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<document id="A1A91065AFEBCBF8D631CCA6A896DA56" ID-DOI="10.11646/zootaxa.4995.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5057385" ID-ZooBank="AF31D6D4-AFD4-46A2-A400-29B0281810E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625172752147" checkinUser="plazi" docAuthor="Hara, Hideho, Smith, David R. &amp; Shinohara, Akihiko" docDate="2021" docId="03F1576FFFC06946FBB5FF3BFD03F992" docLanguage="en" docName="zootaxa.4995.3.4.pdf" docOrigin="Zootaxa 4995 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gilpinia okinawa Hara, Smith &amp; Shinohara 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="485" masterDocId="FFC82F17FFCD6948FB22FFACFF90FA5B" masterDocTitle="Gilpinia hakonensis and similar species in Japan and ovipositors of five European Gilpinia species (Hymenoptera, Diprionidae)" masterLastPageNumber="491" masterPageNumber="471" pageNumber="484" updateTime="1698982284166" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5F5D768683C29E7AC9E3E985EF891BA3">Gilpinia hakonensis and similar species in Japan and ovipositors of five European Gilpinia species (Hymenoptera, Diprionidae)</mods:title>
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<mods:namePart id="1008BF3B2D65929ABC1FD1C0D18DC0E0">Hara, Hideho</mods:namePart>
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<mods:namePart id="8A5DF04CEF54C996E86E1D8207A7090D">Smith, David R.</mods:namePart>
<mods:affiliation id="33507B1D9462BAC0728F188A0CE99902">Systematic Entomology Laboratory, c / o National Museum of Natural History, Smithsonian Institution, P. O. Box 37012, MRC 168, Washington, DC 20013 - 7012, USA.</mods:affiliation>
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<mods:namePart id="35290FAC588C4B4307F721EEDB7FB613">Shinohara, Akihiko</mods:namePart>
<mods:affiliation id="483B47B95B884F3C55C11C74A4447225">National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:date id="73E5109D5CAF9CAE2765AFD558173EFC">2021</mods:date>
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<treatment id="03F1576FFFC06946FBB5FF3BFD03F992" ID-DOI="http://doi.org/10.5281/zenodo.5057416" ID-Zenodo-Dep="5057416" LSID="urn:lsid:plazi:treatment:03F1576FFFC06946FBB5FF3BFD03F992" httpUri="http://treatment.plazi.org/id/03F1576FFFC06946FBB5FF3BFD03F992" lastPageId="14" lastPageNumber="485" pageId="13" pageNumber="484">
<subSubSection id="C342B5F2FFC06945FBB5FF3BFCB1FAEA" box="[151,801,151,178]" pageId="13" pageNumber="484" type="nomenclature">
<paragraph id="8BE7E679FFC06945FBB5FF3BFCB1FAEA" blockId="13.[151,801,151,214]" box="[151,801,151,178]" pageId="13" pageNumber="484">
<heading id="D0AF5115FFC06945FBB5FF3BFCB1FAEA" bold="true" box="[151,801,151,178]" fontSize="11" level="1" pageId="13" pageNumber="484" reason="1">
<emphasis id="B92C3A6BFFC06945FBB5FF3BFCB1FAEA" bold="true" box="[151,801,151,178]" pageId="13" pageNumber="484">
<taxonomicName id="4C589DFAFFC06945FBB5FF3BFD26FAEA" authority="Hara, Smith &amp; Shinohara" authorityName="Hara, Smith &amp; Shinohara" authorityYear="2021" box="[151,694,151,178]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="484" phylum="Arthropoda" rank="species" species="okinawa" status="sp. nov.">
<emphasis id="B92C3A6BFFC06945FBB5FF3BFEF9FAE9" bold="true" box="[151,361,151,178]" italics="true" pageId="13" pageNumber="484">Gilpinia okinawa</emphasis>
Hara, Smith &amp; Shinohara
</taxonomicName>
,
<taxonomicNameLabel id="A21F8710FFC06945F9E1FF3BFCB1FAEA" box="[707,801,151,177]" pageId="13" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C342B5F2FFC06946FBB5FF17FE18F896" lastPageId="14" lastPageNumber="485" pageId="13" pageNumber="484" type="description">
<paragraph id="8BE7E679FFC06945FBB5FF17FD65FA8D" blockId="13.[151,801,151,214]" box="[151,757,187,214]" pageId="13" pageNumber="484">
(
<figureCitation id="1363FAFCFFC06945FBBDFF17FEADFA8D" box="[159,317,187,214]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Figs 2G, H, L</figureCitation>
,
<figureCitation id="1363FAFCFFC06945FA6BFF17FE3BFA8D" box="[329,427,187,214]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="13" pageNumber="484">3H, P, X</figureCitation>
, 4M, N, R, 5K, L, T, U, 6I, J)
</paragraph>
<paragraph id="8BE7E679FFC06945FBB5FEAFFE10F896" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="484">
<emphasis id="B92C3A6BFFC06945FBB5FEAFFF7EFB46" bold="true" box="[151,238,259,285]" pageId="13" pageNumber="484">Female</emphasis>
(
<figureCitation id="1363FAFCFFC06945FBDDFEAFFE15FB45" box="[255,389,259,286]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Figs 2G, H</figureCitation>
). Length
<quantity id="4CA04B9CFFC06945FAD1FEAFFDDFFB45" box="[499,591,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="484" unit="mm" value="9.5">9.5 mm</quantity>
. Head black, reddish yellow on wide ventromedial part of face (
<figureCitation id="1363FAFCFFC06945FE17FEAFFA1FFB46" box="[1333,1423,259,285]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Fig. 2L</figureCitation>
); postocellar area widely dark brown (
<figureCitation id="1363FAFCFFC06945F910FE8BFD17FB1A" box="[562,647,295,321]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="13" pageNumber="484">Fig. 3P</figureCitation>
); labrum red yellow or brown; antenna black, basally narrowly yellow (
<figureCitation id="1363FAFCFFC06945FB82FEE7FF6AFB3E" box="[160,250,331,357]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="13" pageNumber="484">Fig. 3X</figureCitation>
); mandible reddish brown, basally dark brown, with yellow spot on outer basal part; palpi yellow. Thorax red yellow, dark brown to black on most of lateral mesoscutal lobe, most of posterolateral sunken part of mesoscutum, mesoscutellar appendage, anteromedial part of mesopostnotum, metascutum, wide part of anepimeron, anterior margin of katepimeron, and ventral part of metepisternum (
<figureCitation id="1363FAFCFFC06945F87AFE1BFC47FB89" box="[856,983,439,466]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Figs 2G, H</figureCitation>
, 4M, N). Legs red yellow (
<figureCitation id="1363FAFCFFC06945FE20FE1BFACCFB8A" box="[1282,1372,439,465]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Fig. 2H</figureCitation>
); fore and middle coxae basally narrowly blackish; hind coxa widely black; middle and hind tibiae basally pale yellow; hind trochanter widely dark brown. Wings colorless transparent (
<figureCitation id="1363FAFCFFC06945F84AFE53FC52F842" box="[872,962,511,537]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Fig. 2G</figureCitation>
); veins brown to black; most of vein C and bases of vein 1A and 2A+3A yellow; vein R1 basal to stigma widely pale; stigma brown, black on narrow margins and basal third.Abdomen red yellow; second to eighth terga dorsally widely black along posterior margins, and their laterotergites narrowly black along posterior margins; posterior sterna narrowly darkened along posterior margins; ninth tergum posteroventrally black; ovipositor sheath with valvifer 2 widely black, valvula 3 mostly black; cercus dark brown to black.
</paragraph>
<paragraph id="8BE7E679FFC06945FBE5FD7BFCE0F9FE" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="484">
Postocellar area distinctly swollen; lateral furrow distinct (
<figureCitation id="1363FAFCFFC06945F87FFD7BFC23F8AA" box="[861,947,727,753]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="13" pageNumber="484">Fig. 3P</figureCitation>
). OOL:POL:OOCL 1.0:1.0:0.91.0. OOCL 2.83.1× lateral ocellus length. Inner margins of eyes convergent below (
<figureCitation id="1363FAFCFFC06945F8F6FD57FBBCF94E" box="[980,1068,763,789]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="13" pageNumber="484">Fig. 2L</figureCitation>
). Distance between eyes at level of torulus 2.0 × major axis of eye. Supraclypeal area moderately swollen. Clypeus with ventral margin very slightly concave medially; median height 1.0 × torulus height. Malar space 1.01.1 × median ocellus width. Antenna with 2223 antennomeres (
<figureCitation id="1363FAFCFFC06945FAB0FCCBFE7EF9DA" box="[402,494,871,897]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="13" pageNumber="484">Fig. 3X</figureCitation>
); first flagellomere without serration, with apical breadth in lateral view 0.91.0 × dorsal length; serrations of third to seventh flagellomeres largest.
