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<document id="AFB2A05D2E6948D85C77DCA473DC059A" ID-CLB-Dataset="5458" ID-DOI="10.11646/zootaxa.353.1.1" ID-GBIF-Dataset="740af13a-ed5c-40d0-b0c1-8d3fd2b92f56" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019783" checkinTime="1624467278120" checkinUser="felipe" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="03F387B2FFD2531FCD24524EFB70A161" docLanguage="en" docName="zt00353.pdf" docOrigin="Zootaxa 353 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Boreophilia angusticornis Gusarov 2003, comb. nov." docType="treatment" docVersion="9" lastPageNumber="84" masterDocId="FFCAFFCAFF80534CCC2C5352FFD8A23F" masterDocTitle="Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes" masterLastPageNumber="134" masterPageNumber="1" pageNumber="83" updateTime="1698204243681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="20938B0D4B7D1DAE3BBF6DB7B742811F">Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes</mods:title>
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<mods:namePart id="921E387DBA51AC3F39C946866222F52B">Gusarov, Vladimir I.</mods:namePart>
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<treatment id="03F387B2FFD2531FCD24524EFB70A161" ID-DOI="http://doi.org/10.5281/zenodo.5020243" ID-GBIF-Taxon="183542837" ID-Zenodo-Dep="5020243" LSID="urn:lsid:plazi:treatment:03F387B2FFD2531FCD24524EFB70A161" httpUri="http://treatment.plazi.org/id/03F387B2FFD2531FCD24524EFB70A161" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<subSubSection id="C340652FFFD2531ECD24524EFC18A30A" box="[264,960,283,310]" pageId="82" pageNumber="83" type="nomenclature">
<paragraph id="8BE536A4FFD2531ECD24524EFC18A30A" blockId="82.[264,960,283,345]" box="[264,960,283,310]" pageId="82" pageNumber="83">
<heading id="D0AD81C8FFD2531ECD24524EFC18A30A" bold="true" box="[264,960,283,310]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
<emphasis id="B92EEAB6FFD2531ECD24524EFC18A30A" bold="true" box="[264,960,283,310]" pageId="82" pageNumber="83">
<taxonomicName id="4C5A4D27FFD2531ECD24524EFCF4A30A" authority="(Bernhauer, 1907)" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[264,812,283,310]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis" status="comb. nov.">
<emphasis id="B92EEAB6FFD2531ECD24524EFD9AA309" bold="true" box="[264,578,284,310]" italics="true" pageId="82" pageNumber="83">Boreophilia angusticornis</emphasis>
(
<bibRefCitation id="EFCB4B55FFD2531ECE7E5249FCFCA30A" author="Bernhauer, M." box="[594,804,283,309]" pageId="82" pageNumber="83" pagination="381 - 405" refId="ref76008" refString="Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A21D57CDFFD2531ECF165249FC18A30A" box="[826,960,283,309]" pageId="82" pageNumber="83" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD2531ECD24526DFE19A366" box="[264,449,319,345]" pageId="82" pageNumber="83" type="description">
<paragraph id="8BE536A4FFD2531ECD24526DFE19A366" blockId="82.[264,960,283,345]" box="[264,449,319,345]" pageId="82" pageNumber="83">
(
<figureCitation id="13612A21FFD2531ECD3D526DFE60A366" box="[273,440,319,345]" captionStart-0="FIGURES 116119" captionStart-1="FIGURES 120124" captionStartId-0="81.[264,383,1731,1755]" captionStartId-1="82.[264,383,1727,1751]" captionTargetBox-0="[346,1237,974,1699]" captionTargetBox-1="[286,1306,861,1691]" captionTargetId-0="figure-253@81.[341,1245,964,1705]" captionTargetId-1="figure-191@82.[272,1315,852,1701]" captionTargetPageId-0="81" captionTargetPageId-1="82" captionText-0="FIGURES 116119. Abdominal segment 8 of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (116117); and female, lectotype of Metaxya plutonica Casey (118119)). 116 — male tergum 8; 117 — male sternum 8; 118 — female tergum 8; 119 — female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 120124. Genitalia of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (120123); and female, lectotype of Metaxya plutonica Casey (124)). 120 — median lobe, parameral view; 121 — apex of median lobe with retracted internal sac, abparameral view; 122 — median lobe, lateral view; 123 — apex of median lobe with retracted internal sac, lateral view; 124 — spermatheca. Scale bar 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5019864" figureDoi-1="http://doi.org/10.5281/zenodo.5019870" httpUri-0="https://zenodo.org/record/5019864/files/figure.png" httpUri-1="https://zenodo.org/record/5019870/files/figure.png" pageId="82" pageNumber="83">Figs. 116124</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD2531ECD2452DDFB4CA03A" pageId="82" pageNumber="83" type="reference_group">
<paragraph id="8BE536A4FFD2531ECD2452DDFC8EA39A" blockId="82.[264,1218,398,518]" box="[264,854,398,422]" pageId="82" pageNumber="83">
<treatmentCitationGroup id="AB4A118AFFD2531ECD2452DDFC8EA39A" box="[264,854,398,422]" pageId="82" pageNumber="83">
<treatmentCitation id="0AFB10B5FFD2531ECD2452DDFC89A39A" author="Bernhauer, M." box="[264,849,398,422]" page="384" pageId="82" pageNumber="83" year="1907">
<taxonomicName id="4C5A4D27FFD2531ECD2452DDFC89A39A" authority="Bernhauer, 1907: 384" authorityName="Bernhauer" authorityPageNumber="384" authorityYear="1907" box="[264,849,398,422]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis" subGenus="Metaxya">
<emphasis id="B92EEAB6FFD2531ECD2452DDFE96A399" box="[264,334,399,422]" italics="true" pageId="82" pageNumber="83">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFD2531ECD7152DDFE61A399" box="[349,441,399,422]" italics="true" pageId="82" pageNumber="83">Metaxya</emphasis>
)
<emphasis id="B92EEAB6FFD2531ECDE552DDFD82A399" box="[457,602,399,422]" italics="true" pageId="82" pageNumber="83">angusticornis</emphasis>
<bibRefCitation id="EFCB4B55FFD2531ECE4C52DCFC89A39A" author="Bernhauer, M." box="[608,849,398,421]" pageId="82" pageNumber="83" pagination="381 - 405" refId="ref76008" refString="Bernhauer, M. (1907) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405." type="journal article" year="1907">Bernhauer, 1907: 384</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFD2531ECD2452FDFCDBA3F9" blockId="82.[264,1218,398,518]" box="[264,771,430,454]" pageId="82" pageNumber="83">
<treatmentCitationGroup id="AB4A118AFFD2531ECD2452FDFCDBA3F9" box="[264,771,430,454]" pageId="82" pageNumber="83">
<treatmentCitation id="0AFB10B5FFD2531ECD2452FDFD4DA3FA" author="Casey, T. L." box="[264,661,430,454]" page="82" pageId="82" pageNumber="83" year="1910">
<taxonomicName id="4C5A4D27FFD2531ECD2452FDFD4DA3FA" ID-CoL="6RDRR" authority="Casey, 1910a: 82" authorityName="Casey" authorityPageNumber="82" authorityYear="1910" box="[264,661,430,454]" class="Insecta" family="Staphylinidae" genus="Metaxya" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="plutonica" status="syn. nov.">
<emphasis id="B92EEAB6FFD2531ECD2452FDFE17A3F9" box="[264,463,431,454]" italics="true" pageId="82" pageNumber="83">Metaxya plutonica</emphasis>
<bibRefCitation id="EFCB4B55FFD2531ECDFA52FCFD4DA3FA" author="Casey, T. L." box="[470,661,430,453]" pageId="82" pageNumber="83" pagination="1 - 183" refId="ref76473" refString="Casey, T. L. (1910 a) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910a: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B92EEAB6FFD2531ECE8D52FCFCDBA3F9" bold="true" box="[673,771,430,454]" pageId="82" pageNumber="83">
<taxonomicNameLabel id="A21D57CDFFD2531ECE8D52FCFCDBA3F9" box="[673,771,430,454]" pageId="82" pageNumber="83">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFD2531ECD24529DFB1AA3DA" blockId="82.[264,1218,398,518]" box="[264,1218,462,486]" pageId="82" pageNumber="83">
<treatmentCitationGroup id="AB4A118AFFD2531ECD24529DFB1AA3DA" box="[264,1218,462,486]" pageId="82" pageNumber="83">
<treatmentCitation id="0AFB10B5FFD2531ECD24529DFC24A3DA" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[264,1020,462,486]" page="612" pageId="82" pageNumber="83" year="1926">
<taxonomicName id="4C5A4D27FFD2531ECD24529DFC24A3DA" authority=": Bernhauer &amp; Scheerpeltz, 1926: 612" authorityName=": Bernhauer &amp; Scheerpeltz" authorityPageNumber="612" authorityYear="1926" box="[264,1020,462,486]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis" subGenus="Metaxya">
<emphasis id="B92EEAB6FFD2531ECD24529DFE96A3D9" box="[264,334,463,486]" italics="true" pageId="82" pageNumber="83">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFD2531ECD71529DFE61A3D9" box="[349,441,463,486]" italics="true" pageId="82" pageNumber="83">Metaxya</emphasis>
)
<emphasis id="B92EEAB6FFD2531ECDE5529DFD82A3D9" box="[457,602,463,486]" italics="true" pageId="82" pageNumber="83">angusticornis</emphasis>
:
<bibRefCitation id="EFCB4B55FFD2531ECE42529CFC24A3DA" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[622,1020,462,485]" pageId="82" pageNumber="83" pagination="499 - 988" refId="ref76079" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz, 1926: 612</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536A4FFD2531ECD2452BDFB4CA03A" blockId="82.[264,1218,398,518]" box="[264,1172,494,518]" pageId="82" pageNumber="83">
<treatmentCitationGroup id="AB4A118AFFD2531ECD2452BDFB4CA03A" box="[264,1172,494,518]" pageId="82" pageNumber="83">
<treatmentCitation id="0AFB10B5FFD2531ECD2452BDFC16A03A" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[264,974,494,518]" page="616" pageId="82" pageNumber="83" year="1926">
<taxonomicName id="4C5A4D27FFD2531ECD2452BDFC16A03A" authority=": Bernhauer &amp; Scheerpeltz, 1926: 616" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="616" authorityYear="1926" box="[264,974,494,518]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="plutonica" subGenus="Metaxya">
<emphasis id="B92EEAB6FFD2531ECD2452BDFE96A039" box="[264,334,495,518]" italics="true" pageId="82" pageNumber="83">Atheta</emphasis>
(
<emphasis id="B92EEAB6FFD2531ECD7152BDFE61A039" box="[349,441,495,518]" italics="true" pageId="82" pageNumber="83">Metaxya</emphasis>
)
<emphasis id="B92EEAB6FFD2531ECDE552BDFDF5A039" box="[457,557,495,518]" italics="true" pageId="82" pageNumber="83">plutonica</emphasis>
:
<bibRefCitation id="EFCB4B55FFD2531ECE6E52BCFC16A03A" author="Bernhauer, M. &amp; Scheerpeltz, O." box="[578,974,494,517]" pageId="82" pageNumber="83" pagination="499 - 988" refId="ref76079" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz, 1926: 616</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as valid species).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD2531FCD245164FCF1A3BE" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="materials_examined">
<paragraph id="8BE536A4FFD2531ECD245164FBE1A126" blockId="82.[264,1324,566,793]" pageId="82" pageNumber="83">
<emphasis id="B92EEAB6FFD2531ECD245164FE1AA06F" bold="true" box="[264,450,566,592]" pageId="82" pageNumber="83">Type material.</emphasis>
<materialsCitation id="3B323CF9FFD2531ECDF65165FD7FA0F6" ID-GBIF-Occurrence="3321919651" collectionCode="FMNH" collectorName="M. Bernhauer Collection" location="Mt. Washgtn" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1" stateProvince="Washington" typeStatus="lectotype">
<typeStatus id="54E18806FFD2531ECDF65165FD97A06E" box="[474,591,567,593]" pageId="82" pageNumber="83">Lectotype</typeStatus>
of
<taxonomicName id="4C5A4D27FFD2531ECE525165FCAEA06F" authorityName=": Bernhauer &amp; Scheerpeltz" authorityYear="1926" box="[638,886,567,592]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECE525165FCAEA06F" box="[638,886,567,592]" italics="true" pageId="82" pageNumber="83">Atheta angusticornis</emphasis>
</taxonomicName>
(here designated):
<specimenCount id="9D5CFD2DFFD2531EC85C5164FB5CA075" box="[1136,1156,566,586]" pageId="82" pageNumber="83" type="male"></specimenCount>
(with broken antennae), “
<location id="8E85607FFFD2531ECDB2510DFD9AA046" LSID="urn:lsid:plazi:treatment:03F387B2FFD2531FCD24524EFB70A161:8E85607FFFD2531ECDB2510DFD9AA046" box="[414,578,607,633]" name="Mt. Washgtn" pageId="82" pageNumber="83" stateProvince="Washington">Mt. Washgtn</location>
[
<collectingRegion id="499EF846FFD2531ECE76510DFD34A046" box="[602,748,607,633]" country="United States of America" name="Washington" pageId="82" pageNumber="83">Washington</collectingRegion>
] N.[ew] H.[ampshire]”, “July”, white circle, “
<taxonomicName id="4C5A4D27FFD2531ECD3951D5FDECA09E" authority="Brh. Typ." authorityName="Brh. Typ." box="[277,564,647,673]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECD3951D5FE6DA09F" box="[277,437,647,672]" italics="true" pageId="82" pageNumber="83">angusticornis</emphasis>
Brh. Typ.
</taxonomicName>
”, “
<taxonomicName id="4C5A4D27FFD2531ECE7651D5FCEFA09E" authority="Brh." authorityName="Bernhauer" authorityYear="1907" box="[602,823,647,673]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECE7651D5FD21A09F" box="[602,761,647,672]" italics="true" pageId="82" pageNumber="83">angusticornis</emphasis>
Brh.
</taxonomicName>
Typus” (yellow label), “Chicago NHMus
<collectorName id="26AF5372FFD2531ECD2451FDFDF1A0F6" box="[264,553,687,713]" pageId="82" pageNumber="83">M.Bernhauer Collection</collectorName>
” (
<collectionCode id="ED4BAE61FFD2531ECE6F51FDFD46A0F6" box="[579,670,687,713]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="82" pageNumber="83" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B323CF9FFD2531ECE9751FDFBECA126" ID-GBIF-Occurrence="3321919302" collectionCode="FMNH" collectorName="M. Bernhauer Collection" location="Mt. Washgtn" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Washington" typeStatus="paralectotype">
<typeStatus id="54E18806FFD2531ECE9751FDFC87A0F6" box="[699,863,687,713]" pageId="82" pageNumber="83">Paralectotype</typeStatus>
:
