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<document id="452BBD85A7584DC34CE4D7EA67E3B41B" ID-DOI="10.5252/z2013n2a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="5160321" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628102401095" checkinUser="felipe" docAuthor="Bamber, Roger N." docDate="2013" docId="03FE523E833CDA52FCD3FC9A84EB164C" docLanguage="en" docName="Zoosystema.35.2.195-214.pdf" docOrigin="Zoosystema 35 (2)" docSource="http://dx.doi.org/10.5252/z2013n2a5" docStyle="DocumentStyle:BCA1AC1CA2CB189AA8340D8E5C61F956.3:Zoosystema.2013-2014.journal_article" docStyleId="BCA1AC1CA2CB189AA8340D8E5C61F956" docStyleName="Zoosystema.2013-2014.journal_article" docStyleVersion="3" docTitle="Pycnogonum staplesi Bamber 2013, n. sp." docType="treatment" docVersion="2" lastPageNumber="212" masterDocId="FFC72A468333DA43FFCCFFA880531443" masterDocTitle="Deep-water Pycnogonida from recent cruises to Papua New Guinea and Melanesia, with an appendix of new records from Polynesia and descriptions of five new species" masterLastPageNumber="214" masterPageNumber="195" pageNumber="210" updateTime="1699034498488" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="AB6E5B8E4CD78FAE65D6394E435D5425" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="64908A2B78656A96895FE663FD4543BC">
<mods:title id="4BBDA43166DBA17CFE52F44C9C650BBA">Deep-water Pycnogonida from recent cruises to Papua New Guinea and Melanesia, with an appendix of new records from Polynesia and descriptions of five new species</mods:title>
</mods:titleInfo>
<mods:name id="8EB772E6364A6DC23A8511CD08834E81" type="personal">
<mods:role id="59226984F1536EDFAB9B5617C8CD92DA">
<mods:roleTerm id="4AF6FF2C719A13BC247DE3C7FCAA987C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A1A7778914AFCD3365232992EBB547C">Bamber, Roger N.</mods:namePart>
<mods:affiliation id="768570434D2CE66F26C4FC4C1EFE8813">ARTOO Marine Biology Consultants Ocean Quay Marina, Belvidere Road Southampton SO 14 5 QY (United Kingdom)</mods:affiliation>
</mods:name>
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<mods:title id="79352CB2C6D27F6720A766A198C2F589">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="9A67098AE979821D80B7CEF406D89E62">
<mods:date id="6C8BBE094395958CF12755013EFF6A2B">2013</mods:date>
<mods:detail id="3978B11894D3425417755915D9269DDA" type="pubDate">
<mods:number id="6262275C30838286866F7CD57E84B4BC">2013-06-28</mods:number>
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<mods:detail id="068BF3911F35D73EFAF669434E2E0787" type="volume">
<mods:number id="D41284E6BB13DD5393FC0A1DA363FFFA">35</mods:number>
</mods:detail>
<mods:detail id="79E4FE93FE1C3313A3651BF38C307A3A" type="issue">
<mods:number id="492C6A3BC298F17700BA749D9C72295A">2</mods:number>
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<mods:start id="BC101BD922B5255598F204A64F8CE879">195</mods:start>
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<mods:url id="2E42E07B92A1767A0DBB4413228E4B24">http://dx.doi.org/10.5252/z2013n2a5</mods:url>
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<mods:classification id="91E0F0F7967DBD99E19766AE00425B13">journal article</mods:classification>
<mods:identifier id="CCC32BF817DBB1233677B6D3C4650B2F" type="DOI">10.5252/z2013n2a5</mods:identifier>
<mods:identifier id="ABE9F63A3F3DC904B72F4C727764D1E5" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="E4E4CAE5A1EA674D0597FC0C4DB9A6BB" type="Zenodo-Dep">5160321</mods:identifier>
</mods:mods>
<treatment id="03FE523E833CDA52FCD3FC9A84EB164C" LSID="urn:lsid:plazi:treatment:03FE523E833CDA52FCD3FC9A84EB164C" httpUri="http://treatment.plazi.org/id/03FE523E833CDA52FCD3FC9A84EB164C" lastPageId="17" lastPageNumber="212" pageId="15" pageNumber="210">
<subSubSection id="C34DB0A3833CDA4CFCD3FC9A846C170F" box="[799,1087,817,844]" pageId="15" pageNumber="210" type="nomenclature">
<paragraph id="8BE8E328833CDA4CFCD3FC9A846C170F" blockId="15.[799,1087,817,844]" box="[799,1087,817,844]" pageId="15" pageNumber="210">
<heading id="D0A05444833CDA4CFCD3FC9A846C170F" box="[799,1087,817,844]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="210" reason="2">
<taxonomicName id="4C5798AB833CDA4CFCD3FC9A83AE170F" ID-CoL="4QMHF" authority="Bamber, 2013" authorityName="Bamber" authorityYear="2013" box="[799,1021,817,844]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="staplesi" status="sp. nov.">
<emphasis id="B9233F3A833CDA4CFCD3FC9A83AE170F" bold="true" box="[799,1021,817,844]" italics="true" pageId="15" pageNumber="210">Pycnogonum staplesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2108241833CDA4CFBC8FC9A846C170F" box="[1028,1087,818,844]" pageId="15" pageNumber="210" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34DB0A3833CDA4CFD69FCC383CF17DE" pageId="15" pageNumber="210" type="reference_group">
<paragraph id="8BE8E328833CDA4CFD69FCC383CF17DE" blockId="15.[677,1209,875,925]" pageId="15" pageNumber="210">
<treatmentCitationGroup id="AB47C406833CDA4CFD69FCC383CF17DE" pageId="15" pageNumber="210">
<treatmentCitation id="0AF6C539833CDA4CFD69FCC3845117C0" author="STOCK J. H." box="[677,1026,875,899]" page="407" pageId="15" pageNumber="210" year="1997">
<taxonomicName id="4C5798AB833CDA4CFD69FCC3845117C0" ID-CoL="4QMGR" authority="Stock, 1997: 407" authorityName="Stock" authorityPageNumber="407" authorityYear="1997" box="[677,1026,875,899]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A833CDA4CFD69FCC3831817C1" box="[677,843,875,898]" italics="true" pageId="15" pageNumber="210">Pycnogonum occa</emphasis>
<bibRefCitation id="EFC69ED9833CDA4CFC9EFCC3845117C0" author="STOCK J. H." box="[850,1026,875,899]" pageId="15" pageNumber="210" pagination="389 - 409" refId="ref11534" refString="STOCK J. H. 1997. - Pycnogonids collected in recent years around New Caledonia and Vanuatu. In: CROSNIER A. (ed.), Resultats des Campagnes MUSORSTOM, Vol. 18 Memoires du Museum national d'Histoire naturelle, Paris 176: 389 - 409." type="journal article" year="1997">Stock, 1997: 407</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Non
<taxonomicName id="4C5798AB833CDA4CFB8EFCC3830917DE" ID-CoL="4QMGR" authority="Loman, 1908" authorityName="Loman" authorityYear="1908" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A833CDA4CFB8EFCC3829F17DE" italics="true" pageId="15" pageNumber="210">Pycnogonum occa</emphasis>
<bibRefCitation id="EFC69ED9833CDA4CFD1EFC2E830917DE" author="LOMAN J. C. C." box="[722,858,901,925]" pageId="15" pageNumber="210" pagination="1 - 88" refId="ref11253" refString="LOMAN J. C. C. 1908. - Die Pantopoden der Siboga-Expedition. Siboga Expeditie Monographie 40: 1 - 88; pls I - XV." type="journal article" year="1908">Loman, 1908</bibRefCitation>
</taxonomicName>
: fig. 6.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34DB0A3833CDA4CFD69FC1D83B41041" pageId="15" pageNumber="210" type="materials_examined">
<paragraph id="8BE8E328833CDA4CFD69FC1D83B41041" blockId="15.[674,1210,949,1026]" pageId="15" pageNumber="210">
MATERIAL EXAMINED. — 1 “
<specimenCount id="9D5128A1833CDA4CFC68FC1083E6178F" box="[932,949,952,972]" pageId="15" pageNumber="210" type="female">♀</specimenCount>
”,
<typeStatus id="54EC5D8A833CDA4CFC04FC1D844A178E" box="[968,1049,949,973]" pageId="15" pageNumber="210" type="holotype">holotype</typeStatus>
MNHN-Py 922,
<collectingCountry id="F340A3B8833CDA4CFD6EFC78836917A4" box="[674,826,975,999]" name="New Caledonia" pageId="15" pageNumber="210">New Caledonia</collectingCountry>
, Stn DW923,
<geoCoordinate id="EE6385EF833CDA4CFC0EFC67847C17A4" box="[962,1071,975,999]" degrees="18" direction="south" minutes="51.51" orientation="latitude" pageId="15" pageNumber="210" precision="9" value="-18.8585">18°51.51’S</geoCoordinate>
,
<geoCoordinate id="EE6385EF833CDA4CFBF4FC6784E717A4" box="[1080,1204,975,999]" degrees="163" direction="east" minutes="24.17" orientation="longitude" pageId="15" pageNumber="210" precision="9" value="163.40283">163°24.17’E</geoCoordinate>
, 502-
<quantity id="4CAF4ECD833CDA4CFD1CFC4283421042" box="[720,785,1002,1026]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="15" pageNumber="210" unit="m" value="470.0">470 m</quantity>
depth,
<date id="FFE9C5E8833CDA4CFC91FC4283B01041" box="[861,995,1002,1026]" pageId="15" pageNumber="210" value="1994-08-06">06.VIII.1994</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DB0A3833CDA4CFD69FBB2839F1025" pageId="15" pageNumber="210" type="etymology">
<paragraph id="8BE8E328833CDA4CFD69FBB2839F1025" blockId="15.[677,1210,1049,1126]" pageId="15" pageNumber="210">
ETYMOLOGY. — Named after David Staples, in recognition of his comprehensive work on the morphology of
<taxonomicName id="4C5798AB833CDA4CFD69FBE783351025" box="[677,870,1102,1126]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="15" pageNumber="210" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9233F3A833CDA4CFD69FBE783481025" box="[677,795,1103,1126]" italics="true" pageId="15" pageNumber="210">Pycnogonum</emphasis>
species
</taxonomicName>
,
<emphasis id="B9233F3A833CDA4CFCBCFBE783951025" box="[880,966,1102,1126]" italics="true" pageId="15" pageNumber="210">inter alia</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DB0A3833CDA52FD69FB27813B160D" lastPageId="17" lastPageNumber="212" pageId="15" pageNumber="210" type="description">
<paragraph id="8BE8E328833CDA4CFD69FB27839C10E4" blockId="15.[674,1212,1167,1700]" box="[677,975,1167,1193]" pageId="15" pageNumber="210">
<materialsCitation id="3B3FE975833CDA4CFD69FB27839C10E4" box="[677,975,1167,1193]" collectionCode="DESCRIPTION, OF" pageId="15" pageNumber="210" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED467BED833CDA4CFD69FB27830A10E4" box="[677,857,1167,1193]" pageId="15" pageNumber="210">DESCRIPTION OF</collectionCode>
<typeStatus id="54EC5D8A833CDA4CFCACFB3C839C10E4" box="[864,975,1172,1191]" pageId="15" pageNumber="210" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</paragraph>
<paragraph id="8BE8E328833CDA53FD69FB0682D51246" blockId="15.[674,1212,1167,1700]" lastBlockId="16.[111,649,1420,1700]" lastPageId="16" lastPageNumber="211" pageId="15" pageNumber="210">
Trunk length
<quantity id="4CAF4ECD833CDA4CFCFAFB0683DD108B" box="[822,910,1198,1224]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="15" pageNumber="210" unit="mm" value="2.9">2.9 mm</quantity>
, width
<quantity id="4CAF4ECD833CDA4CFC10FB068460108B" box="[988,1075,1198,1224]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="210" unit="mm" value="2.1">2.1 mm</quantity>
, leg span approximately
<quantity id="4CAF4ECD833CDA4CFCEAFB66832510AB" box="[806,886,1230,1256]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="210" unit="mm" value="12.0">12 mm</quantity>
. Cuticle sculptured throughout with small warts.Trunk (
<figureCitation id="136CFFAD833CDA4CFC51FB458457114B" box="[925,1028,1261,1288]" captionStart="FIG" captionStartId="16.[113,124,1317,1335]" captionTargetBox="[113,1209,215,1275]" captionTargetId="figure-220@16.[113,1209,215,1275]" captionTargetPageId="16" captionText="FIG. 6. – Pycnogonum staplesi n. sp., holotype (Py 922): A, habitus, lateral (note, mid-dorsal tubercle on trunk segment 2 broken); B, habitus, dorsal; C, distal articles of third leg, with detail of split spinule. Scale bar: A, B, 3 mm; C, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5160352" httpUri="https://zenodo.org/record/5160352/files/figure.png" pageId="15" pageNumber="210">Fig.6A, B</figureCitation>
) fully segmented, glabrous; lateral processes about 85% as long as segment width, separated by about half of their own width, each bearing fleshy dorsodistal tubercle about 1.5 times as tall as lateral-process width. Cephalon 32% of trunk length, ocular tubercle tall, distally rounded, 1.7 times as high as wide, without eyes; mid-dorsal tubercle near posterior margin similar to lateral process tubercles, tapering, just taller than ocular tubercle; second and third trunk segments each two thirds as long as cephalon and each (?) with a slender, fleshy mid-dorsal tapering tubercle at posterior margin, tubercle on second segment broken, tubercle on third segment as tall mid-dorsal cephalon tubercle; fourth trunk segment slightly shorter than third, without mid-dorsal tubercle. Abdomen naked, not articulating, held horizontally, 0.27 times as long as total trunk length, distally blunt.
</paragraph>
<caption id="DF28B3A08323DA53FFBDFA8D83AB110E" ID-DOI="http://doi.org/10.5281/zenodo.5160352" ID-Zenodo-Dep="5160352" httpUri="https://zenodo.org/record/5160352/files/figure.png" pageId="16" pageNumber="211" startId="16.[113,124,1317,1335]" targetBox="[113,1209,215,1275]" targetPageId="16">
<paragraph id="8BE8E3288323DA53FFBDFA8D83AB110E" blockId="16.[113,1211,1317,1357]" pageId="16" pageNumber="211">
FIG. 6. –
<taxonomicName id="4C5798AB8323DA53FF70FA8D813C1174" authority="Bamber, 2013" authorityName="Bamber" authorityYear="2013" box="[188,367,1317,1335]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="16" pageNumber="211" phylum="Arthropoda" rank="species" species="staplesi" status="sp. nov.">
<emphasis id="B9233F3A8323DA53FF70FA8D813C1174" box="[188,367,1317,1335]" italics="true" pageId="16" pageNumber="211">Pycnogonum staplesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21082418323DA53FEBAFA8D81FA1174" box="[374,425,1317,1335]" pageId="16" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
, holotype (Py 922):
<emphasis id="B9233F3A8323DA53FD9CFA8D820E1174" bold="true" box="[592,605,1317,1335]" pageId="16" pageNumber="211">A</emphasis>
, habitus, lateral (note, mid-dorsal tubercle on trunk segment 2 broken);
<emphasis id="B9233F3A8323DA53FFBDFA93802D110E" bold="true" box="[113,126,1339,1357]" pageId="16" pageNumber="211">B</emphasis>
, habitus, dorsal;
<emphasis id="B9233F3A8323DA53FEC2FA93814F110E" bold="true" box="[270,284,1339,1357]" pageId="16" pageNumber="211">C</emphasis>
, distal articles of third leg, with detail of split spinule. Scale bar: A, B, 3 mm; C, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BE8E3288323DA53FF4AF9A281481227" blockId="16.[111,649,1420,1700]" pageId="16" pageNumber="211">
Proboscis naked, tapering, half as long as cephalon, without tubercles; oral glands (
<emphasis id="B9233F3A8323DA53FE32F98382601207" box="[510,563,1579,1604]" italics="true" pageId="16" pageNumber="211">sensu</emphasis>
<bibRefCitation id="EFC69ED98323DA53FDF7F98280F91227" author="STAPLES D." pageId="16" pageNumber="211" pagination="541 - 553" refId="ref11325" refString="STAPLES D. 2002. - Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria 59: 541 - 553." type="journal article" year="2002">Staples 2002</bibRefCitation>
) not seen.
</paragraph>
<paragraph id="8BE8E3288323DA52FF44F9C1826915D2" blockId="16.[111,649,1420,1700]" lastBlockId="17.[112,648,216,590]" lastPageId="17" lastPageNumber="212" pageId="16" pageNumber="211">
Third leg with sparse dorsal and ventral slender simple spinules on margins of all coxae, femur and tibia 1, dorsally on tibia 2 and propodus (e.g.,
<figureCitation id="136CFFAD8323DA53FD6FFA0382A81185" box="[675,763,1451,1478]" captionStart="FIG" captionStartId="16.[113,124,1317,1335]" captionTargetBox="[113,1209,215,1275]" captionTargetId="figure-220@16.[113,1209,215,1275]" captionTargetPageId="16" captionText="FIG. 6. – Pycnogonum staplesi n. sp., holotype (Py 922): A, habitus, lateral (note, mid-dorsal tubercle on trunk segment 2 broken); B, habitus, dorsal; C, distal articles of third leg, with detail of split spinule. Scale bar: A, B, 3 mm; C, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5160352" httpUri="https://zenodo.org/record/5160352/files/figure.png" pageId="16" pageNumber="211">Fig. 6C</figureCitation>
); ventral margins of tibia 2, tarsus and propodus with dense row of distally squared, split spinules (
<figureCitation id="136CFFAD8323DA53FCC4FA43830F1245" box="[776,860,1515,1542]" captionStart="FIG" captionStartId="16.[113,124,1317,1335]" captionTargetBox="[113,1209,215,1275]" captionTargetId="figure-220@16.[113,1209,215,1275]" captionTargetPageId="16" captionText="FIG. 6. – Pycnogonum staplesi n. sp., holotype (Py 922): A, habitus, lateral (note, mid-dorsal tubercle on trunk segment 2 broken); B, habitus, dorsal; C, distal articles of third leg, with detail of split spinule. Scale bar: A, B, 3 mm; C, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5160352" httpUri="https://zenodo.org/record/5160352/files/figure.png" pageId="16" pageNumber="211">Fig. 6C</figureCitation>
). Coxa-1 without tubercle; coxa- 2 1.3 times as long as coxa-1, coxa-3 just shorter than coxa-2; femur 2.3 times as long as coxa-2, 3.2 times as long as wide, with rounded dorsodistal spur; tibia-1 as long as femur, 2.5 times as long as wide; tibia-2 0.7 times as long as femur, 2.7 times as long as wide; tarsus short, one-quarter as long as propodus, longer ventrally than dorsally; propodus slender, 3.1 times as long as wide, with dorsodistal tuft of three spinules; main claw half as long as propodus; auxiliary claws absent. Gonopores and coxal glands (
<emphasis id="B9233F3A8322DA52FEC8FEDF816B15D3" box="[260,312,375,400]" italics="true" pageId="17" pageNumber="212">sensu</emphasis>
<bibRefCitation id="EFC69ED98322DA52FEF3FEDE819A15D3" author="STAPLES D." box="[319,457,374,400]" pageId="17" pageNumber="212" pagination="541 - 553" refId="ref11325" refString="STAPLES D. 2002. - Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria 59: 541 - 553." type="journal article" year="2002">Staples 2002</bibRefCitation>
) not seen.
</paragraph>
<paragraph id="8BE8E3288322DA52FF44FE3E813B160D" blockId="17.[112,648,216,590]" pageId="17" pageNumber="212">
Measurements of
<typeStatus id="54EC5D8A8322DA52FE86FE3E81FA15F3" box="[330,425,406,432]" pageId="17" pageNumber="212" type="holotype">holotype</typeStatus>
(mm): trunk length 2.91; width across second lateral processes 2.1; proboscis length 1.56; abdomen length 0.8; third leg, coxa-1 0.43, coxa-2 0.57, coxa-3 0.55, femur 1.31, tibia-1 1.31, tibia-2 0.93, tarsus 0.17, propodus 0.68, claw 0.37.
</paragraph>
</subSubSection>
<subSubSection id="C34DB0A38322DA52FFBDFDDC84EB164C" pageId="17" pageNumber="212" type="discussion">
<paragraph id="8BE8E3288322DA52FFBDFDDC808616CF" blockId="17.[111,649,628,1699]" box="[113,213,628,654]" pageId="17" pageNumber="212">REMARKS</paragraph>
<paragraph id="8BE8E3288322DA52FFBDFD3B82DB1049" blockId="17.[111,649,628,1699]" pageId="17" pageNumber="212">
<bibRefCitation id="EFC69ED98322DA52FFBDFD3B815716EE" author="STOCK J. H." box="[113,260,659,685]" pageId="17" pageNumber="212" pagination="389 - 409" refId="ref11534" refString="STOCK J. H. 1997. - Pycnogonids collected in recent years around New Caledonia and Vanuatu. In: CROSNIER A. (ed.), Resultats des Campagnes MUSORSTOM, Vol. 18 Memoires du Museum national d'Histoire naturelle, Paris 176: 389 - 409." type="journal article" year="1997">Stock (1997)</bibRefCitation>
recorded this specimen, which he attributed to
<taxonomicName id="4C5798AB8322DA52FF33FD1C821A168E" authority="Loman, 1908" authorityName="Loman" authorityYear="1908" box="[255,585,691,718]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A8322DA52FF33FD1C81E2168E" box="[255,433,692,717]" italics="true" pageId="17" pageNumber="212">Pycnogonum occa</emphasis>
<bibRefCitation id="EFC69ED98322DA52FE7AFD1C821A168E" author="LOMAN J. C. C." box="[438,585,691,718]" pageId="17" pageNumber="212" pagination="1 - 88" refId="ref11253" refString="LOMAN J. C. C. 1908. - Die Pantopoden der Siboga-Expedition. Siboga Expeditie Monographie 40: 1 - 88; pls I - XV." type="journal article" year="1908">Loman, 1908</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F340A3B88322DA52FFBCFD7B814F16AE" box="[112,284,723,749]" name="New Caledonia" pageId="17" pageNumber="212">New Caledonia</collectingCountry>
.
<bibRefCitation id="EFC69ED98322DA52FEEBFD7B819616AE" author="STAPLES D." box="[295,453,723,749]" pageId="17" pageNumber="212" pagination="541 - 553" refId="ref11325" refString="STAPLES D. 2002. - Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria 59: 541 - 553." type="journal article" year="2002">Staples (2002)</bibRefCitation>
, in his analysis of species of
<taxonomicName id="4C5798AB8322DA52FF15FD5B810A174F" authorityName="Brunnich" authorityYear="1764" box="[217,345,755,780]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B9233F3A8322DA52FF15FD5B810A174F" box="[217,345,755,780]" italics="true" pageId="17" pageNumber="212">Pycnogonum</emphasis>
</taxonomicName>
from
<collectingCountry id="F340A3B88322DA52FE5BFD5A81A4174F" box="[407,503,754,780]" name="Australia" pageId="17" pageNumber="212">Australia</collectingCountry>
, discussed the difficulty in interpreting the various records of
<taxonomicName id="4C5798AB8322DA52FD8AFCBB82D7176F" authorityName="Loman" authorityYear="1908" box="[582,644,787,812]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A8322DA52FD8AFCBB82D7176F" box="[582,644,787,812]" italics="true" pageId="17" pageNumber="212">P.occa</emphasis>
</taxonomicName>
in the literature, and in particular noted that
<bibRefCitation id="EFC69ED98322DA52FD87FC9A80EA172F" author="STOCK J. H." pageId="17" pageNumber="212" pagination="389 - 409" refId="ref11534" refString="STOCK J. H. 1997. - Pycnogonids collected in recent years around New Caledonia and Vanuatu. In: CROSNIER A. (ed.), Resultats des Campagnes MUSORSTOM, Vol. 18 Memoires du Museum national d'Histoire naturelle, Paris 176: 389 - 409." type="journal article" year="1997">Stock (1997)</bibRefCitation>
did not comment on the morphology of his specimen, so no comparison with this specimen could be made. Considering the possibility that this
<collectingCountry id="F340A3B88322DA52FFA3FC1981471789" box="[111,276,944,971]" name="New Caledonia" pageId="17" pageNumber="212">New Caledonia</collectingCountry>
specimen might have been actually of the previous species, Stock’s specimen has been re-examined, and determined to be a distinct species.
</paragraph>
<paragraph id="8BE8E3288322DA52FF4AFBA7811110EA" blockId="17.[111,649,628,1699]" pageId="17" pageNumber="212">
With its lateral-process tubercles, lack of a middorsal tubercle on the fourth trunk segment, blunt ocular tubercle and peculiar spinulation of the propodus, tarsus and tibia-2, this specimen is not
<taxonomicName id="4C5798AB8322DA52FFBDFB27816810EB" authorityName="Bamber" authorityYear="2013" box="[113,315,1167,1192]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="papua">
<emphasis id="B9233F3A8322DA52FFBDFB27816810EB" box="[113,315,1167,1192]" italics="true" pageId="17" pageNumber="212">Pycnogonum papua</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE8E3288322DA52FF44FB0684E91593" blockId="17.[111,649,628,1699]" lastBlockId="17.[674,1212,216,527]" pageId="17" pageNumber="212">
Neither is it
<taxonomicName id="4C5798AB8322DA52FEC9FB0781E1108B" authorityName="Loman" authorityYear="1908" box="[261,434,1199,1224]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A8322DA52FEC9FB0781E1108B" box="[261,434,1199,1224]" italics="true" pageId="17" pageNumber="212">Pycnogonum occa</emphasis>
</taxonomicName>
: in comparison with
<bibRefCitation id="EFC69ED98322DA52FFA3FB66814410AB" author="LOMAN J. C. C." box="[111,279,1230,1256]" pageId="17" pageNumber="212" pagination="1 - 88" refId="ref11253" refString="LOMAN J. C. C. 1908. - Die Pantopoden der Siboga-Expedition. Siboga Expeditie Monographie 40: 1 - 88; pls I - XV." type="journal article" year="1908">Loman’s (1908)</bibRefCitation>
figures and description, the lateral processes are not sufficiently separated, the ocular tubercle is not spired, the lateral-process tubercles are comparatively enormous, there is no middorsal tubercle on the fourth trunk segment, and, in particular, there is an array of distally squared, split spinules along the propodal sole, the ventral margin of the tarsus and the ventral margin of the distal third of tibia-2 (
<figureCitation id="136CFFAD8322DA52FE92FA6381E011A5" box="[350,435,1483,1510]" captionStart="FIG" captionStartId="16.[113,124,1317,1335]" captionTargetBox="[113,1209,215,1275]" captionTargetId="figure-220@16.[113,1209,215,1275]" captionTargetPageId="16" captionText="FIG. 6. – Pycnogonum staplesi n. sp., holotype (Py 922): A, habitus, lateral (note, mid-dorsal tubercle on trunk segment 2 broken); B, habitus, dorsal; C, distal articles of third leg, with detail of split spinule. Scale bar: A, B, 3 mm; C, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5160352" httpUri="https://zenodo.org/record/5160352/files/figure.png" pageId="17" pageNumber="212">Fig. 6C</figureCitation>
). While some other species of
<taxonomicName id="4C5798AB8322DA52FF17FA43810D1247" authorityName="Brunnich" authorityYear="1764" box="[219,350,1515,1540]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B9233F3A8322DA52FF17FA43810D1247" box="[219,350,1515,1540]" italics="true" pageId="17" pageNumber="212">Pycnogonum</emphasis>
</taxonomicName>
have split spines on the legs (e.g.,
<taxonomicName id="4C5798AB8322DA52FF7CF9A381F01266" authority="Staples, 2002" authorityName="Staples" authorityYear="2002" box="[176,419,1546,1573]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B9233F3A8322DA52FF7CF9A381561267" box="[176,261,1546,1572]" italics="true" pageId="17" pageNumber="212">P. clarki</emphasis>
<bibRefCitation id="EFC69ED98322DA52FEC2F9A281F01266" author="STAPLES D." box="[270,419,1546,1573]" pageId="17" pageNumber="212" pagination="541 - 553" refId="ref11325" refString="STAPLES D. 2002. - Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria 59: 541 - 553." type="journal article" year="2002">Staples, 2002</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C5798AB8322DA52FE7CF9A380881207" authority="Bamber, 2008" authorityName="Bamber" authorityYear="2008" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="coninsulum">
<emphasis id="B9233F3A8322DA52FE7CF9A382101267" box="[432,579,1546,1572]" italics="true" pageId="17" pageNumber="212">P. coninsulum</emphasis>
Bamber, 2008
</taxonomicName>
), those spinules are distally pointed, not truncate.
<bibRefCitation id="EFC69ED98322DA52FF16F9E281271227" author="LOMAN J. C. C." box="[218,372,1610,1636]" pageId="17" pageNumber="212" pagination="1 - 88" refId="ref11253" refString="LOMAN J. C. C. 1908. - Die Pantopoden der Siboga-Expedition. Siboga Expeditie Monographie 40: 1 - 88; pls I - XV." type="journal article" year="1908">Loman (1908</bibRefCitation>
: pl. XII, fig. 174) clearly shows sparse and simple ventral spinulation on these articles, consistent with the morphology of Staples’ (2002) specimen. In fact, the density of this spinulation is more in agreement with the “
<taxonomicName id="4C5798AB8322DA52FBA7FF5084FE1552" authorityName="Loman" authorityYear="1908" box="[1131,1197,248,273]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A8322DA52FBA7FF5084FE1552" box="[1131,1197,248,273]" italics="true" pageId="17" pageNumber="212">P. occa</emphasis>
</taxonomicName>
” specimen of
<bibRefCitation id="EFC69ED98322DA52FCE0FEBF84521572" author="STOCK J. H." box="[812,1025,279,305]" pageId="17" pageNumber="212" pagination="7 - 65" refId="ref11413" refString="STOCK J. H. 1968. - Pycnogonida collected by the Galathea and Anton Bruun in the Indian and Pacific Oceans. Vikenskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjobenhavn 131: 7 - 65; 1 - 22." type="journal article" year="1968">Stock (1968: 61-62</bibRefCitation>
, fig. 22c–e) from the Kermadec Trench, although that specimen does have a tubercle on the fourth trunk segment, and also peculiar tubercles along the frontal margin of the cephalon (the latter precluding it from being
<taxonomicName id="4C5798AB8322DA52FD6AFE1E82B6158C" authorityName="Loman" authorityYear="1908" box="[678,741,438,463]" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" kingdom="Animalia" order="Pantopoda" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="occa">
<emphasis id="B9233F3A8322DA52FD6AFE1E82B6158C" box="[678,741,438,463]" italics="true" pageId="17" pageNumber="212">P. occa</emphasis>
</taxonomicName>
; see also
<bibRefCitation id="EFC69ED98322DA52FC8EFE1D83B51593" author="CHILD C. A." box="[834,998,437,464]" pageId="17" pageNumber="212" pagination="i - iv" refId="ref11140" refString="CHILD C. A. 1988. - Pycnogonida of the Western Pacific Islands, III: Recent Smithsonian-Philippine Expeditions. Smithsonian Contributions to Zoology, 468: i - iv + 32 p." type="book chapter" year="1988">Child 1988: 27</bibRefCitation>
;
<bibRefCitation id="EFC69ED98322DA52FC23FE1D84FE1593" author="STAPLES D." box="[1007,1197,437,464]" pageId="17" pageNumber="212" pagination="541 - 553" refId="ref11325" refString="STAPLES D. 2002. - Pycnogonum (Pycnogonida: Pycnogonidae) from Australia with descriptions of two new species. Memoirs of the Museum of Victoria 59: 541 - 553." type="journal article" year="2002">Staples 2002: 547</bibRefCitation>
).
</paragraph>
<paragraph id="8BE8E3288322DA52FD77FE7D84EB164C" blockId="17.[674,1212,216,527]" pageId="17" pageNumber="212">
<bibRefCitation id="EFC69ED98322DA52FD77FE7D831215AC" author="STOCK J. H." box="[699,833,469,495]" pageId="17" pageNumber="212" pagination="389 - 409" refId="ref11534" refString="STOCK J. H. 1997. - Pycnogonids collected in recent years around New Caledonia and Vanuatu. In: CROSNIER A. (ed.), Resultats des Campagnes MUSORSTOM, Vol. 18 Memoires du Museum national d'Histoire naturelle, Paris 176: 389 - 409." type="journal article" year="1997">Stock (1997)</bibRefCitation>
called the present specimen a female, but I can find no gonopores to confirm or deny this.
</paragraph>
</subSubSection>
</treatment>
</document>
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