</paragraph>
<caption id="DF27B6F1FFC06945FBB5F8D7FB47FDEC" ID-DOI="http://doi.org/10.5281/zenodo.5057405" ID-Zenodo-Dep="5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="13" pageNumber="484" startId="13.[151,264,1915,1939]" targetBox="[151,1423,960,1891]" targetPageId="13">
<paragraph id="8BE7E679FFC06945FBB5F8D7FB47FDEC" blockId="13.[151,1436,1915,1975]" pageId="13" pageNumber="484">
<emphasis id="B92C3A6BFFC06945FBB5F8D7FEDCFDC8" bold="true" box="[151,332,1915,1939]" pageId="13" pageNumber="484">FIGURES 7AF.</emphasis>
A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). AC,
<taxonomicName id="4C589DFAFFC06945FAF1F833FC89FDEC" authority=", Honshu. D" authorityName="Honshu. D" box="[467,793,1951,1975]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="484" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC06945FAF1F833FD0FFDEC" box="[467,671,1951,1975]" italics="true" pageId="13" pageNumber="484">Gilpinia hakonensis</emphasis>
, Honshu. D
</taxonomicName>
F,
<taxonomicName id="4C589DFAFFC06945F81CF833FB47FDEC" authority=", Amami Oshima Isl." box="[830,1239,1951,1975]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="484" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC06945F81CF833FB94FDEC" box="[830,1028,1951,1975]" italics="true" pageId="13" pageNumber="484">Gilpinia amamiana</emphasis>
, Amami Oshima Isl.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BE7E679FFC36946FBE5FF3BFB36FB46" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="485">Distance between cenchri 0.70.8 × major axis of cenchrus (Fig. 4M). Metascutellum in dorsal view posterolaterally angulated, with posterior margin truncate; median length 0.40.5 × major axis of cenchrus. Hind leg with posterior tibial spur with length 1.01.1 × dorsal length of first tarsomere, 0.91.1 × apical breadth of tibia in lateral view; dorsal length of second and third tarsomeres combined 1.51.6 × dorsal length of first.</paragraph>
<paragraph id="8BE7E679FFC36946FBE5FE8BFB31F866" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="485">
Valvula
<quantity id="4CA04B9CFFC36946FA07FE8BFEC1FB1A" box="[293,337,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="485" unit="in" value="3.0">3 in</quantity>
dorsal view about 3.0 × as wide as cercus, apically with round lateral projection and small acute medial projection (
<figureCitation id="1363FAFCFFC36946FA4CFEE7FE5AFB3E" box="[366,458,330,357]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="14" pageNumber="485">Fig. 5K</figureCitation>
); scopa about 1.52.0 × as wide as cercus, in posterior view long oval, about 2.5 × as high as wide (
<figureCitation id="1363FAFCFFC36946FA10FEC3FE18FBD2" box="[306,392,366,393]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="14" pageNumber="485">Fig. 5L</figureCitation>
). Lance with posterior projection on processus articularis apically not extending to most basal annular suture of lance (
<figureCitation id="1363FAFCFFC36946FA8BFE3FFDB7FBF6" box="[425,551,402,429]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="14" pageNumber="485">Figs 5T, U</figureCitation>
). Lancet
<figureCitation id="1363FAFCFFC36946F9ADFE3FFD67FBF6" box="[655,759,403,429]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="14" pageNumber="485">Fig. 6I, J</figureCitation>
; lamnium with 9 annuli and length from ventral end of first ctenidium to apex 2.2 × maximum width; ctenidia nearly straight except second arched and third slightly arched; sclerite before first annulus narrow; serrula of second annulus with or without precalcar (absence of precalcar probably due to severe wear,
<figureCitation id="1363FAFCFFC36946FAFDFE53FDBDF841" box="[479,557,511,538]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="14" pageNumber="485">Fig. 6J</figureCitation>
) and serrula of third annulus with very small precalcar or without it; serrula of second annulus much larger than other serrulae, with precalcar relatively close to postcalcar.
</paragraph>
<paragraph id="8BE7E679FFC36946FBE5FDEBFD92F8F2" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="485">
Head and thorax mostly covered with dense punctures (
<figureCitation id="1363FAFCFFC36946F80FFDEBFC12F83A" box="[813,898,583,609]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="14" pageNumber="485">Figs 3P</figureCitation>
, 4M, N); punctures very often fused; interspaces generally narrower than diameters of punctures. Punctures on mesoscutellum much larger than those of mesoscutum. Abdomen microsculptured.
</paragraph>
<paragraph id="8BE7E679FFC36946FBE5FD1FFE18F896" blockId="14.[151,1437,151,969]" box="[199,392,691,717]" pageId="14" pageNumber="485">
<emphasis id="B92C3A6BFFC36946FBE5FD1FFE94F896" bold="true" box="[199,260,691,717]" pageId="14" pageNumber="485">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC36946FBE5FD7AFEFCF962" pageId="14" pageNumber="485" type="materials_examined">
<paragraph id="8BE7E679FFC36946FBE5FD7AFEFCF962" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="485">
<emphasis id="B92C3A6BFFC36946FBE5FD7AFE3AF8AB" bold="true" box="[199,426,726,752]" pageId="14" pageNumber="485">Material examined</emphasis>
.
<materialsCitation id="3B30EC24FFC36946FA9BFD7BFC81F94E" collectingDate="2014-06-16" collectionCode="R, T" collectorName="H. Ogai" country="Japan" elevation="160" location="Okinawa" pageId="14" pageNumber="485" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="54E358DBFFC36946FA9BFD7BFDB1F8AA" box="[441,545,727,753]" pageId="14" pageNumber="485" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1363FAFCFFC36946F912FD7BFD41F8AA" box="[560,721,727,753]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="14" pageNumber="485">Figs 2G, H, L</figureCitation>
,
<figureCitation id="1363FAFCFFC36946F9FFFD7BFCD0F8AA" box="[733,832,727,753]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="14" pageNumber="485">3H, P, X</figureCitation>
, 4M, N,
<collectionCode id="ED497EBCFFC36946F887FD7BFC2AF8AA" box="[933,954,727,753]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="485">R</collectionCode>
, 5K, L,
<collectionCode id="ED497EBCFFC36946FF34FD7BFBB8F8AA" box="[1046,1064,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="485">T</collectionCode>
, U, 6I):
<specimenCount id="9D5E2DF0FFC36946FFA5FD7BFB0CF8AA" box="[1159,1180,727,753]" pageId="14" pageNumber="485" type="female"></specimenCount>
, “
<date id="FFE6C0B9FFC36946FF97FD7BFAABF8AA" box="[1205,1339,727,753]" pageId="14" pageNumber="485" value="2014-06-16">
<collectingDate id="EFA23951FFC36946FF97FD7BFAABF8AA" box="[1205,1339,727,753]" pageId="14" pageNumber="485" value="2014-06-16">16-VI-2014</collectingDate>
</date>
(
<quantity id="4CA04B9CFFC36946FE69FD7BFA1FF8A9" box="[1355,1423,727,754]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="485" unit="m" value="160.0">
<elevation id="0075014AFFC36946FE69FD7BFA1FF8A9" box="[1355,1423,727,754]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="485" unit="m" value="160.0">160m</elevation>
</quantity>
), Nangusuku parc, Nago-shi,
<collectingRegion id="499C289BFFC36946FAF0FD57FDAAF94E" box="[466,570,763,789]" country="Japan" name="Okinawa" pageId="14" pageNumber="485">Okinawa</collectingRegion>
,
<collectingCountry id="F34FA6E9FFC36946F967FD57FD0DF94E" box="[581,669,763,789]" name="Japan" pageId="14" pageNumber="485">JAPAN</collectingCountry>
,
<collectorName id="26AD83AFFFC36946F98AFD57FC97F94E" box="[680,775,763,789]" pageId="14" pageNumber="485">H. Ogai</collectorName>
</materialsCitation>
.
<materialsCitation id="3B30EC24FFC36946F83EFD57FEF8F962" collectingDate="2020-06-04" collectorName="S. Inada" country="Japan" county="Island" location="Genka" municipality="Nago" pageId="14" pageNumber="485" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus id="54E358DBFFC36946F83EFD57FC12F94E" box="[796,898,763,789]" pageId="14" pageNumber="485" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D5E2DF0FFC36946F8AFFD57FC20F94E" box="[909,944,763,789]" pageId="14" pageNumber="485" type="female">1♀</specimenCount>
(
<figureCitation id="1363FAFCFFC36946F8E2FD57FB80F94D" box="[960,1040,763,790]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="14" pageNumber="485">Fig. 6J</figureCitation>
),
<collectingRegion id="499C289BFFC36946FF07FD57FB1AF94E" box="[1061,1162,763,789]" country="Japan" name="Okinawa" pageId="14" pageNumber="485">Okinawa</collectingRegion>
<collectingCounty id="62869EF5FFC36946FFB0FD57FB49F94E" box="[1170,1241,763,789]" pageId="14" pageNumber="485">Island</collectingCounty>
,
<collectingMunicipality id="6B837C03FFC36946FFC7FD57FAB4F94E" box="[1253,1316,763,789]" pageId="14" pageNumber="485">Nago</collectingMunicipality>
,
<location id="8E87B0A2FFC36946FE12FD57FAECF94E" LSID="urn:lsid:plazi:treatment:03F1576FFFC06946FBB5FF3BFD03F992:8E87B0A2FFC36946FE12FD57FAECF94E" box="[1328,1404,763,789]" country="Japan" county="Island" municipality="Nago" name="Genka" pageId="14" pageNumber="485" stateProvince="Okinawa">Genka</location>
,
<date id="FFE6C0B9FFC36946FEA5FD57FF6EF962" pageId="14" pageNumber="485" value="2020-06-04">
<collectingDate id="EFA23951FFC36946FEA5FD57FF6EF962" pageId="14" pageNumber="485" value="2020-06-04">4. VI. 2020</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC36946FA2BFCB3FEF8F962" box="[265,360,799,825]" pageId="14" pageNumber="485">S. Inada</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC36946FBE5FCEFFB96F906" box="[199,1030,835,861]" pageId="14" pageNumber="485" type="etymology">
<paragraph id="8BE7E679FFC36946FBE5FCEFFB96F906" blockId="14.[151,1437,151,969]" box="[199,1030,835,861]" pageId="14" pageNumber="485">
<emphasis id="B92C3A6BFFC36946FBE5FCEFFED8F906" bold="true" box="[199,328,835,861]" pageId="14" pageNumber="485">Etymology</emphasis>
. The species epithet is derived from the locality and is a noun.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC36946FBE5FCCAFDFEF9DA" box="[199,622,870,897]" pageId="14" pageNumber="485" type="distribution">
<paragraph id="8BE7E679FFC36946FBE5FCCAFDFEF9DA" blockId="14.[151,1437,151,969]" box="[199,622,870,897]" pageId="14" pageNumber="485">
<emphasis id="B92C3A6BFFC36946FBE5FCCAFEC9F9DB" bold="true" box="[199,345,870,896]" pageId="14" pageNumber="485">Distribution</emphasis>
.
<collectingCountry id="F34FA6E9FFC36946FA45FCCBFE3BF9DA" box="[359,427,871,897]" name="Japan" pageId="14" pageNumber="485">Japan</collectingCountry>
:
<collectingRegion id="499C289BFFC36946FA94FCCBFD8BF9DA" box="[438,539,871,897]" country="Japan" name="Okinawa" pageId="14" pageNumber="485">Okinawa</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC36946FBE5FC27FD03F992" pageId="14" pageNumber="485" type="discussion">
<paragraph id="8BE7E679FFC36946FBE5FC27FD03F992" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="485">
<emphasis id="B92C3A6BFFC36946FBE5FC27FEA4F9FE" bold="true" box="[199,308,907,933]" pageId="14" pageNumber="485">Remarks</emphasis>
.
<taxonomicName id="4C589DFAFFC36946FA60FC27FD95F9FE" authorityName="Hara, Smith &amp; Shinohara" authorityYear="2021" box="[322,517,907,933]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="okinawa">
<emphasis id="B92C3A6BFFC36946FA60FC27FD95F9FE" box="[322,517,907,933]" italics="true" pageId="14" pageNumber="485">Gilpinia okinawa</emphasis>
</taxonomicName>
is distinguished from other species of
<taxonomicName id="4C589DFAFFC36946F897FC27FB82F9FE" authorityName="Benson" authorityYear="1939" box="[949,1042,907,933]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B92C3A6BFFC36946F897FC27FB82F9FE" box="[949,1042,907,933]" italics="true" pageId="14" pageNumber="485">Gilpinia</emphasis>
</taxonomicName>
as stated in the section of
<taxonomicName id="4C589DFAFFC36946FE18FC27FF4DF993" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="485" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC36946FE18FC27FF4DF993" italics="true" pageId="14" pageNumber="485">G. hakonensis</emphasis>
</taxonomicName>
and similar species and the key above.
</paragraph>
</subSubSection>
</treatment>
</document>