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<mods:title id="4E115F6001F0569EDD3CDC13CCE5C014">A new giant species of Deltochilum subgenus Deltohyboma (Coleoptera, Scarabaeidae, Scarabaeinae) from Colombia, with notes on D. spinipes Paulian, 1938</mods:title>
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<mods:namePart id="9507FDA6575018FD00B63BB1EBD69927">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="7E58A1447F7548FBFD08E8F12B8B1641">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<treatment id="03F387EFFF8FEA28FF35FF02F25EFE58" ID-DOI="http://doi.org/10.5281/zenodo.6137918" ID-GBIF-Taxon="119428679" ID-Zenodo-Dep="6137918" LSID="urn:lsid:plazi:treatment:03F387EFFF8FEA28FF35FF02F25EFE58" httpUri="http://treatment.plazi.org/id/03F387EFFF8FEA28FF35FF02F25EFE58" lastPageId="7" lastPageNumber="283" pageId="5" pageNumber="281">
<subSubSection id="C3406572FF8FEA2AFF35FF02F2A5FC9F" pageId="5" pageNumber="281" type="nomenclature">
<paragraph id="8BE536F9FF8FEA2AFF35FF02F3F1FF1C" blockId="5.[151,743,151,213]" box="[151,743,151,176]" pageId="5" pageNumber="281">
<heading id="D0AD8195FF8FEA2AFF35FF02F3F1FF1C" bold="true" box="[151,743,151,176]" fontSize="10" level="2" pageId="5" pageNumber="281" reason="2">
<taxonomicName id="4C5A4D7AFF8FEA2AFF35FF02F3F1FF1C" ID-CoL="34KCV" authority="Paulian, 1938" authorityName="Paulian" authorityYear="1938" box="[151,743,151,176]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes" subGenus="Deltohyboma">
<emphasis id="B92EEAEBFF8FEA2AFF35FF02F3F1FF1C" bold="true" box="[151,743,151,176]" pageId="5" pageNumber="281">
<emphasis id="B92EEAEBFF8FEA2AFF35FF02F328FF1C" bold="true" box="[151,574,151,176]" italics="true" pageId="5" pageNumber="281">Deltochilum (Deltohyboma) spinipes</emphasis>
<bibRefCitation id="EFCB4B08FF8FEA2AFDE7FF02F3F1FF1C" author="Paulian" box="[581,743,151,176]" pageId="5" pageNumber="281" refString="Paulian, R. (1938) Contribution a lʼetude des Canthonides americains. Annales de la Societe Entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian, 1938</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF35FF29F1FDFF78" blockId="5.[151,743,151,213]" box="[151,235,188,213]" pageId="5" pageNumber="281">
(
<figureCitation id="13612A7CFF8FEA2AFF3DFF29F1F4FF78" box="[159,226,188,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[182,1402,193,1929]" captionTargetId="figure@6.[172,1415,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Deltochilum spinipes. (A) lectotype ♂, dorsal view; (B) paralectotype ♂ 1, dorsal view; (C) paralectotype ♂ 1, ventral view; (D) paralectotype ♀ 1, dorsal view; (E) paralectotype ♀ 1, ventral view; (F) paralectotype ♂ 1, aedeagus (lateral view); (G) paralectotype ♂ 1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN)." httpUri="https://zenodo.org/record/230063/files/figure.png" pageId="5" pageNumber="281">Fig. 3</figureCitation>
)
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF35FE91F043FEB6" blockId="5.[151,341,260,282]" box="[151,341,260,282]" pageId="5" pageNumber="281">
<bibRefCitation id="EFCB4B08FF8FEA2AFF35FE91F033FEB6" author="Paulian" box="[151,293,260,282]" pageId="5" pageNumber="281" refString="Paulian, R. (1938) Contribution a lʼetude des Canthonides americains. Annales de la Societe Entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian, 1938</bibRefCitation>
: 280
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF35FED3F49DFDBF" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
<emphasis id="B92EEAEBFF8FEA2AFF35FED3F1F7FEF3" bold="true" box="[151,225,326,351]" pageId="5" pageNumber="281">Notes.</emphasis>
Paulian described this species based on several specimens from his collection as well as the collections of Oberthür, Boucomont, and the British Museum. He designated a pair of specimens from the Oberthür collection as &quot;
<typeStatus id="54E1885BFF8FEA2AFF02FE1AF1F6FE0B" box="[160,224,399,423]" pageId="5" pageNumber="281">types</typeStatus>
&quot; and the others as &quot;cotypes&quot;&quot;, but there was no formal
<typeStatus id="54E1885BFF8FEA2AFC27FE1BF2FBFE0B" box="[901,1005,398,423]" pageId="5" pageNumber="281" type="holotype">Holotype</typeStatus>
designation and all
<typeStatus id="54E1885BFF8FEA2AFB47FE1AF403FE0B" box="[1253,1301,399,423]" pageId="5" pageNumber="281">type</typeStatus>
material is therefore syntypic. All
<typeStatus id="54E1885BFF8FEA2AFE3CFE26F316FE67" box="[414,512,435,459]" pageId="5" pageNumber="281" type="syntype">syntypes</typeStatus>
in MNHN of
<taxonomicName id="4C5A4D7AFF8FEA2AFD01FE21F234FE67" box="[675,802,436,459]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B92EEAEBFF8FEA2AFD01FE21F234FE67" box="[675,802,436,459]" italics="true" pageId="5" pageNumber="281">D. spinipes</emphasis>
</taxonomicName>
have a body length less than
<quantity id="4CA29B1CFF8FEA2AFBD6FE26F5D1FE66" box="[1140,1223,435,459]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="281" unit="mm" value="20.0">20 mm</quantity>
. However, Paulian states the length for the species as &quot;
<quantity id="4CA29B1CFF8FEA2AFD8FFE42F3BCFE43" box="[557,682,471,495]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="5" pageNumber="281" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">18-24 mm</quantity>
&quot;. Based on this fact, we believe that the
<quantity id="4CA29B1CFF8FEA2AFBD8FE42F5DDFE42" box="[1146,1227,471,495]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="281" unit="mm" value="24.0">24 mm</quantity>
female found in a separate box (female of
<taxonomicName id="4C5A4D7AFF8FEA2AFE07FE69F30DFDBF" box="[421,539,508,531]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="gigante">
<emphasis id="B92EEAEBFF8FEA2AFE07FE69F30DFDBF" box="[421,539,508,531]" italics="true" pageId="5" pageNumber="281">D. gigante</emphasis>
</taxonomicName>
, see above) was regarded by Paulian as
<taxonomicName id="4C5A4D7AFF8FEA2AFC7DFE69F52DFDBF" box="[991,1083,508,531]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B92EEAEBFF8FEA2AFC7DFE69F52DFDBF" box="[991,1083,508,531]" italics="true" pageId="5" pageNumber="281">spinipes</emphasis>
</taxonomicName>
while describing this species.
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65FD8BF593FDF7" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
The
<typeStatus id="54E1885BFF8FEA2AFF5CFD8AF038FD9B" box="[254,302,543,567]" pageId="5" pageNumber="281">type</typeStatus>
series of
<taxonomicName id="4C5A4D7AFF8FEA2AFE0AFD8AF312FD9A" box="[424,516,543,566]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B92EEAEBFF8FEA2AFE0AFD8AF312FD9A" box="[424,516,543,566]" italics="true" pageId="5" pageNumber="281">spinipes</emphasis>
</taxonomicName>
shows variation and we strongly believe it to include more than one species. Therefore, in order to maintain stability of nomenclature, a
<typeStatus id="54E1885BFF8FEA2AFC93FDD7F28EFDF7" box="[817,920,578,603]" pageId="5" pageNumber="281" type="lectotype">lectotype</typeStatus>
is herein designated.
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65FDF2F2A5FC9F" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
Two of the
<typeStatus id="54E1885BFF8FEA2AFEE5FDF3F0FFFDD3" box="[327,489,614,639]" pageId="5" pageNumber="281" type="paralectotype">paralectotypes</typeStatus>
(♀4 and ♀5) appear to be non-conspecific with the remainder of the
<typeStatus id="54E1885BFF8FEA2AFB57FDF2F433FDD3" box="[1269,1317,615,639]" pageId="5" pageNumber="281">type</typeStatus>
series and likely represent two different species. They differ in being blue (not bright green), much darker and dull; one specimen is bigger and duller than the other; both specimens were collected in
<collectingCountry id="F34D7669FF8FEA2AFB9CFD3BF58DFD6B" box="[1086,1179,686,711]" name="Ecuador" pageId="5" pageNumber="281">Ecuador</collectingCountry>
(not
<collectingCountry id="F34D7669FF8FEA2AFB7CFD3BF45AFD6B" box="[1246,1356,686,711]" name="Colombia" pageId="5" pageNumber="281">Colombia</collectingCountry>
as the remainder of the
<typeStatus id="54E1885BFF8FEA2AFEF7FD46F093FD47" box="[341,389,723,747]" pageId="5" pageNumber="281">type</typeStatus>
series). These two specimens, at a first glance, most likely belong to the species-complex that includes
<taxonomicName id="4C5A4D7AFF8FEA2AFEA6FD62F0DEFCA3" box="[260,456,758,783]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="aequinoctiale">
<emphasis id="B92EEAEBFF8FEA2AFEA6FD62F0DEFCA3" box="[260,456,758,783]" italics="true" pageId="5" pageNumber="281">D. aequinoctiale</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7AFF8FEA2AFDB3FD62F3FCFCA2" box="[529,746,759,782]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="speciosissimum">
<emphasis id="B92EEAEBFF8FEA2AFDB3FD62F3FCFCA2" box="[529,746,759,782]" italics="true" pageId="5" pageNumber="281">D. speciosissimum</emphasis>
</taxonomicName>
. However, we cannot be sure about the identity of
<taxonomicName id="4C5A4D7AFF8FEA2AFA23FD62F039FC9F" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="aequinoctiale">
<emphasis id="B92EEAEBFF8FEA2AFA23FD62F039FC9F" italics="true" pageId="5" pageNumber="281">D. aequinoctiale</emphasis>
</taxonomicName>
since its
<typeStatus id="54E1885BFF8FEA2AFE3AFC8EF0DEFC9F" box="[408,456,795,819]" pageId="5" pageNumber="281">type</typeStatus>
has not yet been located (see
<bibRefCitation id="EFCB4B08FF8FEA2AFCBAFC8FF2B1FC9F" author="Genier" box="[792,935,794,819]" pageId="5" pageNumber="281" refString="Genier, F. (2001) Note sur les especes de Deltochilum Eschscholtz decrites en 1939 par V. Balthasar (Coleoptera: Scarabaeidae, Scarabaeinae). Fabreries, 26, 1 - 8." type="journal article" year="2001">Génier 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3406572FF8FEA28FF65FCABF25EFE58" lastPageId="7" lastPageNumber="283" pageId="5" pageNumber="281" type="materials_examined">
<paragraph id="8BE536F9FF8FEA2AFF65FCABF3CFFB83" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
<emphasis id="B92EEAEBFF8FEA2AFF65FCABF21EFCFB" bold="true" box="[199,776,830,855]" pageId="5" pageNumber="281">
Material examined.
<typeStatus id="54E1885BFF8FEA2AFE19FCABF327FCFB" box="[443,561,830,855]" pageId="5" pageNumber="281" type="lectotype">Lectotype</typeStatus>
(here designated)
</emphasis>
: ♂. (old white label, handwritten). Bogota. / 2 (old white label, printed and bordered in black). Ex-Musaeo Mniszech./ 3 (white label, handwritten).
<taxonomicName id="4C5A4D7AFF8FEA2AFB2FFCF6F41AFCD7" box="[1165,1292,866,891]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes" status="sp. nov.">D. spinipes</taxonomicName>
<taxonomicNameLabel id="A21D5790FF8FEA2AFAB1FCF6F459FCD6" box="[1299,1359,867,890]" pageId="5" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="54E1885BFF8FEA2AFAF4FCF6F486FCD7" box="[1366,1424,867,891]" pageId="5" pageNumber="281">Type</typeStatus>
./ 4 (white label, bordered in black) R. PAULIAN Vidit./ 5 (white label, printed in black). MUSÉUM
<collectionCode id="ED4BAE3CFF8FEA2AFAADFC12F44AFC33" box="[1295,1372,903,927]" pageId="5" pageNumber="281">PARIS</collectionCode>
1952 coll. R. OBERTHÜR./ 6 (red label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFCBDFC3EF2B6FC6F" box="[799,928,939,963]" pageId="5" pageNumber="281" type="syntype">SYNTYPE</typeStatus>
./ 7 (white label, printed in black)
<collectionCode id="ED4BAE3CFF8FEA2AFAE5FC3EF48AFC6E" LSID="urn:lsid:biocol.org:col:34988" box="[1351,1436,939,962]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
<collectionCode id="ED4BAE3CFF8FEA2AFF35FC5BF1AAFC4B" box="[151,188,974,999]" pageId="5" pageNumber="281">EC</collectionCode>
2509./ 8 (red label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFDF5FC5AF3ECFC4B" box="[599,762,975,999]" pageId="5" pageNumber="281" type="lectotype">LECTOTYPE</typeStatus>
./ 9 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFF35FC67F316FBA7" authority="Paulian" authorityName="Paulian" box="[151,512,1010,1035]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
<collectionCode id="ED4BAE3CFF8FEA2AFDB5FC66F362FBA7" box="[535,628,1011,1035]" pageId="5" pageNumber="281">LECTO</collectionCode>
. F Vaz-de-Mello des. 2013.
<figureCitation id="13612A7CFF8FEA2AFC5DFC67F57DFBA7" box="[1023,1131,1010,1035]" captionStart="FIGURE 3" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[182,1402,193,1929]" captionTargetId="figure@6.[172,1415,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Deltochilum spinipes. (A) lectotype ♂, dorsal view; (B) paralectotype ♂ 1, dorsal view; (C) paralectotype ♂ 1, ventral view; (D) paralectotype ♀ 1, dorsal view; (E) paralectotype ♀ 1, ventral view; (F) paralectotype ♂ 1, aedeagus (lateral view); (G) paralectotype ♂ 1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN)." httpUri="https://zenodo.org/record/230063/files/figure.png" pageId="5" pageNumber="281">Figure 3</figureCitation>
A (copyright
<collectionCode id="ED4BAE3CFF8FEA2AFA81FC66F46DFBA7" LSID="urn:lsid:biocol.org:col:34988" box="[1315,1403,1011,1035]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
/A. Mantilleri).
<typeStatus id="54E1885BFF8FEA2AFEB5FB83F0D1FB83" box="[279,455,1046,1071]" pageId="5" pageNumber="281" type="paralectotype">
<emphasis id="B92EEAEBFF8FEA2AFEB5FB83F0D1FB83" bold="true" box="[279,455,1046,1071]" pageId="5" pageNumber="281">Paralectotypes</emphasis>
</typeStatus>
: 2 ♂♂, 5 ♀♀ (
<collectionCode id="ED4BAE3CFF8FEA2AFDD3FB82F3D8FB83" LSID="urn:lsid:biocol.org:col:34988" box="[625,718,1047,1071]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
):
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65FBAEF383FAAB" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
♂1. 1 (old white label, printed and bordered in black).
<collectingCountry id="F34D7669FF8FEA2AFC9DFBAEF2C7FBFF" box="[831,977,1083,1107]" name="Colombia" pageId="5" pageNumber="281">COLOMBIE</collectingCountry>
COLLECTION LE
<collectionCode id="ED4BAE3CFF8FEA2AFB1FFBAEF436FBFF" box="[1213,1312,1083,1107]" pageId="5" pageNumber="281">MOULT</collectionCode>
. / 2 (white label, handwritten and printed in black).
<taxonomicName id="4C5A4D7AFF8FEA2AFD22FBCAF2D9FBDB" authority="Paul. R. Paulian" authorityName="Paul. R. Paulian" box="[640,975,1118,1143]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">D. spinipes Paul. R. Paulian</taxonomicName>
det./ 3 (white label, printed in black). MUSÉUM
<collectionCode id="ED4BAE3CFF8FEA2AFEBAFB16F073FB37" box="[280,357,1155,1179]" pageId="5" pageNumber="281">PARIS</collectionCode>
1952 coll. R. OBERTHÜR./ 4 (red label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFC55FB16F56EFB37" box="[1015,1144,1155,1179]" pageId="5" pageNumber="281" type="syntype">SYNTYPE</typeStatus>
./ 5 (white label, printed in black)
<collectionCode id="ED4BAE3CFF8FEA2AFF4BFB32F028FB12" LSID="urn:lsid:biocol.org:col:34988" box="[233,318,1191,1214]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
<collectionCode id="ED4BAE3CFF8FEA2AFEE8FB33F079FB13" box="[330,367,1190,1215]" pageId="5" pageNumber="281">EC</collectionCode>
2508./ 6 (yellow label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFCF4FB32F557FB13" box="[854,1089,1191,1215]" pageId="5" pageNumber="281" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 7 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFE4CFB5FF220FB4F" authority="Paulian" authorityName="Paulian" box="[494,822,1226,1251]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013.
<figureCitation id="13612A7CFF8FEA2AFAB8FB5FF495FB4F" box="[1306,1411,1226,1251]" captionStart="FIGURE 3" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[182,1402,193,1929]" captionTargetId="figure@6.[172,1415,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Deltochilum spinipes. (A) lectotype ♂, dorsal view; (B) paralectotype ♂ 1, dorsal view; (C) paralectotype ♂ 1, ventral view; (D) paralectotype ♀ 1, dorsal view; (E) paralectotype ♀ 1, ventral view; (F) paralectotype ♂ 1, aedeagus (lateral view); (G) paralectotype ♂ 1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN)." httpUri="https://zenodo.org/record/230063/files/figure.png" pageId="5" pageNumber="281">Figures 3</figureCitation>
B, C, F and G (copyright
<collectionCode id="ED4BAE3CFF8FEA2AFE36FB7AF0FAFAAB" LSID="urn:lsid:biocol.org:col:34988" box="[404,492,1263,1287]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
/A. Mantilleri).
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65FA86F580FA3B" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
♂2. 1 (old white label, handwritten). Fusagasugá./ 2 (old white label, handwritten). pretiosum Harold./ 3 (old white label, printed and bordered in black). Ex-Musaeo E. Steinheil./ 4 (white label, bordered in black) R. PAULIAN Vidit./ 5 (yellow label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFD42FACEF2DDFADF" box="[736,971,1371,1395]" pageId="5" pageNumber="281" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 6 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFEC9FAEBF3A3FA3B" authority="Paulian" authorityName="Paulian" box="[363,693,1406,1431]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013.
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65FA36F305F93F" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
♀1. 1 (old white label, printed and bordered in black).
<collectingCountry id="F34D7669FF8FEA2AFC8FFA36F2A9FA17" box="[813,959,1443,1467]" name="Colombia" pageId="5" pageNumber="281">COLOMBIE</collectingCountry>
ANTIOQUIA./ 2 (old white label, printed and bordered in black) COLLON LE
<collectionCode id="ED4BAE3CFF8FEA2AFDE2FA52F3B7FA73" box="[576,673,1479,1503]" pageId="5" pageNumber="281">MOULT</collectionCode>
Naturaliste Paris. / 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (white label, printed in black). MUSÉUM
<collectionCode id="ED4BAE3CFF8FEA2AFD41FA7EF226F9AF" box="[739,816,1515,1539]" pageId="5" pageNumber="281">PARIS</collectionCode>
1952 coll. R. OBERTHÜR./ 5 (white label, printed in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFF47F99BF321F98B" authority="Paulian." authorityName="Paulian." box="[229,567,1550,1575]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B92EEAEBFF8FEA2AFF47F99BF0C2F98A" box="[229,468,1550,1575]" italics="true" pageId="5" pageNumber="281">Deltochilum spinipes</emphasis>
Paulian.
</taxonomicName>
/ 6 (red label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFC3CF99AF509F98B" box="[926,1055,1551,1575]" pageId="5" pageNumber="281" type="syntype">SYNTYPE</typeStatus>
./ 7 (white label, printed in black)
<collectionCode id="ED4BAE3CFF8FEA2AFF35F9A6F1FAF9E6" LSID="urn:lsid:biocol.org:col:34988" box="[151,236,1587,1610]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
<collectionCode id="ED4BAE3CFF8FEA2AFF56F9A7F00FF9E7" box="[244,281,1586,1611]" pageId="5" pageNumber="281">EC</collectionCode>
2510./ 8 (yellow label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFD42F9A6F2DDF9E7" box="[736,971,1587,1611]" pageId="5" pageNumber="281" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 9 (white label, handwritten, and printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFE3FF9C3F3FFF9C3" authority="Paulian" authorityName="Paulian" box="[413,745,1622,1647]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013.
<figureCitation id="13612A7CFF8FEA2AFB77F9C3F428F9C3" box="[1237,1342,1622,1647]" captionStart="FIGURE 3" captionStartId="6.[151,250,1951,1973]" captionTargetBox="[182,1402,193,1929]" captionTargetId="figure@6.[172,1415,193,1930]" captionTargetPageId="6" captionText="FIGURE 3. Deltochilum spinipes. (A) lectotype ♂, dorsal view; (B) paralectotype ♂ 1, dorsal view; (C) paralectotype ♂ 1, ventral view; (D) paralectotype ♀ 1, dorsal view; (E) paralectotype ♀ 1, ventral view; (F) paralectotype ♂ 1, aedeagus (lateral view); (G) paralectotype ♂ 1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN)." httpUri="https://zenodo.org/record/230063/files/figure.png" pageId="5" pageNumber="281">Figures 3</figureCitation>
D and E (copyright
<collectionCode id="ED4BAE3CFF8FEA2AFEB1F9EEF07DF93F" LSID="urn:lsid:biocol.org:col:34988" box="[275,363,1659,1683]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="281">MNHN</collectionCode>
/A. Mantilleri).
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65F90AF367F88F" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
♀2. 1 (old white label, handwritten). Muzo,
<collectingCountry id="F34D7669FF8FEA2AFD1BF90BF23DF91B" box="[697,811,1694,1719]" name="United States of America" pageId="5" pageNumber="281">Columbia</collectingCountry>
./ 2 (old white label, printed and bordered in black). Ex- Musaeo D. Sharp 1890./ 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (yellow label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFF35F972F094F953" box="[151,386,1767,1791]" pageId="5" pageNumber="281" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 5 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFBEEF973F48AF953" authority="Paulian" authorityName="Paulian" box="[1100,1436,1766,1791]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013.
</paragraph>
<paragraph id="8BE536F9FF8FEA2AFF65F8BAF01CF81F" blockId="5.[151,1437,326,1971]" pageId="5" pageNumber="281">
♀3. 1 (old white label, printed and bordered in black).
<collectingCountry id="F34D7669FF8FEA2AFC9DF8BAF2C7F8EB" box="[831,977,1839,1863]" name="Colombia" pageId="5" pageNumber="281">COLOMBIE</collectingCountry>
COLLECTION LE
<collectionCode id="ED4BAE3CFF8FEA2AFB1FF8BAF436F8EB" box="[1213,1312,1839,1863]" pageId="5" pageNumber="281">MOULT</collectionCode>
. / 2 (white label, bordered in black) R. PAULIAN Vidit./ 3 (yellow label, printed in black)
<typeStatus id="54E1885BFF8FEA2AFB9EF8C6F431F8C7" box="[1084,1319,1875,1899]" pageId="5" pageNumber="281" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 4 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8FEA2AFD7FF8E3F53AF823" authority="Paulian" authorityName="Paulian" box="[733,1068,1910,1935]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013.
</paragraph>
<caption id="DF256671FF8CEA29FF35F80AF58AF85E" httpUri="https://zenodo.org/record/230063/files/figure.png" pageId="6" pageNumber="282" targetBox="[182,1402,193,1929]" targetPageId="6">
<paragraph id="8BE536F9FF8CEA29FF35F80AF58AF85E" blockId="6.[151,1436,1950,2034]" pageId="6" pageNumber="282">
<emphasis id="B92EEAEBFF8CEA29FF35F80AF0EBF818" bold="true" box="[151,509,1950,1973]" pageId="6" pageNumber="282">
FIGURE 3.
<taxonomicName id="4C5A4D7AFF8CEA29FEBCF80BF0EBF818" box="[286,509,1950,1973]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="282" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B92EEAEBFF8CEA29FEBCF80BF0EBF818" bold="true" box="[286,509,1950,1973]" italics="true" pageId="6" pageNumber="282">Deltochilum spinipes</emphasis>
</taxonomicName>
</emphasis>
. (A) lectotype ♂, dorsal view; (B) paralectotype ♂1, dorsal view; (C) paralectotype ♂1, ventral view; (D) paralectotype ♀1, dorsal view; (E) paralectotype ♀1, ventral view; (F) paralectotype ♂1, aedeagus (lateral view); (G) paralectotype ♂1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN).
</paragraph>
</caption>
<paragraph id="8BE536F9FF8DEA28FF65FF0DF045FEEC" blockId="7.[151,1437,151,501]" pageId="7" pageNumber="283">
♀4. 1 (white label, printed and bordered in black)
<collectingCountry id="F34D7669FF8DEA28FCA0FF0DF271FF03" box="[770,871,152,175]" name="Ecuador" pageId="7" pageNumber="283">Equateur</collectingCountry>
Env. d'Ambato R.P. Irenée Blanc./ 2 (white label, bordered in black) R. PAULIAN Vidit./ 3 (yellow label, printed in black)
<typeStatus id="54E1885BFF8DEA28FC56FF29F5C9FF78" box="[1012,1247,188,212]" pageId="7" pageNumber="283" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 4 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8DEA28FD3CFF4AF2E6FF54" authority="Paulian" authorityName="Paulian" box="[670,1008,223,248]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013./ 5 (white label, handwritten, printed and bordered in black) DIFFERENT of
<taxonomicName id="4C5A4D7AFF8DEA28FBFBFE90F5C5FEB1" box="[1113,1235,260,285]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spinipes">D spinipes</taxonomicName>
lectot. F Vaz-de- Mello des. 2013.
</paragraph>
<paragraph id="8BE536F9FF8DEA28FF65FED9F25EFE58" blockId="7.[151,1437,151,501]" pageId="7" pageNumber="283">
♀5. 1 (old white label, handwritten)
<collectingCountry id="F34D7669FF8DEA28FDD5FED9F3C2FEC9" box="[631,724,332,357]" name="Ecuador" pageId="7" pageNumber="283">Ecuador</collectingCountry>
Buckley./ 2 (white label, printed and bordered in black) Ex- Musaeo H.W.Bates 1892./ 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (yellow label, printed in black)
<typeStatus id="54E1885BFF8DEA28FF35FE01F094FE00" box="[151,386,404,428]" pageId="7" pageNumber="283" type="paralectotype">PARALECTOTYPE</typeStatus>
./ 5 (white label, handwritten, printed and bordered in black)
<taxonomicName id="4C5A4D7AFF8DEA28FBEEFE01F48AFE01" authority="Paulian" authorityName="Paulian" box="[1100,1436,404,429]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spinipes">Deltochilum spinipes Paulian</taxonomicName>
PARALECTO. F Vaz-de-Mello des. 2013./ 6 (white label, handwritten, printed and bordered in black) DIFFERENT of
<taxonomicName id="4C5A4D7AFF8DEA28FEF1FE48F0DCFE59" box="[339,458,476,501]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="283" phylum="Arthropoda" rank="species" species="spinipes">D spinipes</taxonomicName>
lectot. F Vaz-de-Mello des. 2013.
</paragraph>
</subSubSection>
</treatment>
</document>