<specimenCount id="9D5CFD2DFFD2531ECF5E51FCFC5EA0FD" box="[882,902,686,706]" pageId="82" pageNumber="83" type="female"></specimenCount>
(with missing head and prothorax), “
<location id="8E85607FFFD2531ECD3F5185FE75A0CE" LSID="urn:lsid:plazi:treatment:03F387B2FFD2531FCD24524EFB70A161:8E85607FFFD2531ECD3F5185FE75A0CE" box="[275,429,727,753]" name="Mt. Washgtn" pageId="82" pageNumber="83" stateProvince="Washington">Mt. Washgtn</location>
[
<collectingRegion id="499EF846FFD2531ECD905185FD96A0CE" box="[444,590,727,753]" country="United States of America" name="Washington" pageId="82" pageNumber="83">Washington</collectingRegion>
] N.[ew] H.[ampshire] Fenyes”, “
<taxonomicName id="4C5A4D27FFD2531ECFF45185FAF9A0CE" authority="Brh. Cotypus" authorityName="Brh. Cotypus" box="[984,1313,727,753]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECFF45185FBA0A0CF" box="[984,1144,727,752]" italics="true" pageId="82" pageNumber="83">angusticornis</emphasis>
Brh. Cotypus
</taxonomicName>
” (yellow label), “Chicago NHMus
<collectorName id="26AF5372FFD2531ECEBB51ADFC6FA126" box="[663,951,767,793]" pageId="82" pageNumber="83">M.Bernhauer Collection</collectorName>
” (
<collectionCode id="ED4BAE61FFD2531ECFFD51ADFBF5A126" box="[977,1069,767,793]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="82" pageNumber="83" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF25662CFFD2531ECD2455EDFCE5A55E" ID-DOI="http://doi.org/10.5281/zenodo.5019870" ID-Zenodo-Dep="5019870" httpUri="https://zenodo.org/record/5019870/files/figure.png" pageId="82" pageNumber="83" startId="82.[264,383,1727,1751]" targetBox="[286,1306,861,1691]" targetPageId="82">
<paragraph id="8BE536A4FFD2531ECD2455EDFCE5A55E" blockId="82.[264,1323,1727,1889]" pageId="82" pageNumber="83">
<emphasis id="B92EEAB6FFD2531ECD2455EDFE28A4E8" bold="true" box="[264,496,1727,1751]" pageId="82" pageNumber="83">FIGURES 120124.</emphasis>
Genitalia of
<taxonomicName id="4C5A4D27FFD2531ECEB15592FB9FA4E9" authority="(Bernhauer)" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[669,1095,1727,1751]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECEB15592FC61A4E8" box="[669,953,1728,1751]" italics="true" pageId="82" pageNumber="83">Boreophilia angusticornis</emphasis>
(Bernhauer)
</taxonomicName>
(male, lectotype of
<taxonomicName id="4C5A4D27FFD2531ECD2455B0FE3DA4C6" box="[264,485,1762,1785]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD2531ECD2455B0FE3DA4C6" box="[264,485,1762,1785]" italics="true" pageId="82" pageNumber="83">Atheta angusticornis</emphasis>
</taxonomicName>
(120123); and female, lectotype of
<taxonomicName id="4C5A4D27FFD2531ECF5855B0FB5BA4C6" authority="Casey" authorityName="Casey" authorityYear="1910" box="[884,1155,1762,1785]" class="Polypodiopsida" family="Metaxyaceae" genus="Metaxya" kingdom="Plantae" order="Cyatheales" pageId="82" pageNumber="83" phylum="Tracheophyta" rank="species" species="plutonica">
<emphasis id="B92EEAB6FFD2531ECF5855B0FBE3A4C6" box="[884,1083,1762,1785]" italics="true" pageId="82" pageNumber="83">Metaxya plutonica</emphasis>
Casey
</taxonomicName>
(124)). 120 — median lobe, parameral view; 121 — apex of median lobe with retracted internal sac, abparameral view; 122 — median lobe, lateral view; 123 — apex of median lobe with retracted internal sac, lateral view; 124 — spermatheca. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BE536A4FFD3531FCD14524EFCF1A3BE" blockId="83.[264,1323,283,862]" pageId="83" pageNumber="84">
<materialsCitation id="3B323CF9FFD3531FCD14524EFCF1A3BE" ID-GBIF-Occurrence="3321919546" collectionCode="NMNH" location="Mt. Wn." municipality="Subalpine Mt." pageId="83" pageNumber="84" specimenCount="1" specimenCount-female="1" stateProvince="Washington" typeStatus="lectotype">
<typeStatus id="54E18806FFD3531FCD14524EFE75A309" box="[312,429,284,310]" pageId="83" pageNumber="84">Lectotype</typeStatus>
of
<taxonomicName id="4C5A4D27FFD3531FCDC2524EFD0FA30A" authorityName="Casey" authorityYear="1910" box="[494,727,284,309]" class="Polypodiopsida" family="Metaxyaceae" genus="Metaxya" kingdom="Plantae" order="Cyatheales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="plutonica">
<emphasis id="B92EEAB6FFD3531FCDC2524EFD0FA30A" box="[494,727,284,309]" italics="true" pageId="83" pageNumber="84">Metaxya plutonica</emphasis>
</taxonomicName>
(here designated):
<specimenCount id="9D5CFD2DFFD3531FCFD85249FBD0A310" box="[1012,1032,283,303]" pageId="83" pageNumber="84" type="female"></specimenCount>
, “
<collectingMunicipality id="6B81ACDEFFD3531FC802524EFB3CA309" box="[1070,1252,284,310]" pageId="83" pageNumber="84">
Subalpine
<location id="8E85607FFFD3531FC897524EFB3CA309" LSID="urn:lsid:plazi:treatment:03F387B2FFD2531FCD24524EFB70A161:8E85607FFFD3531FC897524EFB3CA309" box="[1211,1252,284,310]" municipality="Subalpine Mt." name="Mt. Wn." pageId="83" pageNumber="84" stateProvince="Washington">Mt.</location>
</collectingMunicipality>
Wn. [
<collectingRegion id="499EF846FFD3531FCD225213FE47A364" box="[270,415,321,347]" country="United States of America" name="Washington" pageId="83" pageNumber="84">Washington</collectingRegion>
] N.[ew] H.[ampshire] vii.7.1899”, “
<taxonomicName id="4C5A4D27FFD3531FCF7C5213FC27A364" authority="Csy." authorityName="Casey" authorityYear="1910" box="[848,1023,321,347]" class="Polypodiopsida" family="Metaxyaceae" genus="Metaxya" kingdom="Plantae" order="Cyatheales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="plutonica">
<emphasis id="B92EEAB6FFD3531FCF7C5213FC66A365" box="[848,958,321,346]" italics="true" pageId="83" pageNumber="84">plutonica</emphasis>
Csy.
</taxonomicName>
”, “TYPE USNM 39064” (red label), “CASEY bequest 1925” (
<collectionCode id="ED4BAE61FFD3531FCE905235FCC5A3BE" box="[700,797,359,385]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="83" pageNumber="84" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD3531FCD1452D9FD50A021" pageId="83" pageNumber="84" type="diagnosis">
<paragraph id="8BE536A4FFD3531FCD1452D9FD50A021" blockId="83.[264,1323,283,862]" pageId="83" pageNumber="84">
<emphasis id="B92EEAB6FFD3531FCD1452D9FE6FA39A" bold="true" box="[312,439,395,421]" pageId="83" pageNumber="84">Diagnosis.</emphasis>
<taxonomicName id="4C5A4D27FFD3531FCDFC52DEFCD0A39A" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[464,776,396,421]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD3531FCDFC52DEFCD0A39A" box="[464,776,396,421]" italics="true" pageId="83" pageNumber="84">Boreophilia angusticornis</emphasis>
</taxonomicName>
is closely related to Palaearctic
<taxonomicName id="4C5A4D27FFD3531FC88C52DEFDBEA3F1" authority="(J. Sahlberg, 1880)" baseAuthorityName="J. Sahlberg" baseAuthorityYear="1880" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="subplana">
<emphasis id="B92EEAB6FFD3531FC88C52DEFEABA3F2" italics="true" pageId="83" pageNumber="84">Boreophilia subplana</emphasis>
(J.
<bibRefCitation id="EFCB4B55FFD3531FCD8B52E6FD86A3F1" author="Sahlberg, J." box="[423,606,436,462]" pageId="83" pageNumber="84" pagination="1 - 115" refId="ref80630" refString="Sahlberg, J. (1880) Bidrag till Nordvestra Sibiriens Insektfauna. Coleoptera. Insamlade under Expeditionerna till obi och Jenessej 1876 och 1877. I. Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, Dryopidae, Georyssidae, Limnichidae, Heteroceridae, Staphylinidae och Micropeplidae. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 17 (4), 1 - 115." type="journal article" year="1880">Sahlberg, 1880</bibRefCitation>
)
</taxonomicName>
but differs in the shape of the aedeagus (
<figureCitation id="13612A21FFD3531FC84252E6FAC4A3F1" box="[1134,1308,436,462]" captionStart="FIGURES 120124" captionStartId="82.[264,383,1727,1751]" captionTargetBox="[286,1306,861,1691]" captionTargetId="figure-191@82.[272,1315,852,1701]" captionTargetPageId="82" captionText="FIGURES 120124. Genitalia of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (120123); and female, lectotype of Metaxya plutonica Casey (124)). 120 — median lobe, parameral view; 121 — apex of median lobe with retracted internal sac, abparameral view; 122 — median lobe, lateral view; 123 — apex of median lobe with retracted internal sac, lateral view; 124 — spermatheca. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5019870" httpUri="https://zenodo.org/record/5019870/files/figure.png" pageId="83" pageNumber="84">Figs. 120123</figureCitation>
), particularly a narrower apex of the median lobe (in parameral view) (cf.
<figureCitation id="13612A21FFD3531FC878528EFB7EA3C9" box="[1108,1190,476,502]" captionStart="FIGURES 7074" captionStartId="56.[264,382,1344,1368]" captionTargetBox="[295,1288,322,1307]" captionTargetId="figure-167@56.[271,1315,307,1318]" captionTargetPageId="56" captionText="FIGURES 7074. Abdominal segment 8 of Atheta frosti Bernhauer (male, lectotype of At. modiella Casey (7072); and female, lectotype of At. frosti (7374)). 70 — male tergum 8; 71 — male sternum 8; 72 — apex of male tergum 8; 73 — female tergum 8; 74 — female sternum 8. Scale bar 0.1 mm (72), 0.2 mm (7071, 7374)." figureDoi="http://doi.org/10.5281/zenodo.5019834" httpUri="https://zenodo.org/record/5019834/files/figure.png" pageId="83" pageNumber="84">
Fig.
<quantity id="4CA29B41FFD3531FC8A5528EFB7EA3C9" box="[1161,1190,476,502]" metricMagnitude="0" metricUnit="m" metricValue="1.8542" pageId="83" pageNumber="84" unit="in" value="73.0">73</quantity>
</figureCitation>
in Brundin 1953 and
<figureCitation id="13612A21FFD3531FCD505156FE06A021" box="[380,478,516,542]" captionStart="FIGURES 120124" captionStartId="82.[264,383,1727,1751]" captionTargetBox="[286,1306,861,1691]" captionTargetId="figure-191@82.[272,1315,852,1701]" captionTargetPageId="82" captionText="FIGURES 120124. Genitalia of Boreophilia angusticornis (Bernhauer) (male, lectotype of Atheta angusticornis (120123); and female, lectotype of Metaxya plutonica Casey (124)). 120 — median lobe, parameral view; 121 — apex of median lobe with retracted internal sac, abparameral view; 122 — median lobe, lateral view; 123 — apex of median lobe with retracted internal sac, lateral view; 124 — spermatheca. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5019870" httpUri="https://zenodo.org/record/5019870/files/figure.png" pageId="83" pageNumber="84">
Fig.
<quantity id="4CA29B41FFD3531FCD9E5156FE06A021" box="[434,478,516,542]" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="83" pageNumber="84" unit="in" value="120.0">120</quantity>
</figureCitation>
in this paper).
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD3531FCD145179FDB9A081" pageId="83" pageNumber="84" type="discussion">
<paragraph id="8BE536A4FFD3531FCD145179FDB9A081" blockId="83.[264,1323,283,862]" pageId="83" pageNumber="84">
<emphasis id="B92EEAB6FFD3531FCD145179FE1BA07A" bold="true" box="[312,451,555,581]" pageId="83" pageNumber="84">Discussion.</emphasis>
The
<typeStatus id="54E18806FFD3531FCE2A517EFDA9A079" box="[518,625,556,582]" pageId="83" pageNumber="84">lectotype</typeStatus>
of
<emphasis id="B92EEAB6FFD3531FCEB4517EFCE2A07A" box="[664,826,556,581]" italics="true" pageId="83" pageNumber="84">
Me.
<taxonomicName id="4C5A4D27FFD3531FCEE0517EFCE2A07A" authorityName="Casey" authorityYear="1910" box="[716,826,556,581]" class="Polypodiopsida" family="Metaxyaceae" genus="Metaxya" kingdom="Plantae" order="Cyatheales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="plutonica">plutonica</taxonomicName>
</emphasis>
and the
<typeStatus id="54E18806FFD3531FCFB2517EFBE4A079" box="[926,1084,556,582]" pageId="83" pageNumber="84">paralectotype</typeStatus>
of
<taxonomicName id="4C5A4D27FFD3531FC84E517EFAF2A07A" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[1122,1322,556,581]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD3531FC84E517EFAF2A07A" box="[1122,1322,556,581]" italics="true" pageId="83" pageNumber="84">At. angusticornis</emphasis>
</taxonomicName>
are similar in external characters and also in the shape of the spermatheca. The two species were described from the same locality. I consider
<emphasis id="B92EEAB6FFD3531FCF97512EFB87A0AA" box="[955,1119,636,661]" italics="true" pageId="83" pageNumber="84">
Me.
<taxonomicName id="4C5A4D27FFD3531FCFDC512EFB87A0AA" authorityName="Casey" authorityYear="1910" box="[1008,1119,636,661]" class="Polypodiopsida" family="Metaxyaceae" genus="Metaxya" kingdom="Plantae" order="Cyatheales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="plutonica">plutonica</taxonomicName>
</emphasis>
to be a synonym of
<taxonomicName id="4C5A4D27FFD3531FCD0451F6FD81A082" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[296,601,676,701]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD3531FCD0451F6FD81A082" box="[296,601,676,701]" italics="true" pageId="83" pageNumber="84">Boreophilia angusticornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFD3531FCD14519EFB70A161" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BE536A4FFD3531FCD14519EFB70A161" blockId="83.[264,1323,283,862]" pageId="83" pageNumber="84">
<emphasis id="B92EEAB6FFD3531FCD14519EFE00A0D9" bold="true" box="[312,472,716,742]" pageId="83" pageNumber="84">Distribution.</emphasis>
<taxonomicName id="4C5A4D27FFD3531FCDC5519EFCC3A0DA" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[489,795,716,741]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD3531FCDC5519EFCC3A0DA" box="[489,795,716,741]" italics="true" pageId="83" pageNumber="84">Boreophilia angusticornis</emphasis>
</taxonomicName>
is known only from the Mount
<collectingRegion id="499EF846FFD3531FC8B5519EFAFFA0D9" box="[1177,1319,716,742]" country="United States of America" name="Washington" pageId="83" pageNumber="84">Washington</collectingRegion>
,
<collectingRegion id="499EF846FFD3531FCD2451A6FE12A131" box="[264,458,756,782]" country="United States of America" name="New Hampshire" pageId="83" pageNumber="84">New Hampshire</collectingRegion>
. This area was affected by the last glaciation and is unlikely to have relict endemic species.
<taxonomicName id="4C5A4D27FFD3531FCDC5504EFCC7A10A" authorityName="Gusarov" authorityYear="2003" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[489,799,796,821]" class="Insecta" family="Staphylinidae" genus="Boreophilia" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="angusticornis">
<emphasis id="B92EEAB6FFD3531FCDC5504EFCC7A10A" box="[489,799,796,821]" italics="true" pageId="83" pageNumber="84">Boreophilia angusticornis</emphasis>
</taxonomicName>
may occur in other high mountains of the north­eastern
<collectingCountry id="F34D7634FFD3531FCD865016FD9EA161" box="[426,582,836,862]" name="United States of America" pageId="83" pageNumber="84">United States</collectingCountry>
and also in
<collectingCountry id="F34D7634FFD3531FCEF85016FCF7A161" box="[724,815,836,862]" name="Canada" pageId="83" pageNumber="84">Canada</collectingCountry>
, at lower altitudes further north.
</paragraph>
</subSubSection>
</treatment>
</document>