
locality) and GSR license areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |