306 lines
50 KiB
XML
306 lines
50 KiB
XML
<document id="AD07E03C4804F648120AED6E554FE9BB" ID-CLB-Dataset="45677" ID-DOI="10.11646/zootaxa.3802.2.8" ID-GBIF-Dataset="d1e35dc1-d06f-4212-b764-8d021c1518a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="230060" ID-ZooBank="2797E064-00BF-4113-8F84-0AEDE0C991DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460426224671" checkinUser="plazi" docAuthor="Silva, Fernando A. B. & Vaz-De-Mello, Fernando Z." docDate="2014" docId="03F387EFFF8BEA2DFF35FD25F508F815" docLanguage="en" docName="zt03802p284.pdf" docOrigin="Zootaxa 3802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Deltohyboma) gigante Silva & Vaz-De-Mello, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="278" masterDocId="FFCAFF97FF8AEA2FFFA2FF95F116FFAC" masterDocTitle="A new giant species of Deltochilum subgenus Deltohyboma (Coleoptera, Scarabaeidae, Scarabaeinae) from Colombia, with notes on D. spinipes Paulian, 1938" masterLastPageNumber="284" masterPageNumber="276" pageNumber="277" updateTime="1698330713618" updateUser="ExternalLinkService">
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<mods:mods id="91895D1FCF9E0580A7F60A1C91D0929F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="611B128C020CC762A0242121BFECDD88">
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<mods:title id="7EBC665CA9D1C3A0AC23ACC051136F16">A new giant species of Deltochilum subgenus Deltohyboma (Coleoptera, Scarabaeidae, Scarabaeinae) from Colombia, with notes on D. spinipes Paulian, 1938</mods:title>
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<mods:name id="22FA507CD62D6D78450BB64811C539D0" type="personal">
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<mods:roleTerm id="81D7E318EA86C542255A12BC71DC15C9">Author</mods:roleTerm>
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<mods:namePart id="97D20B04BFB47968DADC7587C594CD58">Silva, Fernando A. B.</mods:namePart>
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<mods:name id="9E0A3C0FA91CC73B2B63872E8721FADD" type="personal">
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<mods:role id="75FEBEE4FD25BC72D1DF8CFD59CE13AE">
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<mods:roleTerm id="3ED0F25E830292D3991C45AE261777A1">Author</mods:roleTerm>
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<mods:namePart id="9A3EF28608C688AD5CC604CD38B94E90">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="5F0D3427D5D28D29305AC4229E8A647F">text</mods:typeOfResource>
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<mods:title id="72F7A36F4B4DD1F284CA72F0D43A5F80">Zootaxa</mods:title>
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<mods:part id="1A78284CF43A5654045889EB05C3109C">
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<mods:date id="971E6134F3C14A55737F90CC7B4820FA">2014</mods:date>
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<mods:detail id="3F36CB124CE6AB6471C554944F6767C7" type="volume">
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<mods:number id="2D3ED998A7EDC30FC07EB73988956BFA">3802</mods:number>
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<mods:number id="2302E52C70A6E4CCE8BFC2A8C231F67D">2</mods:number>
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<mods:classification id="4AFD55100FE8320572BA90632C0A7B1F">journal article</mods:classification>
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<mods:identifier id="18E7A51990E3192052E710946EC56C30" type="CLB-Dataset">45677</mods:identifier>
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<mods:identifier id="DDCDAC5EBC71C8C4D12CDE43A3223983" type="DOI">10.11646/zootaxa.3802.2.8</mods:identifier>
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<mods:identifier id="A454299FE9C839E692AB7D6A086292C0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F387EFFF8BEA2DFF35FD25F508F815" ID-DOI="http://doi.org/10.5281/zenodo.6137916" ID-GBIF-Taxon="119428684" ID-Zenodo-Dep="6137916" LSID="urn:lsid:plazi:treatment:03F387EFFF8BEA2DFF35FD25F508F815" httpUri="http://treatment.plazi.org/id/03F387EFFF8BEA2DFF35FD25F508F815" lastPageId="2" lastPageNumber="278" pageId="1" pageNumber="277">
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<subSubSection id="C3406572FF8BEA2EFF35FD25F011FD40" pageId="1" pageNumber="277" type="nomenclature">
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<paragraph id="8BE536F9FF8BEA2EFF35FD25F3B8FD67" blockId="1.[151,686,688,749]" box="[151,686,688,715]" pageId="1" pageNumber="277">
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<heading id="D0AD8195FF8BEA2EFF35FD25F3B8FD67" bold="true" box="[151,686,688,715]" fontSize="11" level="1" pageId="1" pageNumber="277" reason="1">
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<emphasis id="B92EEAEBFF8BEA2EFF35FD25F3B8FD67" bold="true" box="[151,686,688,715]" pageId="1" pageNumber="277">
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<taxonomicName id="4C5A4D7AFF8BEA2EFF35FD25F35FFD66" ID-CoL="34KC6" box="[151,585,688,715]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="gigante" status="sp. nov." subGenus="Deltohyboma">
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<emphasis id="B92EEAEBFF8BEA2EFF35FD25F03BFD66" bold="true" box="[151,301,688,714]" italics="true" pageId="1" pageNumber="277">Deltochilum</emphasis>
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(
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<emphasis id="B92EEAEBFF8BEA2EFE9FFD25F35FFD66" bold="true" box="[317,585,688,714]" italics="true" pageId="1" pageNumber="277">Deltohyboma) gigante</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D5790FF8BEA2EFDF3FD24F3B8FD67" box="[593,686,689,715]" pageId="1" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE536F9FF8BEA2EFF35FD41F011FD40" blockId="1.[151,686,688,749]" box="[151,263,724,749]" pageId="1" pageNumber="277">
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(
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<figureCitation id="13612A7CFF8BEA2EFF3DFD41F1E8FD40" box="[159,254,724,749]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1925,1947]" captionStartId-1="4.[151,250,1909,1931]" captionTargetBox-0="[222,1365,195,1884]" captionTargetBox-1="[213,1364,209,1864]" captionTargetId-0="figure@3.[216,1370,195,1884]" captionTargetId-1="figure@4.[202,1369,193,1868]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." captionText-1="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri-0="https://zenodo.org/record/230061/files/figure.png" httpUri-1="https://zenodo.org/record/230062/files/figure.png" pageId="1" pageNumber="277">Fig. 1–2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3406572FF8BEA2EFF35FC89F3EEFCF4" pageId="1" pageNumber="277" type="etymology">
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<paragraph id="8BE536F9FF8BEA2EFF35FC89F3EEFCF4" blockId="1.[151,1437,796,2009]" pageId="1" pageNumber="277">
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<emphasis id="B92EEAEBFF8BEA2EFF35FC89F008FC99" bold="true" box="[151,286,796,821]" pageId="1" pageNumber="277">Etymology.</emphasis>
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The species name refers to the body size,
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<taxonomicName id="4C5A4D7AFF8BEA2EFD5AFC88F25AFC98" box="[760,844,797,820]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="gigante">
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<emphasis id="B92EEAEBFF8BEA2EFD5AFC88F25AFC98" box="[760,844,797,820]" italics="true" pageId="1" pageNumber="277">gigante</emphasis>
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</taxonomicName>
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meaning "giant" in both the Portuguese and Spanish languages. The species epithet is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3406572FF8BEA2EFF65FCF1F3BEFB49" pageId="1" pageNumber="277" type="diagnosis">
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<paragraph id="8BE536F9FF8BEA2EFF65FCF1F3BEFB49" blockId="1.[151,1437,796,2009]" pageId="1" pageNumber="277">
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<emphasis id="B92EEAEBFF8BEA2EFF65FCF1F057FCD1" bold="true" box="[199,321,868,893]" pageId="1" pageNumber="277">Diagnosis.</emphasis>
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The new species is placed in the group of species that comprises
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<taxonomicName id="4C5A4D7AFF8BEA2EFBD4FCF0F5EDFCD0" box="[1142,1275,869,892]" class="Insecta" family="Scarabaeidae" genus="Deltohyboma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="spinipes">
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<emphasis id="B92EEAEBFF8BEA2EFBD4FCF0F5EDFCD0" box="[1142,1275,869,892]" italics="true" pageId="1" pageNumber="277">D. spinipes</emphasis>
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</taxonomicName>
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, probably
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<taxonomicName id="4C5A4D7AFF8BEA2EFA23FCF0F0CEFC0C" authority="Buquet 1844" authorityName="Buquet" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="aequinoctiale">
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<emphasis id="B92EEAEBFF8BEA2EFA23FCF0F026FC0C" italics="true" pageId="1" pageNumber="277">D. aequinoctiale</emphasis>
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(Buquet 1844)
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</taxonomicName>
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(
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<typeStatus id="54E1885BFF8BEA2EFE44FC1DF30EFC0C" box="[486,536,904,928]" pageId="1" pageNumber="277">type</typeStatus>
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specimens not examined), and
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<taxonomicName id="4C5A4D7AFF8BEA2EFCD5FC1CF5E9FC0C" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[887,1279,903,928]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="speciosissimum">
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<emphasis id="B92EEAEBFF8BEA2EFCD5FC1CF551FC0C" box="[887,1095,905,928]" italics="true" pageId="1" pageNumber="277">D. speciosissimum</emphasis>
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Balthasar, 1939
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</taxonomicName>
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. This group is characterized by the following characters: size equal or greater than
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<quantity id="4CA29B1CFF8BEA2EFC70FC39F53CFC68" box="[978,1066,940,964]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="277" unit="mm" value="13.0">13 mm</quantity>
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; apical tubercles or carinae on interstriae 3-4-5-6, very weak or lacking on interstria 7; elytra lacking foveate punctures on discal striae. In this group of species,
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<taxonomicName id="4C5A4D7AFF8BEA2EFEF9FC60F303FBA1" box="[347,533,1012,1037]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="aequinoctiale">
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<emphasis id="B92EEAEBFF8BEA2EFEF9FC60F303FBA1" box="[347,533,1012,1037]" italics="true" pageId="1" pageNumber="277">D. aequinoctiale</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5A4D7AFF8BEA2EFDEEFC60F20DFBA0" box="[588,795,1013,1036]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="speciosissimum">
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<emphasis id="B92EEAEBFF8BEA2EFDEEFC60F20DFBA0" box="[588,795,1013,1036]" italics="true" pageId="1" pageNumber="277">D. speciosissimum</emphasis>
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</taxonomicName>
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have a microgranulated opaque dorsal integument, while
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<taxonomicName id="4C5A4D7AFF8BEA2EFF35FB8CF00FFB9C" box="[151,281,1049,1072]" class="Insecta" family="Scarabaeidae" genus="Deltohyboma" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="spinipes">
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<emphasis id="B92EEAEBFF8BEA2EFF35FB8CF00FFB9C" box="[151,281,1049,1072]" italics="true" pageId="1" pageNumber="277">D. spinipes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5A4D7AFF8BEA2EFEF5FB8CF0C7FB9C" box="[343,465,1049,1072]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="gigante">
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<emphasis id="B92EEAEBFF8BEA2EFEF5FB8CF0C7FB9C" box="[343,465,1049,1072]" italics="true" pageId="1" pageNumber="277">D. gigante</emphasis>
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</taxonomicName>
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share a smooth and shiny surface of the elytra and pronotum (punctures might be present but surface lacks microgranulation).
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<taxonomicName id="4C5A4D7AFF8BEA2EFD06FBA9F280FBF8" box="[676,918,1084,1109]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="spinipes">
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<emphasis id="B92EEAEBFF8BEA2EFD06FBA9F280FBF8" box="[676,918,1084,1109]" italics="true" pageId="1" pageNumber="277">Deltochilum spinipes</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C5A4D7AFF8BEA2EFB98FBA8F5A3FBF8" box="[1082,1205,1085,1108]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="gigante">
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<emphasis id="B92EEAEBFF8BEA2EFB98FBA8F5A3FBF8" box="[1082,1205,1085,1108]" italics="true" pageId="1" pageNumber="277">D. gigante</emphasis>
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</taxonomicName>
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by its significantly smaller size and the lack of the posterior metasternal spiniform projections as well as the internal tooth on protibia. In addition,
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<taxonomicName id="4C5A4D7AFF8BEA2EFE81FB10F0B5FB30" box="[291,419,1157,1180]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="spinipes">
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<emphasis id="B92EEAEBFF8BEA2EFE81FB10F0B5FB30" box="[291,419,1157,1180]" italics="true" pageId="1" pageNumber="277">D. spinipes</emphasis>
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</taxonomicName>
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has a stronger mesofemur with a larger tubercle on posterior margin, and a conspicuous apical carina on the seventh elytral stria. Furthermore,
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<taxonomicName id="4C5A4D7AFF8BEA2EFCBBFB3CF285FB6C" box="[793,915,1193,1216]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="gigante">
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<emphasis id="B92EEAEBFF8BEA2EFCBBFB3CF285FB6C" box="[793,915,1193,1216]" italics="true" pageId="1" pageNumber="277">D. gigante</emphasis>
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</taxonomicName>
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can be differentiated from all the remaining species in this subgenus by its larger body size.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3406572FF8BEA2DFF65FB7AF2ACF911" lastPageId="2" lastPageNumber="278" pageId="1" pageNumber="277" type="description">
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<paragraph id="8BE536F9FF8BEA2DFF65FB7AF271FBF4" blockId="1.[151,1437,796,2009]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="278" pageId="1" pageNumber="277">
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<emphasis id="B92EEAEBFF8BEA2EFF65FB7AF0CCFAA4" bold="true" box="[199,474,1263,1288]" pageId="1" pageNumber="277">
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Description (
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<typeStatus id="54E1885BFF8BEA2EFEC0FB7AF0C4FAA4" box="[354,466,1263,1288]" pageId="1" pageNumber="277" type="holotype">Holotype</typeStatus>
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)
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</emphasis>
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.
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<emphasis id="B92EEAEBFF8BEA2EFE4BFB7AF32DFAA4" bold="true" box="[489,571,1263,1288]" pageId="1" pageNumber="277">Colour</emphasis>
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. Completely metallic blue, with green to violaceous sheen (
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<figureCitation id="13612A7CFF8BEA2EFB4BFB7AF447FAA4" box="[1257,1361,1263,1288]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figures 1</figureCitation>
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A, B).
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<emphasis id="B92EEAEBFF8BEA2EFF35FA81F1FAFA81" bold="true" box="[151,236,1300,1325]" pageId="1" pageNumber="277">Length</emphasis>
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.
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<quantity id="4CA29B1CFF8BEA2EFF5FFA81F045FA80" box="[253,339,1300,1324]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="277" unit="mm" value="24.0">24 mm</quantity>
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.
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<emphasis id="B92EEAEBFF8BEA2EFEC0FA81F0B4FA81" bold="true" box="[354,418,1300,1325]" pageId="1" pageNumber="277">Head</emphasis>
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. Dorsal surface with dense foveiform punctures, closer on lateral portions (
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<figureCitation id="13612A7CFF8BEA2EFAB7FA81F460FA80" box="[1301,1398,1300,1325]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
C). Clypeal margin with two small, triangular, sharp, central teeth, separated by approximately five times the width of their basal portion. Margin between clypeal teeth and clypeo-genal suture almost straight. Margin of gena strongly curved outward. Clypeo-genal suture slightly impressed. Clypeo-frontal suture brighter than surrounding surface. Dorsal interocular width eight times the eye width (
|
||
<figureCitation id="13612A7CFF8BEA2EFCB3FA31F263FA10" box="[785,885,1444,1469]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
C). Antennae 9-segmented, club 3-segmented. Posterior dorsal margin of head (occipital region of front) completely marginated by fine carina.
|
||
<emphasis id="B92EEAEBFF8BEA2EFB4AFA52F457FA4C" bold="true" box="[1256,1345,1479,1504]" pageId="1" pageNumber="277">Thorax</emphasis>
|
||
. W/L = 1.3; pronotum strongly convex, with a smooth longitudinal depression medially (
|
||
<figureCitation id="13612A7CFF8BEA2EFB92FA79F598F9A8" box="[1072,1166,1516,1541]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
A). Dorsal surface with dense and small foveiform punctures, randomly spaced; punctures larger and closer on anterior angles and lateral portions of pronotum. Anterior margin deeply emarginated. Anterior angles acute, directed forward (
|
||
<figureCitation id="13612A7CFF8BEA2EFAB5F9A1F460F9E0" box="[1303,1398,1588,1613]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
C). Lateral pronotal impressions with a small rounded tubercle, poorly differentiated from surrounding surface. Margin between anterior and lateral angles oblique, slightly curved outward. Margin between lateral and posterior angles distinctly curved outward. Hypomera not excavated (
|
||
<figureCitation id="13612A7CFF8BEA2EFCB0F90AF260F914" box="[786,886,1695,1720]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
D); lacking carina separating the anterior and posterior area; foveiform punctures denser on anterior and posterior portions, with some long setae on anterior portion; median portion with sparse or absent punctures. Meso- and metepisternum with foveiform punctures. Mesosternum with dense foveiform punctures, almost touching on lateral portion. Mesometasternal suture arched, angulate medially. Metasternum with dense foveiform punctures on lateral and anterior portions; disc smooth, with a strong triangular depression (
|
||
<figureCitation id="13612A7CFF8BEA2EFDA7F8C1F371F8C0" box="[517,615,1876,1901]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
B); anterior portion of depression with a strong spiniform projection; posterior portion with two smaller spiniform projections.
|
||
<emphasis id="B92EEAEBFF8BEA2EFC8BF8E2F263F83C" bold="true" box="[809,885,1911,1936]" pageId="1" pageNumber="277">Elytra</emphasis>
|
||
. Elongated, with lateral margins slightly curved outward (
|
||
<figureCitation id="13612A7CFF8BEA2EFEABF809F078F818" box="[265,366,1948,1973]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
A). Elytral striae weakly impressed (
|
||
<figureCitation id="13612A7CFF8BEA2EFC91F809F28EF818" box="[819,920,1948,1973]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="1" pageNumber="277">Figure 1</figureCitation>
|
||
E), with foveiform punctures separated by approximately four or five times their diameter on elytral disc. Interstriae flattened. Interstrial surface with setigerous foveiform punctures, randomly spaced. Basal carina of seventh interstria inconspicuous. Basal carina of ninth interstria reaches approximately half the length of elytra. Apical tubercle of second elytral interstria absent. Apical tubercle of third interstria rounded. Apical carina of fifth and sixth interstriae longer than fourth. Apical carina of seventh interstria inconspicuous. Pseudepipleural carina complete, visible for entire length of epipleuron.
|
||
<emphasis id="B92EEAEBFF88EA2DFF35FEB2F01AFEEC" bold="true" box="[151,268,295,320]" pageId="2" pageNumber="278">Abdomen</emphasis>
|
||
. First sternite expanded medially, occupying 2/3 of the abdominal length (
|
||
<figureCitation id="13612A7CFF88EA2DFBEBFEB2F5B3FEEC" box="[1097,1189,295,320]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
F); anterior portion of sternite with two tubercles on its medial portion (
|
||
<figureCitation id="13612A7CFF88EA2DFD78FED9F22CFEC8" box="[730,826,332,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
F). Sternites 2, 3 and 4 not visible medially. Lateral region of sternites 1 – 5 with numerous foveiform punctures closer to anterior margin. Sixth sternite smaller medially than fourth. Pygidium separate from propygidium by carina, with dense setigerous foveiform punctures on its surface, punctures closer along the midline. Pygidium as wide as high, apically rounded, clearly margined.
|
||
<emphasis id="B92EEAEBFF88EA2DFF35FE49F1C0FE59" bold="true" box="[151,214,476,501]" pageId="2" pageNumber="278">Legs.</emphasis>
|
||
Protrochanter with a cluster of bristles on the anterior edge. Meso and metatrochanter with a cluster of bristles on the posterior edge (
|
||
<figureCitation id="13612A7CFF88EA2DFE47FE6AF354FDB4" box="[485,578,511,536]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
B). Femora with numerous foveiform punctures, randomly spaced. Profemur with clusters of bristles on basal third of anterior margin (
|
||
<figureCitation id="13612A7CFF88EA2DFC8BFDB1F291FD90" box="[809,903,548,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
B). Mesofemur widened at the middle portion, with a small denticle on basal third of posterior margin (
|
||
<figureCitation id="13612A7CFF88EA2DFCABFDD2F270FDCC" box="[777,870,583,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
B). Metafemur widened at its apical third. Protibia widened at 2/3 of the internal margin; apical third of the internal margin angularly lobed, with a concave curvature, forming a strong triangular tooth and fourteen spiniform denticles between tibial base and that tooth (
|
||
<figureCitation id="13612A7CFF88EA2DFAB4FD1AF462FD04" box="[1302,1396,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
G). External margin with three teeth on its apical third, apical one larger than basal two (
|
||
<figureCitation id="13612A7CFF88EA2DFB2FFD21F5F8FD60" box="[1165,1262,692,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
G). Meso- and metatibiae arched toward body. Mesotibia with an internal, apical, spur-like denticle, about the length of the first tarsomere; smaller spur shorter than the sum of the length of second and third metatarsomeres; larger spur approximately the sum of three basal tarsomeres in length (
|
||
<figureCitation id="13612A7CFF88EA2DFCC1FC8AF2D0FC94" box="[867,966,799,824]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
H). Basal third of metatibia with small denticles on the internal margin (
|
||
<figureCitation id="13612A7CFF88EA2DFDB4FCD1F363FCF0" box="[534,629,836,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figure 1</figureCitation>
|
||
A). Internal angle of apex of metatibia prolonged beyond insertion of tarsus in spiniform process, with the same length of tarsus (
|
||
<figureCitation id="13612A7CFF88EA2DFC95FCF2F28EFC2C" box="[823,920,871,896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
A). Length of metatibial spur less than that of tarsal claws. Protarsus absent. Length of first meso- and metatarsomeres short, about half the second tarsomere, obliquely truncated at apex (
|
||
<figureCitation id="13612A7CFF88EA2DFE7BFC3AF354FC64" box="[473,578,943,968]" captionStart="FIGURE 1" captionStartId="3.[151,250,1925,1947]" captionTargetBox="[222,1365,195,1884]" captionTargetId="figure@3.[216,1370,195,1884]" captionTargetPageId="3" captionText="FIGURE 1. Deltochilum gigante sp. nov. (holotype ♂). (A) dorsal view; (B) ventral view (spiniform projections arrowed); (C) head; (D) hypomera; (E) elytral disc; (F) male abdominal sternites (tubercles arrowed); (G) male protibia; (H) apex of mesotibia (spiniform spurs arrowed)." httpUri="https://zenodo.org/record/230061/files/figure.png" pageId="2" pageNumber="278">Figures 1</figureCitation>
|
||
H, 2A). Tarsal claws without basal teeth.
|
||
<emphasis id="B92EEAEBFF88EA2DFBB8FC3AF5E8FC64" bold="true" box="[1050,1278,943,968]" pageId="2" pageNumber="278">Male genital organ</emphasis>
|
||
. Symmetrical parameres (
|
||
<figureCitation id="13612A7CFF88EA2DFEBFFC41F098FC40" box="[285,398,980,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figures 2</figureCitation>
|
||
B, C). Genital sclerite with internal arms elongated and weakly sclerotized (
|
||
<figureCitation id="13612A7CFF88EA2DFAB1FC41F46FFC40" box="[1299,1401,980,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
D). Endophallus with small basal lobe (
|
||
<figureCitation id="13612A7CFF88EA2DFD80FC62F394FBBC" box="[546,642,1015,1040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
E). Lamella copulatrix (LC): left and right lobes subrectangular (
|
||
<figureCitation id="13612A7CFF88EA2DFAF3FC62F1B0FB98" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
F). Superior right peripheral sclerite (SRP) circular (
|
||
<figureCitation id="13612A7CFF88EA2DFCB2FB89F260FB98" box="[784,886,1052,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
G). Fronto-lateral peripheral sclerite (FLP) and complex of axial and subaxial sclerites (A+
|
||
<collectingCountry id="F34D7669FF88EA2DFDDCFBD5F3B5FBF4" box="[638,675,1088,1112]" name="American Samoa" pageId="2" pageNumber="278">AS</collectingCountry>
|
||
) as in
|
||
<figureCitation id="13612A7CFF88EA2DFD4FFBAAF25AFBF4" box="[749,844,1087,1112]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1931]" captionTargetBox="[213,1364,209,1864]" captionTargetId="figure@4.[202,1369,193,1868]" captionTargetPageId="4" captionText="FIGURE 2. Deltochilum gigante sp. nov. (holotype ♂). (A) apex of metatibia (arrow shows the internal angle of apex of metatibia prolonged beyond insertion of tarsus); (B) aedeagus (lateral view); (C) aedeagus (dorsal and ventral views); (D) genital sclerite; (E) endophallus; (F) lamella copulatrix (top left lobe; bottom right lobe); (G) superior right peripheral sclerite (SRP); (H) complex of axial and subaxial sclerites (A + AS) (arrow left), fronto-lateral peripheral sclerite (FLP) (arrow right)." httpUri="https://zenodo.org/record/230062/files/figure.png" pageId="2" pageNumber="278">Figure 2</figureCitation>
|
||
H.
|
||
</paragraph>
|
||
<paragraph id="8BE536F9FF88EA2DFF65FBF1F0F9FAD5" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="278">
|
||
<emphasis id="B92EEAEBFF88EA2DFF65FBF1F023FBD1" bold="true" box="[199,309,1124,1149]" pageId="2" pageNumber="278">Variation</emphasis>
|
||
: In the collection of the MNHN (Paris), we found a female lacking an abdomen and without locality data. We strongly believe that
|
||
<bibRefCitation id="EFCB4B08FF88EA2DFE50FB12F38FFB0C" author="Paulian" box="[498,665,1159,1184]" pageId="2" pageNumber="278" refString="Paulian, R. (1938) Contribution a lʼetude des Canthonides americains. Annales de la Societe Entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian (1938)</bibRefCitation>
|
||
considered this female,
|
||
<quantity id="4CA29B1CFF88EA2DFC13FB1DF517FB0C" box="[945,1025,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="278" unit="mm" value="24.0">24 mm</quantity>
|
||
in length, to be conspecific with
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFA23FB1CF1FEFB68" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spinipe">
|
||
<emphasis id="B92EEAEBFF88EA2DFA23FB1CF1FEFB68" italics="true" pageId="2" pageNumber="278">D. spinipe</emphasis>
|
||
</taxonomicName>
|
||
s specimens due to their body size overlap (see below notes for
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFC08FB38F50AFB68" box="[938,1052,1197,1220]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spinipe">
|
||
<emphasis id="B92EEAEBFF88EA2DFC08FB38F50AFB68" box="[938,1052,1197,1220]" italics="true" pageId="2" pageNumber="278">D. spinipe</emphasis>
|
||
</taxonomicName>
|
||
s), because he states in his original description that specimens belonging to this species reach up to
|
||
<quantity id="4CA29B1CFF88EA2DFCCFFB45F2A9FB44" box="[877,959,1232,1256]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="278" unit="mm" value="24.0">24 mm</quantity>
|
||
, and specimens of that size (or near) were found identified as
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFED0FB60F0E6FAA0" box="[370,496,1269,1292]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spinipes">
|
||
<emphasis id="B92EEAEBFF88EA2DFED0FB60F0E6FAA0" box="[370,496,1269,1292]" italics="true" pageId="2" pageNumber="278">D. spinipes</emphasis>
|
||
</taxonomicName>
|
||
by Paulian in MNHN. Although this female was found in a box of specimens used by Paulian in his revision (
|
||
<bibRefCitation id="EFCB4B08FF88EA2DFE60FA82F34CFA9C" author="Paulian" box="[450,602,1303,1328]" pageId="2" pageNumber="278" refString="Paulian, R. (1938) Contribution a lʼetude des Canthonides americains. Annales de la Societe Entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCB4B08FF88EA2DFDC7FA8DF3B6FA9C" author="Paulian" box="[613,672,1304,1328]" pageId="2" pageNumber="278" refString="Paulian, R. (1939) Contribution a l`etude des Canthonides americains. Annales de la Societe Entomologique de France, 108, 1 - 40." type="journal article" year="1939">1939</bibRefCitation>
|
||
), since it lacks any label, we cannot be sure that it is a
|
||
<typeStatus id="54E1885BFF88EA2DFAA6FA8CF44DFA9D" box="[1284,1371,1305,1329]" pageId="2" pageNumber="278" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFA23FA8CF1E5FAF8" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="spinipes">
|
||
<emphasis id="B92EEAEBFF88EA2DFA23FA8CF1E5FAF8" italics="true" pageId="2" pageNumber="278">D. spinipes</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, we are confident that this female belongs to the species herein described and have made this unlabeled female the
|
||
<typeStatus id="54E1885BFF88EA2DFE25FAF4F0FDFAD5" box="[391,491,1377,1401]" pageId="2" pageNumber="278" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE536F9FF88EA2DFF65FA11F2ACF911" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="278">
|
||
This female differs from the male
|
||
<typeStatus id="54E1885BFF88EA2DFDC4FA11F3DEFA31" box="[614,712,1412,1437]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
|
||
in the following aspects: body sheen green, brighter than in
|
||
<typeStatus id="54E1885BFF88EA2DFF35FA32F1EBFA6C" box="[151,253,1447,1472]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
|
||
. Head with longer clypeal teeth and more protruding clypeo-genal angle. Pronotum smaller in relation to elytra, with four rounded feeble depressions on each side of disc. Metathorax feebly concave on disc, with only indications of anterior and posterior spiniform projections. Profemur less strong, protibiae lacking internal tooth, with reduced basal internal denticles, somewhat dilated in the apical 2/5, and with three external teeth in the apical 2/5, the middle tooth smaller than the others, apical teeth less developed than in male. Meso- and metafemur straighter and weaker than in male. Meso- and metatibia less curved than in male, internal and apical denticles absent. Internal angle of apex of metatibia not prolonged in spiniform process. Middle and hind tibial spurs much longer than in male. Meso- and metatarsi about 1/7 longer than in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3406572FF88EA2DFF65F952F508F815" pageId="2" pageNumber="278" type="materials_examined">
|
||
<paragraph id="8BE536F9FF88EA2DFF65F952F508F815" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="278">
|
||
<emphasis id="B92EEAEBFF88EA2DFF65F952F0E3F94C" bold="true" box="[199,501,1735,1760]" pageId="2" pageNumber="278">
|
||
<typeStatus id="54E1885BFF88EA2DFF65F952F014F94C" box="[199,258,1735,1760]" pageId="2" pageNumber="278">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54E1885BFF88EA2DFEDDF952F0E6F94C" box="[383,496,1735,1760]" pageId="2" pageNumber="278" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D5CFD70FF88EA2DFE5DF95DF33CF94C" box="[511,554,1736,1760]" pageId="2" pageNumber="278" type="generic">1 ♂</specimenCount>
|
||
(white label, handwritten).
|
||
<collectingCountry id="F34D7669FF88EA2DFCCBF95DF515F94C" box="[873,1027,1736,1760]" name="Colombia" pageId="2" pageNumber="278">COLOMBIA</collectingCountry>
|
||
:
|
||
<emphasis id="B92EEAEBFF88EA2DFBADF952F544F94C" bold="true" box="[1039,1106,1735,1760]" pageId="2" pageNumber="278">Huila</emphasis>
|
||
, Nat. Park Las Cuevas, 1,80 dm,
|
||
<date id="FFE41039FF88EA2DFF6FF979F030F8A9" box="[205,294,1772,1797]" pageId="2" pageNumber="278" value="1976-05-23">23.V.76</date>
|
||
, M. Cooper./ 2 (white label, handwritten). Parque Nacional Las Cuevas de los Guácharos, Cloud forest, Forest of Quercus humboldtii/ 3 (white label, printed in black).
|
||
<collectionCode id="ED4BAE3CFF88EA2DFC1BF885F51BF88B" box="[953,1037,1808,1831]" pageId="2" pageNumber="278">BMNH</collectionCode>
|
||
{E} 2004275 M. Cooper/ 4 (white label, printed in black).
|
||
<collectionCode id="ED4BAE3CFF88EA2DFE0DF8A0F315F8E0" box="[431,515,1845,1868]" pageId="2" pageNumber="278">BMNH</collectionCode>
|
||
(E) #943732 / 5 (red label, printed and handwritten in black).
|
||
<typeStatus id="54E1885BFF88EA2DFB40F8A1F460F8E0" box="[1250,1398,1844,1868]" pageId="2" pageNumber="278" type="holotype">HOLOTYPE</typeStatus>
|
||
♂,
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFF35F8C2F068F8DC" box="[151,382,1879,1904]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="gigante" status="sp. nov.">
|
||
<emphasis id="B92EEAEBFF88EA2DFF35F8C2F068F8DC" box="[151,382,1879,1904]" italics="true" pageId="2" pageNumber="278">Deltochilum gigante</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D5790FF88EA2DFE25F8C2F0F4F8DC" box="[391,482,1879,1904]" pageId="2" pageNumber="278" rank="species">
|
||
<emphasis id="B92EEAEBFF88EA2DFE25F8C2F0F4F8DC" bold="true" box="[391,482,1879,1904]" pageId="2" pageNumber="278">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Silva and Vaz-de-Mello, des. 2013 (
|
||
<collectionCode id="ED4BAE3CFF88EA2DFC2CF8CDF2FEF8DC" box="[910,1000,1880,1904]" pageId="2" pageNumber="278">BMNH</collectionCode>
|
||
).
|
||
<emphasis id="B92EEAEBFF88EA2DFC5FF8C2F565F8DC" bold="true" box="[1021,1139,1879,1904]" pageId="2" pageNumber="278">
|
||
<typeStatus id="54E1885BFF88EA2DFC5FF8C2F578F8DC" box="[1021,1134,1879,1904]" pageId="2" pageNumber="278" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
1 ♀, no locality label, no abdomen (
|
||
<collectionCode id="ED4BAE3CFF88EA2DFEB2F8E9F07AF838" LSID="urn:lsid:biocol.org:col:34988" box="[272,364,1916,1940]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="278">MNHN</collectionCode>
|
||
). 1 (yellow label, printed in black).
|
||
<typeStatus id="54E1885BFF88EA2DFCB1F8E9F2B4F838" box="[787,930,1916,1940]" pageId="2" pageNumber="278" type="paratype">PARATYPE</typeStatus>
|
||
./ 2 (white label, printed and handwritten in black)
|
||
<taxonomicName id="4C5A4D7AFF88EA2DFF46F835F0DFF814" box="[228,457,1952,1977]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="gigante" status="sp. nov.">
|
||
<emphasis id="B92EEAEBFF88EA2DFF46F835F0DFF814" box="[228,457,1952,1977]" italics="true" pageId="2" pageNumber="278">Deltochilum gigante</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D5790FF88EA2DFE72F835F33CF815" box="[464,554,1952,1977]" pageId="2" pageNumber="278" rank="species">
|
||
<emphasis id="B92EEAEBFF88EA2DFE72F835F33CF815" bold="true" box="[464,554,1952,1977]" pageId="2" pageNumber="278">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Silva and Vaz-de-Mello, des. 2013
|
||
<collectionCode id="ED4BAE3CFF88EA2DFC60F834F50CF815" box="[962,1050,1953,1977]" pageId="2" pageNumber="278">PARAT</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |