162 lines
18 KiB
XML
162 lines
18 KiB
XML
<document id="33773B573D8C8F92F1CFBD539B0B79DC" ID-DOI="10.5281/zenodo.196882" ID-GBIF-Dataset="5f7352b6-e07e-4e40-95b9-2a5c2daec87e" ID-ISSN="1175-5326" ID-Zenodo-Dep="196882" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460255369181" checkinUser="plazi" docAuthor="Fend, Steven V. & Lenat, David R." docDate="2010" docId="03EEF0432A3B6B65A1D3FE89FDB8A788" docLanguage="en" docName="zt02554p022.pdf" docOrigin="Zootaxa 2554" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pararhynchelmis Fend & Lenat, 2010, n. gen." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FFD7883B2A366B68A144FFFEFFB9A22C" masterDocTitle="New southeastern Nearctic Rhynchelmis (Rhynchelmoides) species and the description of Pararhynchelmis n. gen. (Annelida: Clitellata: Lumbriculidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="14" updateTime="1698243314012" updateUser="plazi">
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<mods:mods id="DCBB4D32AEA7B367021B70ABBA6BF75C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6AFC87BF49ED4B33A403A118375999B8">
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<mods:title id="6375BDF56733A988F0E0627EB321FB9E">New southeastern Nearctic Rhynchelmis (Rhynchelmoides) species and the description of Pararhynchelmis n. gen. (Annelida: Clitellata: Lumbriculidae)</mods:title>
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<mods:name id="11D53D97DA65ACA688FF55F485F299C2" type="personal">
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<mods:namePart id="B4ADD7087DDFD459513CD710BA3C47BC">Fend, Steven V.</mods:namePart>
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<mods:roleTerm id="1B80E83056E617732E7E85FCB55D758A">Author</mods:roleTerm>
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<mods:namePart id="5908159E0AAA31F7BB4807069C9A9E77">Lenat, David R.</mods:namePart>
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<mods:typeOfResource id="BEF61979D6A85643C5A3588A277F4CB2">text</mods:typeOfResource>
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<mods:title id="A4796395964AA775410B74F8050A9571">Zootaxa</mods:title>
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<mods:part id="D3300C2BCDA4A3077567A9A78D417D60">
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<mods:date id="DC59431C05FA571249192A79742E0982">2010</mods:date>
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<mods:detail id="F9523C5B8A6AB01D6063D9C5CDACED55" type="volume">
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<mods:number id="B027DBF014982F09D06D23EB1B1004E8">2554</mods:number>
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<mods:start id="86C77217B028A729353E61C2A8318A6D">1</mods:start>
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<mods:classification id="8A096B696EBA67EE82706EFC45AD73C1">journal article</mods:classification>
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<mods:identifier id="C8E1A4AF205DB9892589B64D594AC045" type="DOI">10.5281/zenodo.196882</mods:identifier>
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<mods:identifier id="BF163FFC88EDBD18A91BA00268D2F2D6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="397C22006A51BF440F6DA2171B29AC81" type="Zenodo-Dep">196882</mods:identifier>
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<treatment id="03EEF0432A3B6B65A1D3FE89FDB8A788" ID-DOI="http://doi.org/10.5281/zenodo.6206485" ID-GBIF-Taxon="119392351" ID-Zenodo-Dep="6206485" LSID="urn:lsid:plazi:treatment:03EEF0432A3B6B65A1D3FE89FDB8A788" httpUri="http://treatment.plazi.org/id/03EEF0432A3B6B65A1D3FE89FDB8A788" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="C35D12DE2A3B6B65A1D3FE89FD45A3F5" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BF841552A3B6B65A1D3FE89FE79A3BD" blockId="13.[151,448,375,401]" box="[151,448,375,401]" pageId="13" pageNumber="14">
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<heading id="D0B0F6392A3B6B65A1D3FE89FE79A3BD" bold="true" box="[151,448,375,401]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B9339D472A3B6B65A1D3FE89FE79A3BD" bold="true" box="[151,448,375,401]" pageId="13" pageNumber="14">
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<taxonomicName id="4C473AD62A3B6B65A1D3FE89FED1A3BD" ID-CoL="84RQX" box="[151,360,375,401]" class="Clitellata" family="Lumbriculidae" genus="Pararhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B9339D472A3B6B65A1D3FE89FED1A3BD" bold="true" box="[151,360,375,401]" italics="true" pageId="13" pageNumber="14">Pararhynchelmis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A200203C2A3B6B65A02AFE89FE79A3BD" box="[366,448,375,401]" pageId="13" pageNumber="14" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF841552A3B6B65A1D3FE41FD45A3F5" blockId="13.[151,1437,447,1444]" box="[151,764,447,473]" pageId="13" pageNumber="14">
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<emphasis id="B9339D472A3B6B65A1D3FE41FED4A3F5" bold="true" box="[151,365,447,473]" pageId="13" pageNumber="14">Included species:</emphasis>
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<taxonomicName id="4C473AD62A3B6B65A030FE3EFD0AA3F5" box="[372,691,448,473]" class="Clitellata" family="Lumbriculidae" genus="Pararhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="murdocki" status="sp. nov.">
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<emphasis id="B9339D472A3B6B65A030FE3EFD0AA3F5" box="[372,691,448,473]" italics="true" pageId="13" pageNumber="14">Pararhynchelmis murdocki</emphasis>
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</taxonomicName>
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<emphasis id="B9339D472A3B6B65A3FEFE41FD45A3F5" bold="true" box="[698,764,447,473]" pageId="13" pageNumber="14">
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<taxonomicNameLabel id="A200203C2A3B6B65A3FEFE41FD45A3F5" box="[698,764,447,473]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D12DE2A3B6B65A181FE1BFE17A106" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8BF841552A3B6B65A181FE1BFE17A106" blockId="13.[151,1437,447,1444]" pageId="13" pageNumber="14">
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<emphasis id="B9339D472A3B6B65A181FE1BFEF7A3D3" bold="true" box="[197,334,485,511]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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Small worms with rounded prostomium. Lateral blood vessels in middle and posterior segments unbranched. Preclitellar nephridia on 6/7. Pharynx without a distinct dorsal pad. Male pores in X, behind and slightly medial to chaetae; spermathecal pores inconspicuous, paired in VIII and IX, behind and slightly medial to chaetae; female pores on chaetal line, intersegmental on 11/12. Testes in IX and X, ovaries in XI. Spermathecae simple, roughly cylindrical; without ampulla or diverticula, connected to the gut entally. Atria petiolate, with thin duct, and club-shaped ampulla covered with multicellular prostate glands. Very short, retractable penes. Semiprosoporous; each atrium with two thin vasa deferentia; both anterior and posterior male funnels functional; posterior vasa deferentia do not form a loop in the postatrial segment or penetrate septum 10/11.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D12DE2A3B6B65A181FCCBFA8CA163" box="[197,1333,821,847]" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="8BF841552A3B6B65A181FCCBFA8CA163" blockId="13.[151,1437,447,1444]" box="[197,1333,821,847]" pageId="13" pageNumber="14">
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<emphasis id="B9339D472A3B6B65A181FCCBFEE8A163" bold="true" box="[197,337,821,847]" pageId="13" pageNumber="14">Etymology.</emphasis>
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From Latin
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<emphasis id="B9339D472A3B6B65A0A1FCCBFDA5A162" box="[485,540,821,846]" italics="true" pageId="13" pageNumber="14">para</emphasis>
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, “beside”, indicating its possible close relationship to
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<taxonomicName id="4C473AD62A3B6B65A5DDFCCBFA97A162" box="[1177,1326,821,846]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A5DDFCCBFA97A162" box="[1177,1326,821,846]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D12DE2A3B6B65A181FCA4FDB8A788" pageId="13" pageNumber="14" type="discussion">
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<paragraph id="8BF841552A3B6B65A181FCA4FB97A718" blockId="13.[151,1437,447,1444]" pageId="13" pageNumber="14">
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<emphasis id="B9339D472A3B6B65A181FCA4FEFCA158" bold="true" box="[197,325,858,884]" pageId="13" pageNumber="14">Remarks.</emphasis>
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The single known
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<taxonomicName id="4C473AD62A3B6B65A37EFCA4FCA8A15F" box="[570,785,858,883]" class="Clitellata" family="Lumbriculidae" genus="Pararhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B9339D472A3B6B65A37EFCA4FCA8A15F" box="[570,785,858,883]" italics="true" pageId="13" pageNumber="14">Pararhynchelmis</emphasis>
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</taxonomicName>
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species, described herein, has male pores in X and spermathecal pores paired in both VIII and IX, distinguishing it from all other lumbriculids except for two Palearctic
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<taxonomicName id="4C473AD62A3B6B65A053FC5BFE08A192" box="[279,433,933,958]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A053FC5BFE08A192" box="[279,433,933,958]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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species (
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<taxonomicName id="4C473AD62A3B6B65A360FC5BFC6CA193" authority="Michaelsen" authorityName="Michaelsen" box="[548,981,933,959]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="tetratheca">
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<emphasis id="B9339D472A3B6B65A360FC5BFCF8A192" box="[548,833,933,958]" italics="true" pageId="13" pageNumber="14">Rhynchelmis tetratheca</emphasis>
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Michaelsen
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</taxonomicName>
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and
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<taxonomicName id="4C473AD62A3B6B65A555FC5BFA33A193" authority="Hrabe" authorityName="Hrabe" box="[1041,1418,933,959]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="granuensis">
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<emphasis id="B9339D472A3B6B65A555FC5BFA81A192" box="[1041,1336,933,958]" italics="true" pageId="13" pageNumber="14">Rhynchelmis granuensis</emphasis>
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Hrabĕ
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</taxonomicName>
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). Four described genera (
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<taxonomicName id="4C473AD62A3B6B65A0F3FC34FC24A1C8" authority="Pseudorhynchelmis Hrabe" authorityName="Pseudorhynchelmis Hrabe" box="[439,925,970,996]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A0F3FC34FDE9A1CF" box="[439,592,970,995]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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,
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<emphasis id="B9339D472A3B6B65A325FC34FCF5A1CF" box="[609,844,970,995]" italics="true" pageId="13" pageNumber="14">Pseudorhynchelmis</emphasis>
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Hrabĕ
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</taxonomicName>
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,
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<taxonomicName id="4C473AD62A3B6B65A2EAFC34FADAA1C8" authority="Fend & Gustafson 2001" authorityName="Fend & Gustafson" authorityYear="2001" box="[942,1379,970,996]" class="Clitellata" family="Lumbriculidae" genus="Secubelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A2EAFC34FB8CA1CF" box="[942,1077,970,995]" italics="true" pageId="13" pageNumber="14">Secubelmis</emphasis>
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<bibRefCitation id="EFD63CA42A3B6B65A57AFC34FADAA1C8" author="Fend" box="[1086,1379,970,996]" pageId="13" pageNumber="22" refString="Fend, S. V. & Gustafson, D. L. (2001) A new lumbriculid genus and species from North America (Clitellata, Lumbriculidae). Hydrobiologia, 463, 13 - 22." type="journal article" year="2001">Fend & Gustafson 2001</bibRefCitation>
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</taxonomicName>
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, and
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<taxonomicName id="4C473AD62A3B6B65A1D3FC0EFEF0A625" authority="Hrabe" authorityName="Hrabe" box="[151,329,1007,1033]" class="Clitellata" family="Lumbriculidae" genus="Tatriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A1D3FC0EFF42A625" box="[151,251,1008,1033]" italics="true" pageId="13" pageNumber="14">Tatriella</emphasis>
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Hrabĕ
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</taxonomicName>
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), plus two undescribed southeastern Nearctic species (S. Fend & D. Lenat, unpublished) have the first spermathecae located two segments anterior to the atria. Of these, only
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<taxonomicName id="4C473AD62A3B6B65A505FBEBFB6EA602" box="[1089,1239,1045,1070]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A505FBEBFB6EA602" box="[1089,1239,1045,1070]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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includes species in which the spermathecae connect to the gut (
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<bibRefCitation id="EFD63CA42A3B6B65A3B3FBC4FBC1A678" author="Martin" box="[759,1144,1082,1108]" pageId="13" pageNumber="22" refString="Martin, P. & Kaygorodova, I. (2008) A new species of Pseudorhynchelmis Hrabe, 1982 (Clitellata: Lumbriculidae) from Lake Baikal, with re-descriptions of P. parva and P. olchonensis. Zootaxa, 1938, 23 - 39." type="journal article" year="2008">Martin & Kaygorodova 2008</bibRefCitation>
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). Almost all described
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<taxonomicName id="4C473AD62A3B6B65A1D3FB9EFE8AA655" box="[151,307,1120,1145]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A1D3FB9EFE8AA655" box="[151,307,1120,1145]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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species have a long, filiform proboscis. The few species or subspecies without a proboscis (
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<taxonomicName id="4C473AD62A3B6B65A1E5FB7BFCFAA6B3" authority="Michaelsen" authorityName="Michaelsen" box="[161,835,1157,1183]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="subSpecies" species="brachycephala" subSpecies="brachycephala">
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<emphasis id="B9339D472A3B6B65A1E5FB7BFD17A6B2" box="[161,686,1157,1182]" italics="true" pageId="13" pageNumber="14">Rhynchelmis brachycephala brachycephala</emphasis>
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Michaelsen
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</taxonomicName>
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,
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<taxonomicName id="4C473AD62A3B6B65A214FB7BFA85A6B3" authority="Hrabe" authorityName="Hrabe" box="[848,1340,1157,1183]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="subSpecies" species="komareki" subSpecies="brevirostra">
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<emphasis id="B9339D472A3B6B65A214FB7BFB55A6B2" box="[848,1260,1157,1182]" italics="true" pageId="13" pageNumber="14">Rhynchelmis komareki brevirostra</emphasis>
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Hrabĕ
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</taxonomicName>
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, and an undescribed Nearctic species [S. Fend, unpublished]) closely resemble proboscis-bearing species: they are all large worms, usually found in profundal lake habitats. The petiolate atria of
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<taxonomicName id="4C473AD62A3B6B65A555FB2EFB63A6C5" box="[1041,1242,1232,1257]" class="Clitellata" family="Lumbriculidae" genus="Pararhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B9339D472A3B6B65A555FB2EFB63A6C5" box="[1041,1242,1232,1257]" italics="true" pageId="13" pageNumber="14">Pararhynchelmis</emphasis>
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</taxonomicName>
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do not resemble those of any described
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<taxonomicName id="4C473AD62A3B6B65A0E3FB0BFD85A722" box="[423,572,1269,1294]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A0E3FB0BFD85A722" box="[423,572,1269,1294]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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species, and the very short, duct-like spermathecae are unique within the family. Other distinguishing characters are evaluated below, in the Discussion.
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</paragraph>
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<paragraph id="8BF841552A3B6B65A181FABEFDB8A788" blockId="13.[151,1437,447,1444]" pageId="13" pageNumber="14">
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<taxonomicName id="4C473AD62A3B6B65A181FABEFE2FA775" box="[197,406,1344,1369]" class="Clitellata" family="Lumbriculidae" genus="Pararhynchelmis" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B9339D472A3B6B65A181FABEFE2FA775" box="[197,406,1344,1369]" italics="true" pageId="13" pageNumber="14">Pararhynchelmis</emphasis>
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</taxonomicName>
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has a narrower geographic range than any of the
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<taxonomicName id="4C473AD62A3B6B65A2BEFABEFB2FA775" box="[1018,1174,1344,1369]" class="Clitellata" family="Lumbriculidae" genus="Rhynchelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="13" pageNumber="14" phylum="Annelida" rank="genus">
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<emphasis id="B9339D472A3B6B65A2BEFABEFB2FA775" box="[1018,1174,1344,1369]" italics="true" pageId="13" pageNumber="14">Rhynchelmis</emphasis>
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</taxonomicName>
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subgenera or species groups. The single known species has been collected only within a single spring system in the southern Appalachian Mountains,
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<collectingCountry id="F35001C52A3B6B65A0FAFA74FE44A788" box="[446,509,1418,1444]" name="United States of America" pageId="13" pageNumber="14">USA</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |