treatments-xml/data/03/F3/87/03F387C57A25FFB3FC0C07CA34DDB2E9.xml
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<document id="C4963015C0C8361F84ECD668B0981931" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A25FFB3FC0C07CA34DDB2E9" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria ceylonensis Hebauer 2001" docType="treatment" docVersion="9" lastPageNumber="242" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="240" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="3800D72B7C9B889FAED7A6326BE2520E">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:namePart id="E21382267E61E295C2341C7444173A5E">Fikáček, Martin</mods:namePart>
<mods:affiliation id="596A59929952B21D03349CF4BF8E13C8">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="BA72FE7B968BED3141616C562677EAC9" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3F7006E0C0E405F93EDDA2558D196D5C">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="0D8B630CF39632215A78806D1F96608F">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<treatment id="03F387C57A25FFB3FC0C07CA34DDB2E9" ID-DOI="http://doi.org/10.5281/zenodo.4548831" ID-GBIF-Taxon="177423423" ID-Zenodo-Dep="4548831" LSID="urn:lsid:plazi:treatment:03F387C57A25FFB3FC0C07CA34DDB2E9" httpUri="http://treatment.plazi.org/id/03F387C57A25FFB3FC0C07CA34DDB2E9" lastPageId="13" lastPageNumber="242" pageId="11" pageNumber="240">
<subSubSection id="C34065587A25FFB5FC0C07CA3163B1F0" box="[901,1362,1762,1785]" pageId="11" pageNumber="240" type="nomenclature">
<paragraph id="8BE536D37A25FFB5FC0C07CA3163B1F0" blockId="11.[901,1362,1762,1785]" box="[901,1362,1762,1785]" pageId="11" pageNumber="240">
<heading id="D0AD81BF7A25FFB5FC0C07CA3163B1F0" box="[901,1362,1762,1785]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="240" reason="3">
<taxonomicName id="4C5A4D507A25FFB5FC0C07CA3163B1F0" ID-CoL="56TH3" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[901,1362,1762,1785]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="240" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B92EEAC17A25FFB5FC0C07CA3163B1F0" bold="true" box="[901,1362,1762,1785]" pageId="11" pageNumber="240">
<emphasis id="B92EEAC17A25FFB5FC0C07CA3099B1F0" bold="true" box="[901,1192,1762,1785]" italics="true" pageId="11" pageNumber="240">Thysanarthria ceylonensis</emphasis>
<bibRefCitation id="EFCB4B227A25FFB5FB2607CA3163B1F0" author="Hebauer" box="[1199,1362,1762,1785]" firstAuthor="Hebauer" pageId="11" pageNumber="240" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C34065587A25FFB5FBAF06283083B01A" box="[1062,1202,1792,1811]" pageId="11" pageNumber="240" type="description">
<paragraph id="8BE536D37A25FFB5FBAF06283083B01A" blockId="11.[1062,1202,1792,1811]" box="[1062,1202,1792,1811]" pageId="11" pageNumber="240">
(
<figureCitation id="13612A567A25FFB5FBA506283050B01A" box="[1068,1121,1792,1811]" captionStart="Fig" captionStartId="12.[151,182,1980,1999]" captionTargetBox="[155,1405,208,1868]" captionTargetId="figure-13@12.[153,1435,189,1873]" captionTargetPageId="12" captionText="Fig. 6. Male genitalia of Thysanarthria species, holotypes (AI, KO) and paratype (J). AE T.bifida sp. nov. (AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. trifida sp. nov. (FH whole aedeagus of the holotype in dorsal, lateral and ventral view; I details of parameres of the holotype; J same of the paratype). KO T. ceylonensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488934" httpUri="https://zenodo.org/record/4488934/files/figure.png" pageId="11" pageNumber="240">Figs 6</figureCitation>
KO,
<figureCitation id="13612A567A25FFB5FB1F0628309AB01A" box="[1174,1195,1792,1811]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (AB) and Near East (C), and examples of habitats of Thysanarthria (DF). Localities: D India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; FG Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, FG by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="11" pageNumber="240">11</figureCitation>
)
</paragraph>
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<paragraph id="8BE536D37A25FFB5FCB5060230FAB034" blockId="11.[828,1438,1834,2081]" box="[828,1227,1834,1853]" pageId="11" pageNumber="240">
<treatmentCitationGroup id="AB4A11FD7A25FFB5FCB5060230FAB034" box="[828,1227,1834,1853]" pageId="11" pageNumber="240">
<treatmentCitation id="0AFB10C27A25FFB5FCB5060230FAB034" author="HEBAUER F." box="[828,1227,1834,1853]" page="396" pageId="11" pageNumber="240" year="2001">
<taxonomicName id="4C5A4D507A25FFB5FCB5060230FAB034" ID-CoL="56TH3" authority="Hebauer, 2001: 396" authorityName="Hebauer" authorityPageNumber="396" authorityYear="2001" box="[828,1227,1834,1853]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="240" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B92EEAC17A25FFB5FCB50602302CB034" box="[828,1053,1834,1853]" italics="true" pageId="11" pageNumber="240">Thysanarthria ceylonensis</emphasis>
<bibRefCitation id="EFCB4B227A25FFB5FBAA060230FAB034" author="Hebauer" box="[1059,1227,1834,1853]" firstAuthor="Hebauer" page="396" pageId="11" pageNumber="240" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001: 396</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="C34065587A25FFB3FCB5067B35BEB678" lastPageId="13" lastPageNumber="242" pageId="11" pageNumber="240" type="materials_examined">
<paragraph id="8BE536D37A25FFB5FCB5067B31B9B0D8" blockId="11.[828,1438,1834,2081]" pageId="11" pageNumber="240">
<emphasis id="B92EEAC17A25FFB5FCB5067B3001B06F" bold="true" box="[828,1072,1875,1894]" pageId="11" pageNumber="240">Type material examined.</emphasis>
<materialsCitation id="3B323C8E7A25FFB5FBB3067B31B9B0D8" collectingDate="1962-02-13" collectionCode="MZLU" country="Sri Lanka" location="Kudattanai" municipality="Kudattanai" pageId="11" pageNumber="240" specimenCount="1" specimenCount-male="1" stateProvince="NORTHERN PROVINCE" typeStatus="holotype">
<typeStatus id="54E188717A25FFB5FBB3067B30A3B06C" box="[1082,1170,1875,1894]" pageId="11" pageNumber="240" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5CFD5A7A25FFB5FB2B06653083B06F" box="[1186,1202,1869,1894]" pageId="11" pageNumber="240" type="male"></specimenCount>
(
<collectionCode id="ED4BAE167A25FFB5FB49067B3135B06F" box="[1216,1284,1875,1894]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="11" pageNumber="240">MZLU</collectionCode>
),
<emphasis id="B92EEAC17A25FFB5FA9E067B37C6B088" bold="true" pageId="11" pageNumber="240">
<collectingCountry id="F34D76437A25FFB5FA9E067B31A9B06F" box="[1303,1432,1875,1894]" name="Sri Lanka" pageId="11" pageNumber="240">SRI LANKA</collectingCountry>
:
<collectingRegion id="499EF8317A25FFB5FCB5064637C0B076" box="[828,1009,1902,1921]" pageId="11" pageNumber="240">NORTHERN PROVINCE</collectingRegion>
:
</emphasis>
ʻCeylon, N. Prov. /
<collectingMunicipality id="6B81ACA97A25FFB5FB2006463136B088" box="[1193,1287,1902,1921]" pageId="11" pageNumber="240">Kudattanai</collectingMunicipality>
/
<locationDeviation id="9A572FF17A25FFB5FA900646375FB095" pageId="11" pageNumber="240">6 mls SE Point Pedro</locationDeviation>
/ 13.II.62, Loc. 70 // At pond in semi- / desert // Lund University / Ceylon Expedition 1962 / Brinck-Andersson- / Cederholm // MZLU / Type no. / 3120:1 // Photo 2017 / by MZLU // MZLU / 2017 / 511.
</materialsCitation>
</paragraph>
<paragraph id="8BE536D37A25FFB3FCB506F135BEB678" blockId="11.[828,1438,1834,2081]" lastBlockId="13.[151,761,190,369]" lastPageId="13" lastPageNumber="242" pageId="11" pageNumber="240">
<emphasis id="B92EEAC17A25FFB5FCB506F13061B0E5" bold="true" box="[828,1104,2009,2028]" pageId="11" pageNumber="240">Additional material examined.</emphasis>
<materialsCitation id="3B323C8E7A25FFB5FBDA06F1302DBF0F" ID-GBIF-Occurrence="3756360353" pageId="11" pageNumber="240">
<emphasis id="B92EEAC17A25FFB5FBDA06F1310CB0E5" bold="true" box="[1107,1341,2009,2028]" pageId="11" pageNumber="240">
<collectingCountry id="F34D76437A25FFB5FBDA06F130A4B0E5" box="[1107,1173,2009,2028]" name="India" pageId="11" pageNumber="240">INDIA</collectingCountry>
:
<collectingRegion id="499EF8317A25FFB5FB1706F130D6B0E3" box="[1182,1255,2009,2028]" country="India" name="Madhya Pradesh" pageId="11" pageNumber="240">MADHYA</collectingRegion>
PRADESH:
</emphasis>
1 ♂,
<specimenCount id="9D5CFD5A7A25FFB5FAE306F13749BF0F" pageId="11" pageNumber="240" type="generic">2 specimens</specimenCount>
(
<collectionCode id="ED4BAE167A25FFB5FC0B06DB37FBBF0F" box="[898,970,2035,2054]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="240" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED4BAE167A25FFB5FC5806DB3023BF0F" box="[977,1042,2035,2054]" country="Czech Republic" name="National Museum Prague" pageId="11" pageNumber="240">NMPC</collectionCode>
):
</materialsCitation>
<materialsCitation id="3B323C8E7A25FFB5FB9606DB3129BF28" ID-GBIF-Occurrence="3756360330" pageId="11" pageNumber="240">
River Denwa, ca.
<quantity id="4CA29B367A25FFB5FB3F06DB30D1BF0F" box="[1206,1248,2035,2054]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="240" unit="km" value="8.0">8 km</quantity>
SSE Matkuli, Satpura Range,
<quantity id="4CA29B367A25FFB5FCF409263784BF28" box="[893,949,2062,2081]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="240" unit="m" value="400.0">400 m</quantity>
,
<date id="FFE410137A25FFB5FC3709263027BF28" box="[958,1046,2062,2081]" pageId="11" pageNumber="240" value="2008-02-28">28.ii.2008</date>
, lgt. M. Jäch, S. &amp; P. Sharma;
</materialsCitation>
<materialsCitation id="3B323C8E7A25FFB3FA940926353FB60F" ID-GBIF-Occurrence="3756360319" lastPageId="13" lastPageNumber="242" pageId="11" pageNumber="240">
<specimenCount id="9D5CFD5A7A25FFB5FA940926310CBF28" box="[1309,1341,2058,2081]" pageId="11" pageNumber="240" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4BAE167A25FFB5FAC1092631A5BF28" box="[1352,1428,2062,2081]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="240" type="Museum">NHMW</collectionCode>
): Hoshangabad Distr., Bandrabhan, ca.
<quantity id="4CA29B367A23FFB3FE5501963623B7D8" box="[476,530,190,209]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="242" unit="km" value="60.0">60 km</quantity>
SSE Bhopal, ca.
<quantity id="4CA29B367A23FFB3FD23019636E4B7D8" box="[682,725,190,209]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="242" unit="km" value="5.0">5 km</quantity>
NE Hoshangabad, River Narmada,
<quantity id="4CA29B367A23FFB3FE2101F135D0B7E5" box="[424,481,217,236]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="242" unit="m" value="280.0">280 m</quantity>
,
<date id="FFE410137A23FFB3FE6201F13658B7E5" box="[491,617,217,236]" pageId="13" pageNumber="242" value="2008-02-23" valueMax="2008-02-24" valueMin="2008-02-23">23.24.ii.2008</date>
, lgt. M. Jäch, S. &amp; P. Sharma;
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FE9C01DB3559B635" ID-GBIF-Occurrence="3756360331" pageId="13" pageNumber="242">
<specimenCount id="9D5CFD5A7A23FFB3FE9C01DB3565B60F" box="[277,340,243,262]" pageId="13" pageNumber="242" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FEE101DB3585B60F" box="[360,436,243,262]" pageId="13" pageNumber="242">NHWM</collectionCode>
): Chhindwara Distr., Bhadhua Chora (stream), ca.
<quantity id="4CA29B367A23FFB3FE810026350EB628" box="[264,319,270,289]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="242" unit="km" value="10.0">10 km</quantity>
E Matkuli near Mahul Jhir,
<quantity id="4CA29B367A23FFB3FDBE00263641B628" box="[567,624,270,289]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="242" unit="m" value="400.0">400 m</quantity>
,
<date id="FFE410137A23FFB3FDF3002636E3B628" box="[634,722,270,289]" pageId="13" pageNumber="242" value="2008-02-28">28.ii.2008</date>
, lgt. M. Jäch, S. &amp; P.Sharma;
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FEE5000135BEB678" ID-GBIF-Occurrence="3756360344" pageId="13" pageNumber="242">
<specimenCount id="9D5CFD5A7A23FFB3FEE5000135B8B635" box="[364,393,293,316]" pageId="13" pageNumber="242" type="female">1♀</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FE1D000135D1B635" box="[404,480,297,316]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="242" type="Museum">NHMW</collectionCode>
): Hoshangabad Distr., Dhobighat Nala (stream), ca.
<quantity id="4CA29B367A23FFB3FEA4006B3567B65F" box="[301,342,323,342]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="242" unit="km" value="2.0">2 km</quantity>
SE Pachmarhi,Saphura Range,
<quantity id="4CA29B367A23FFB3FDE8006B36A7B65F" box="[609,662,323,342]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="242" unit="m" value="900.0">900 m</quantity>
,
<date id="FFE410137A23FFB3FD14006B36C4B65F" box="[669,757,323,342]" pageId="13" pageNumber="242" value="2008-02-27">27.ii.2008</date>
, lgt. M. Jäch, S. &amp; P. Sharma.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF25665B7A22FFB2FF1E069430B0BF16" ID-DOI="http://doi.org/10.5281/zenodo.4488934" ID-Zenodo-Dep="4488934" httpUri="https://zenodo.org/record/4488934/files/figure.png" pageId="12" pageNumber="241" startId="12.[151,182,1980,1999]" targetBox="[155,1405,208,1868]" targetPageId="12">
<paragraph id="8BE536D37A22FFB2FF1E069430B0BF16" blockId="12.[151,1436,1980,2079]" pageId="12" pageNumber="241">
Fig. 6. Male genitalia of
<taxonomicName id="4C5A4D507A22FFB2FEEE069435D0B0C6" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[359,481,1980,1999]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A22FFB2FEEE069435D0B0C6" box="[359,481,1980,1999]" italics="true" pageId="12" pageNumber="241">Thysanarthria</emphasis>
</taxonomicName>
species, holotypes (AI, KO) and paratype (J). AE
<taxonomicName id="4C5A4D507A22FFB2FC3E069437CAB0C6" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[951,1019,1980,1999]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="241" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="B92EEAC17A22FFB2FC3E069437CAB0C6" box="[951,1019,1980,1999]" italics="true" pageId="12" pageNumber="241">T.bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A22FFB2FB8906943073B0C6" box="[1024,1090,1980,1999]" pageId="12" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ
<taxonomicName id="4C5A4D507A22FFB2FC3706FF3037B0E3" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[958,1030,2007,2026]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="241" phylum="Arthropoda" rank="species" species="trifida" status="sp. nov.">
<emphasis id="B92EEAC17A22FFB2FC3706FF3037B0E3" box="[958,1030,2007,2026]" italics="true" pageId="12" pageNumber="241">T. trifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A22FFB2FB8206FF307CB0E3" box="[1035,1101,2007,2026]" pageId="12" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
(FH whole aedeagus of the holotype in dorsal, lateral and ventral view; I details of parameres of the holotype; J same of the paratype). KO
<taxonomicName id="4C5A4D507A22FFB2FBAC06D9312DBF0D" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[1061,1308,2033,2052]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="241" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B92EEAC17A22FFB2FBAC06D930AABF0D" box="[1061,1179,2033,2052]" italics="true" pageId="12" pageNumber="241">T. ceylonensis</emphasis>
<bibRefCitation id="EFCB4B227A22FFB2FB2906D9312DBF0D" author="Hebauer" box="[1184,1308,2033,2052]" firstAuthor="Hebauer" pageId="12" pageNumber="241" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</taxonomicName>
(KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view).
</paragraph>
</caption>
<subSubSection id="C34065587A23FFB3FF1E00A135DFB349" pageId="13" pageNumber="242" type="description">
<paragraph id="8BE536D37A23FFB3FF1E00A135DFB349" blockId="13.[151,760,393,1504]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF1E00A135A0B6A9" bold="true" box="[151,401,393,416]" pageId="13" pageNumber="242">
Redescription.
<emphasis id="B92EEAC17A23FFB3FEDF00A135A0B6A9" bold="true" box="[342,401,393,416]" italics="true" pageId="13" pageNumber="242">Body</emphasis>
</emphasis>
length
<quantity id="4CA29B367A23FFB3FE7800A1364FB6A9" box="[497,638,393,416]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="13" pageNumber="242" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
(
<typeStatus id="54E188717A23FFB3FD1800A136C6B6A9" box="[657,759,393,416]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
<quantity id="4CA29B367A23FFB3FF1E008134C1B6C9" box="[151,240,425,448]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="242" unit="mm" value="1.5">1.5 mm</quantity>
), maximum body width 0.91.0 mm (
<typeStatus id="54E188717A23FFB3FD11008136C6B6C9" box="[664,759,425,448]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
<quantity id="4CA29B367A23FFB3FF1E00E134DDB6E9" box="[151,236,457,480]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="242" unit="mm" value="0.9">0.9 mm</quantity>
). Head and labrum black and pronotum uniformly yellowish; elytra yellowish with slightly darker lateral parts; legs reddish to yellowish.
<emphasis id="B92EEAC17A23FFB3FE790321361AB529" bold="true" box="[496,555,521,544]" italics="true" pageId="13" pageNumber="242">Head</emphasis>
with strong microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.5× the width of one eye in dorsal view.
<emphasis id="B92EEAC17A23FFB3FE1103413636B589" bold="true" box="[408,519,617,640]" italics="true" pageId="13" pageNumber="242">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices without microsculpture.
<emphasis id="B92EEAC17A23FFB3FECE038135BAB5C9" bold="true" box="[327,395,681,704]" italics="true" pageId="13" pageNumber="242">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
<emphasis id="B92EEAC17A23FFB3FD04020136C9B449" bold="true" box="[653,760,809,832]" italics="true" pageId="13" pageNumber="242">Aedeagus</emphasis>
(
<figureCitation id="13612A567A23FFB3FF16026134D4B469" box="[159,229,841,864]" captionStart="Fig" captionStartId="12.[151,182,1980,1999]" captionTargetBox="[155,1405,208,1868]" captionTargetId="figure-13@12.[153,1435,189,1873]" captionTargetPageId="12" captionText="Fig. 6. Male genitalia of Thysanarthria species, holotypes (AI, KO) and paratype (J). AE T.bifida sp. nov. (AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. trifida sp. nov. (FH whole aedeagus of the holotype in dorsal, lateral and ventral view; I details of parameres of the holotype; J same of the paratype). KO T. ceylonensis Hebauer, 2001 (KM whole aedeagus in dorsal, lateral and ventral view; NO details of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488934" httpUri="https://zenodo.org/record/4488934/files/figure.png" pageId="13" pageNumber="242">Figs 6</figureCitation>
KO)
<quantity id="4CA29B367A23FFB3FEA5026135B3B469" box="[300,386,841,864]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="242" unit="mm" value="0.4">0.4 mm</quantity>
long. Phallobase not much wider at base of parameres than more basally, only indistinctly narrowed at midlength; arcuate in lateral view. Paremere moderately wide basally, slightly narrowing towards apex, apex widely angulate. Median lobe widely bottle-shaped, rounded apically, without paired subapical projections; apex not reaching level of parameral apices; gonopore rounded, situated far from apex.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FF1E056136C9B3E9" pageId="13" pageNumber="242" type="diagnosis">
<paragraph id="8BE536D37A23FFB3FF1E056136C9B3E9" blockId="13.[151,760,393,1504]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF1E056135BEB369" bold="true" box="[151,399,1097,1120]" pageId="13" pageNumber="242">Differential diagnosis.</emphasis>
<taxonomicName id="4C5A4D507A23FFB3FE1A0561369BB369" authorityName="Hebauer" authorityYear="2001" box="[403,682,1097,1120]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B92EEAC17A23FFB3FE1A0561369BB369" box="[403,682,1097,1120]" italics="true" pageId="13" pageNumber="242">Thysanarthria ceylonensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C5A4D507A23FFB3FF4E0541356FB389" authorityName="Hebauer" authorityYear="2001" box="[199,350,1129,1152]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B92EEAC17A23FFB3FF4E0541356FB389" box="[199,350,1129,1152]" italics="true" pageId="13" pageNumber="242">T. bengalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A4D507A23FFB3FEE3054135FFB389" authorityName="Hebauer" authorityYear="2001" box="[362,462,1129,1152]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="brincki">
<emphasis id="B92EEAC17A23FFB3FEE3054135FFB389" box="[362,462,1129,1152]" italics="true" pageId="13" pageNumber="242">T. brincki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D507A23FFB3FE76054136A1B389" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[511,656,1129,1152]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="saurahana">
<emphasis id="B92EEAC17A23FFB3FE76054136A1B389" box="[511,656,1129,1152]" italics="true" pageId="13" pageNumber="242">H. saurahana</emphasis>
</taxonomicName>
in having strong mesh-like microsculpture on the head, but it can be easily distinguished from all these species as well as from all other
<taxonomicName id="4C5A4D507A23FFB3FF7C05E135BFB3E9" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[245,398,1225,1248]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A23FFB3FF7C05E135BFB3E9" box="[245,398,1225,1248]" italics="true" pageId="13" pageNumber="242">Thysanarthria</emphasis>
</taxonomicName>
by a very characteristic aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FF1E05C13556B269" pageId="13" pageNumber="242" type="biology_ecology">
<paragraph id="8BE536D37A23FFB3FF1E05C13556B269" blockId="13.[151,760,393,1504]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF1E05C134C2B209" bold="true" box="[151,243,1257,1280]" pageId="13" pageNumber="242">Biology.</emphasis>
Specimens from
<collectingRegion id="499EF8317A23FFB3FE2005C13668B209" box="[425,601,1257,1280]" country="India" name="Madhya Pradesh" pageId="13" pageNumber="242">Madhya Pradesh</collectingRegion>
were collected at the sides of small to large rivers with stony banks (e.g.,
<figureCitation id="13612A567A23FFB3FF1E040134C9B249" box="[151,248,1321,1344]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (AB) and Near East (C), and examples of habitats of Thysanarthria (DF). Localities: D India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; FG Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, FG by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="13" pageNumber="242">Fig. 11D</figureCitation>
) but precise microhabitat is not known (M. A. Jäch, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FF1E044134DDB2E9" pageId="13" pageNumber="242" type="distribution">
<paragraph id="8BE536D37A23FFB3FF1E044134DDB2E9" blockId="13.[151,760,393,1504]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF1E0441351BB289" bold="true" box="[151,298,1385,1408]" pageId="13" pageNumber="242">Distribution.</emphasis>
Described from
<collectingCountry id="F34D76437A23FFB3FE5404413677B289" box="[477,582,1385,1408]" name="Sri Lanka" pageId="13" pageNumber="242">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFCB4B227A23FFB3FDDD044136C1B289" author="HEBAUER" box="[596,752,1385,1408]" firstAuthor="HEBAUER" pageId="13" pageNumber="242" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER 2001</bibRefCitation>
) but here recorded from central
<collectingCountry id="F34D76437A23FFB3FE6504A13612B2A9" box="[492,547,1417,1440]" name="India" pageId="13" pageNumber="242">India</collectingCountry>
(
<collectingRegion id="499EF8317A23FFB3FDBB04A136DCB2A9" box="[562,749,1417,1440]" country="India" name="Madhya Pradesh" pageId="13" pageNumber="242">Madhya Pradesh</collectingRegion>
), hence the species is likely widespread in the Indian Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>