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<document id="DABD7FA931A2DE53D456AB21EFDD7867" ID-CLB-Dataset="55105" ID-DOI="10.5852/ejt.2023.879.2165" ID-GBIF-Dataset="d90d882a-7caa-4e31-b2e4-6e41e19e3545" ID-ISSN="2118-9773" ID-Zenodo-Dep="8155427" ID-ZooBank="7727E45F-0108-4810-8FC2-C52D038ED572" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1689603286317" checkinUser="felipe" docAuthor="Hamer, Matthew T., Lee, Roger H. &amp; Guénard, Benoit" docDate="2023" docId="03E99E542626B838FE58D1E701D0A7B1" docLanguage="en" docName="ejt-879_116-135.pdf" docOrigin="European Journal of Taxonomy 879 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.879.2165" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Temnothorax haveni Lee, Hamer &amp; Guenard 2023, sp. nov." docType="treatment" docUuid="D5640CE1-BA4A-469B-959B-444695F094C3" docUuidSource="ZooBank" docVersion="4" lastPageNumber="129" masterDocId="FFD0E62C262EB835FF85D3750533A513" masterDocTitle="First record of the genus Temnothorax Mayr, 1861 (Formicidae: Myrmicinae) in Hong Kong, with descriptions of two new species" masterLastPageNumber="135" masterPageNumber="116" pageNumber="124" updateTime="1699540638648" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0A254B24911B5D6D3D3E1A034B7DA967">First record of the genus Temnothorax Mayr, 1861 (Formicidae: Myrmicinae) in Hong Kong, with descriptions of two new species</mods:title>
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<mods:namePart id="03F0F1493F414AE45339C63DA609717B">Hamer, Matthew T.</mods:namePart>
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<mods:affiliation id="72912BFEDB227975F9D19C4AF9E23799">School of Biological Sciences, The University of Hong Kong, Kadoorie Biological Sciences Building, Pok Fu Lam Road, Hong Kong SAR, China.</mods:affiliation>
<mods:nameIdentifier id="6348741054449A49A5AD2E78D8C3C3EF" type="email">matt.hamer@hotmail.co.uk</mods:nameIdentifier>
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<mods:namePart id="10E120D66B61212FCF63684BD834D6A0">Lee, Roger H.</mods:namePart>
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<mods:affiliation id="065B42456C512F16BA0EC69F7F8C12C4">School of Biological Sciences, The University of Hong Kong, Kadoorie Biological Sciences Building, Pok Fu Lam Road, Hong Kong SAR, China. &amp; Science Unit, Lingnan University, 8 Castle Peak Road, Tuen Mun, Hong Kong SAR, China.</mods:affiliation>
<mods:nameIdentifier id="3179B6EF740C2FEFA7EB47C6BE785D06" type="email">rogerlh86@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C4EC9506A48FDA44F4A6083526119C56">Guénard, Benoit</mods:namePart>
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<mods:affiliation id="F870B062769512D5E72DD2CABB52FD2B">School of Biological Sciences, The University of Hong Kong, Kadoorie Biological Sciences Building, Pok Fu Lam Road, Hong Kong SAR, China.</mods:affiliation>
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<treatment id="03E99E542626B838FE58D1E701D0A7B1" ID-DOI="http://doi.org/10.5281/zenodo.8158715" ID-GBIF-Taxon="211662906" ID-Zenodo-Dep="8158715" LSID="urn:lsid:zoobank.org:act:D5640CE1-BA4A-469B-959B-444695F094C3" httpUri="http://treatment.plazi.org/id/03E99E542626B838FE58D1E701D0A7B1" lastPageId="13" lastPageNumber="129" pageId="8" pageNumber="124">
<subSubSection id="C35A7CC92626B83DFE58D1E70165A7BE" box="[477,1110,658,685]" pageId="8" pageNumber="124" type="nomenclature">
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<heading id="D0B7982E2626B83DFE58D1E70165A7BE" box="[477,1110,658,685]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="124" reason="2">
<taxonomicName id="4C4054C12626B83DFE58D1E706C7A7BF" authority="Lee, Hamer &amp; Guenard" authorityName="Lee, Hamer &amp; Guenard" authorityYear="2023" box="[477,1012,658,685]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="124" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502626B83DFE58D1E707EBA7BE" bold="true" box="[477,728,658,685]" italics="true" pageId="8" pageNumber="124">Temnothorax haveni</emphasis>
Lee, Hamer &amp; Guénard
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2626B83DFC79D1E60165A7BE" box="[1020,1110,659,685]" pageId="8" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BFF2F422626B83DFD62D1AD067EA7E0" blockId="8.[368,1219,658,755]" box="[743,845,728,755]" pageId="8" pageNumber="124">
<figureCitation id="137B33C72626B83DFD62D1AD067EA7E0" box="[743,845,728,755]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[189,232,1963,1989]" captionStartId-1="11.[188,231,1928,1954]" captionStartId-2="12.[189,232,1963,1989]" captionTargetBox-0="[393,1194,265,1922]" captionTargetBox-1="[189,1398,992,1886]" captionTargetBox-2="[219,1391,1101,1915]" captionTargetId-0="figure-11@10.[393,1194,265,1922]" captionTargetId-1="figure-232@11.[189,1398,992,1886]" captionTargetId-2="figure-310@12.[189,1398,1067,1922]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Fig. 3. Temnothorax haveni Lee, Hamer &amp; Guénard sp. nov., holotype (RHL03433). A. Lateral view. B. Dorsal view. C. Head in full face view." captionText-1="Fig. 4. Temnothorax haveni Lee, Hamer &amp; Guénard sp. nov., holotype (RHL03433), morphological characters. A. Petiole and postpetiole in lateral view. B. Petiole and postpetiole in dorsal view. C. Mesosoma in dorsal view. D. First gastral tergite in dorsal view." captionText-2="Fig. 5. Distribution map of Temnothorax Mayr, 1861 records from Hong Kong. The darker the green areas, the greater the tree canopy cover." figureDoi-0="http://doi.org/10.5281/zenodo.8155433" figureDoi-1="http://doi.org/10.5281/zenodo.8155435" figureDoi-2="http://doi.org/10.5281/zenodo.8155437" httpUri-0="https://zenodo.org/record/8155433/files/figure.png" httpUri-1="https://zenodo.org/record/8155435/files/figure.png" httpUri-2="https://zenodo.org/record/8155437/files/figure.png" pageId="8" pageNumber="124">Figs 3–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A7CC92626B83DFF38D06F0420A125" pageId="8" pageNumber="124" type="diagnosis">
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<heading id="D0B7982E2626B83DFF38D06F0405A627" bold="true" box="[189,310,794,820]" fontSize="11" level="3" pageId="8" pageNumber="124" reason="3">
<emphasis id="B934F3502626B83DFF38D06F0405A627" bold="true" box="[189,310,794,820]" pageId="8" pageNumber="124">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422626B83DFF38D03D0420A125" blockId="8.[188,1400,840,1078]" pageId="8" pageNumber="124">Head subquadrate; lateral margins of head subparallel in full face view; clypeus with longitudinal carinae extending only in the anterior half; scapes not reaching occipital head margin; in lateral view promesonotum convex, followed by a concave mesopropodeal depression at the junction with the propodeum; promesonotal suture visible in lateral view only; metanotal groove absent; propodeal spines well-developed with thick base, long and curved pointing backwards; head and mesosoma glabrate; head, mesosoma and gaster covered with scarce erect, stout setae. Core body concolorous ochreous-yellow.</paragraph>
</subSubSection>
<subSubSection id="C35A7CC92626B83DFF38D728002FA1FF" pageId="8" pageNumber="124" type="etymology">
<paragraph id="8BFF2F422626B83DFF38D7280477A164" blockId="8.[189,324,1117,1143]" box="[189,324,1117,1143]" pageId="8" pageNumber="124">
<heading id="D0B7982E2626B83DFF38D7280477A164" bold="true" box="[189,324,1117,1143]" fontSize="11" level="3" pageId="8" pageNumber="124" reason="3">
<emphasis id="B934F3502626B83DFF38D7280477A164" bold="true" box="[189,324,1117,1143]" pageId="8" pageNumber="124">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422626B83DFF38D7F9002FA1FF" blockId="8.[189,1399,1164,1260]" pageId="8" pageNumber="124">
The specific epithet ‘haveni’ is a noun in English. This was the first species of
<taxonomicName id="4C4054C12626B83DFBD4D7F901D8A1B5" authorityName="Mayr" authorityYear="1861" box="[1105,1259,1164,1190]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B934F3502626B83DFBD4D7F901D8A1B5" box="[1105,1259,1164,1190]" italics="true" pageId="8" pageNumber="124">Temnothorax</emphasis>
</taxonomicName>
recorded in
<collectingCountry id="F3576FD22626B83DFF38D7DA047AA1DA" box="[189,329,1199,1225]" name="Hong Kong" pageId="8" pageNumber="124">Hong Kong</collectingCountry>
, one of the most urbanized cities but with 40% of the land designated as protected areas for biodiversity conservation, providing havens for countless species and those awaiting discovery.
</paragraph>
</subSubSection>
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<heading id="D0B7982E2626B83DFF38D6660499A03E" bold="true" box="[189,426,1299,1325]" fontSize="11" level="3" pageId="8" pageNumber="124" reason="3">
<emphasis id="B934F3502626B83DFF38D6660499A03E" bold="true" box="[189,426,1299,1325]" pageId="8" pageNumber="124">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422626B83DFF6FD637047AA0D5" blockId="8.[189,1399,1346,1478]" pageId="8" pageNumber="124">
<materialsCitation id="3B28251F2626B83DFF6FD637047AA0D5" ID-GBIF-Occurrence="4161170302" collectingDate="2017-06-27" collectionCode="ZRC" collectorName="Roger H. Lee &amp; Yuet Yin Ling" country="China" county="Aberdeen Reservoir" elevation="190" latitude="22.2589" location="Hong Kong" longLatPrecision="1" longitude="114.1596" municipality="Hong Kong Island" pageId="8" pageNumber="124" specimenCode="RHL03433" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54FB91E02626B83DFF6FD637046FA04F" box="[234,348,1346,1372]" pageId="8" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3576FD22626B83DFF38D610042BA06C" box="[189,280,1381,1407]" name="China" pageId="8" pageNumber="124">CHINA</collectingCountry>
• worker;
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SAR,
<collectingMunicipality id="6B9BB5382626B83DFD24D61306BDA06C" box="[673,910,1381,1408]" pageId="8" pageNumber="124">Hong Kong Island</collectingMunicipality>
,
<collectingCounty id="629E57CE2626B83DFC24D61001AAA06C" box="[929,1177,1381,1407]" pageId="8" pageNumber="124">Aberdeen Reservoir</collectingCounty>
;
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,
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;
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<quantity id="4CB882A72626B83DFE05D6FD04F8A0B0" box="[384,459,1416,1443]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="124" unit="m" value="190.0">190 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFD9CD6FD0787A0B1" box="[537,692,1416,1443]" pageId="8" pageNumber="124" value="2017-06-27">
<collectingDate id="EFBAF06A2626B83DFD9CD6FD0787A0B1" box="[537,692,1416,1443]" pageId="8" pageNumber="124" value="2017-06-27">27 Jun. 2017</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFD45D6FC0653A0B0" box="[704,864,1417,1443]" pageId="8" pageNumber="124">Roger H. Lee</collectorName>
and
<collectorName id="26B54A942626B83DFC1CD6FC010DA0B1" box="[921,1086,1416,1443]" pageId="8" pageNumber="124">Yuet Yin Ling</collectorName>
leg.; ground baiting;
<collectionCode id="ED51B7872626B83DFAB8D6FD0045A0B1" box="[1341,1398,1416,1442]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="124">ZRC</collectionCode>
<specimenCode id="DBE687392626B83DFF38D6D90476A0D5" box="[189,325,1452,1478]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="124">RHL03433</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BFF2F422626B83CFF6FD698070EA47A" blockId="8.[186,1399,1517,2001]" lastBlockId="9.[189,1399,265,362]" lastPageId="9" lastPageNumber="125" pageId="8" pageNumber="124">
<materialsCitation id="3B28251F2626B83DFF6FD698044BA362" ID-GBIF-Occurrence="4161170303" collectingDate="2022-08-15" collectionCode="HKBM" collectorName="Andre Ibanez &amp; Matthew T. Hamer" country="China" elevation="38" latitude="22.2616" location="Hong Kong" longLatPrecision="1" longitude="114.0019" municipality="Mui Wo" pageId="8" pageNumber="124" specimenCode="ANTWEB1010974" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
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<typeStatus id="54FB91E02626B83DFF6FD6980454A314" box="[234,359,1517,1543]" pageId="8" pageNumber="124" type="paratype">Paratypes</typeStatus>
</emphasis>
(n = 8)
<collectingCountry id="F3576FD22626B83DFF38D565042BA339" box="[189,280,1552,1578]" name="China" pageId="8" pageNumber="124">CHINA</collectingCountry>
–
<emphasis id="B934F3502626B83DFEC5D565072DA339" bold="true" box="[320,542,1552,1578]" pageId="8" pageNumber="124">
<collectingRegion id="4984E1A02626B83DFEC5D56504E6A339" box="[320,469,1552,1578]" pageId="8" pageNumber="124">Hong Kong</collectingRegion>
SAR
</emphasis>
•
<specimenCount id="9D46E4CB2626B83DFDC4D5650780A339" box="[577,691,1552,1578]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382626B83DFD41D565061AA338" box="[708,809,1552,1579]" pageId="8" pageNumber="124">Mui Wo</collectingMunicipality>
(
<location id="8E9F79992626B83DFCBBD56506E9A339" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFCBBD56506E9A339" box="[830,986,1552,1578]" country="China" latitude="22.2616" longLatPrecision="1" longitude="114.0019" municipality="Mui Wo" name="Lantau Trail" pageId="8" pageNumber="124" stateProvince="Hong Kong">Lantau Trail</location>
);
<geoCoordinate id="EE7449852626B83DFC71D57A0180A338" box="[1012,1203,1551,1579]" degrees="22" direction="north" minutes="15" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="41.76" value="22.2616">22°15′41.76″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFB41D57A0042A338" box="[1220,1393,1551,1579]" degrees="114" direction="east" minutes="0" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="6.84" value="114.0019">114°0′6.84″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFF38D5410407A35E" box="[189,308,1587,1614]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="124" unit="m" value="38.0">
<quantity id="4CB882A72626B83DFF38D54105C5A35D" box="[189,246,1588,1614]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="124" unit="m" value="38.0">38 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFEBAD54604EFA35D" box="[319,476,1587,1614]" pageId="8" pageNumber="124" value="2022-08-15">
<collectingDate id="EFBAF06A2626B83DFEBAD54604EFA35D" box="[319,476,1587,1614]" pageId="8" pageNumber="124" value="2022-08-15">15 Aug. 2022</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFE60D54607B2A35E" box="[485,641,1587,1613]" pageId="8" pageNumber="124">André Ibáñez</collectorName>
and
<collectorName id="26B54A942626B83DFD30D54606BDA35D" box="[693,910,1587,1614]" pageId="8" pageNumber="124">Matthew T. Hamer</collectorName>
leg.;
<collectorName id="26B54A942626B83DFC4ED546011DA35E" box="[971,1070,1587,1613]" pageId="8" pageNumber="124">Winkler</collectorName>
;
<collectionCode id="ED51B7872626B83DFBB2D54101BCA35D" box="[1079,1167,1588,1614]" pageId="8" pageNumber="124">HKBM</collectionCode>
<specimenCode id="DBE687392626B83DFB17D5410044A35D" box="[1170,1399,1588,1614]" collectionCode="HKBM" pageId="8" pageNumber="124">ANTWEB1010974</specimenCode>
[MW1T1W4-5]
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83DFE18D5220044A387" ID-GBIF-Occurrence="4161170306" collectingDate="2022-04-14" collectionCode="HKBM" collectorName="Matthew T. Hamer" country="China" elevation="231" latitude="22.2793" location="Lung Fu Shan" longLatPrecision="1" longitude="114.1372" municipality="Hong Kong Island" pageId="8" pageNumber="124" specimenCode="ANTWEB1010976" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFE18D5220723A362" box="[413,528,1623,1649]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382626B83DFDA7D5220639A362" box="[546,778,1623,1649]" pageId="8" pageNumber="124">Hong Kong Island</collectingMunicipality>
,
<location id="8E9F79992626B83DFC9ED52206E2A362" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFC9ED52206E2A362" box="[795,977,1623,1649]" country="China" latitude="22.2793" longLatPrecision="1" longitude="114.1372" municipality="Hong Kong Island" name="Lung Fu Shan" pageId="8" pageNumber="124" stateProvince="Hong Kong">Lung Fu Shan</location>
;
<geoCoordinate id="EE7449852626B83DFC66D5200190A362" box="[995,1187,1621,1649]" degrees="22" direction="north" minutes="16" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="45.48" value="22.2793">22°16′45.48″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFB30D5200042A362" box="[1205,1393,1621,1649]" degrees="114" direction="east" minutes="8" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="13.92" value="114.1372">114°8′13.92″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFF38D50F0471A387" box="[189,322,1658,1684]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="8" pageNumber="124" unit="m" value="231.0">
<quantity id="4CB882A72626B83DFF38D50F0436A387" box="[189,261,1658,1684]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="8" pageNumber="124" unit="m" value="231.0">231 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFEC9D50F04D1A387" box="[332,482,1658,1684]" pageId="8" pageNumber="124" value="2022-04-14">
<collectingDate id="EFBAF06A2626B83DFEC9D50F04D1A387" box="[332,482,1658,1684]" pageId="8" pageNumber="124" value="2022-04-14">14Apr. 2022</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFE69D50F07F7A387" box="[492,708,1658,1684]" pageId="8" pageNumber="124">Matthew T. Hamer</collectorName>
leg.; hand collection on ground;
<collectionCode id="ED51B7872626B83DFBB2D50F01BCA387" box="[1079,1167,1658,1684]" pageId="8" pageNumber="124">HKBM</collectionCode>
<specimenCode id="DBE687392626B83DFB17D50F0044A387" box="[1170,1399,1658,1684]" collectionCode="HKBM" pageId="8" pageNumber="124">ANTWEB1010976</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83DFF55D5E80150A3C8" ID-GBIF-Occurrence="4161170309" collectingDate="2017-06-27" collectionCode="HKBM" collectorName="Roger H. Lee" country="China" elevation="192" latitude="22.2589" location="Aberdeen Reservoir" longLatPrecision="1" longitude="114.1595" municipality="Hong Kong Island" pageId="8" pageNumber="124" specimenCode="RHL03467" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFF55D5E80470A3A4" box="[208,323,1693,1719]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382626B83DFED6D5EB0705A3A4" box="[339,566,1693,1720]" pageId="8" pageNumber="124">Hong Kong Island</collectingMunicipality>
,
<location id="8E9F79992626B83DFDC1D5E80604A3A4" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFDC1D5E80604A3A4" box="[580,823,1693,1719]" country="China" latitude="22.2589" longLatPrecision="1" longitude="114.1595" municipality="Hong Kong Island" name="Aberdeen Reservoir" pageId="8" pageNumber="124" stateProvince="Hong Kong">Aberdeen Reservoir</location>
;
<geoCoordinate id="EE7449852626B83DFCC3D5EE0136A3AB" box="[838,1029,1691,1720]" degrees="22" direction="north" minutes="15" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="32.04" value="22.2589">22°15′32.04″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFB91D5EE01FDA3AB" box="[1044,1230,1691,1720]" degrees="114" direction="east" minutes="9" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="34.20" value="114.1595">114°9′34.20″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFB5AD5E80043A3A4" box="[1247,1392,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="1.92" pageId="8" pageNumber="124" unit="m" value="192.0">
<quantity id="4CB882A72626B83DFB5AD5E8001EA3AB" box="[1247,1325,1693,1720]" metricMagnitude="2" metricUnit="m" metricValue="1.92" pageId="8" pageNumber="124" unit="m" value="192.0">192 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFF38D5B50467A3C9" box="[189,340,1728,1755]" pageId="8" pageNumber="124" value="2017-06-27">
<collectingDate id="EFBAF06A2626B83DFF38D5B50467A3C9" box="[189,340,1728,1755]" pageId="8" pageNumber="124" value="2017-06-27">27 Jun. 2017</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFEDAD5B404C8A3C8" box="[351,507,1729,1755]" pageId="8" pageNumber="124">Roger H. Lee</collectorName>
leg.; ground baiting;
<collectionCode id="ED51B7872626B83DFD77D5B40679A3C8" box="[754,842,1729,1755]" pageId="8" pageNumber="124">HKBM</collectionCode>
<specimenCode id="DBE687392626B83DFCD5D5B506E7A3C9" box="[848,980,1728,1754]" collectionCode="HKBM" pageId="8" pageNumber="124">RHL03467</specimenCode>
[RHL5265]
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83DFBFCD5B5078FA232" ID-GBIF-Occurrence="4161170307" collectingDate="2017-07-11" collectionCode="HKBM" collectorName="Roger H. Lee" country="China" elevation="391" latitude="22.2736" location="The Peak" longLatPrecision="1" longitude="114.1389" municipality="Hong Kong Island" pageId="8" pageNumber="124" specimenCode="RHL03474" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFBFCD5B501D7A3C9" box="[1145,1252,1728,1754]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382626B83DFB6BD5B4043BA3EE" pageId="8" pageNumber="124">Hong Kong Island</collectingMunicipality>
,
<location id="8E9F79992626B83DFE91D59604B5A3EE" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFE91D59604B5A3EE" box="[276,390,1763,1789]" country="China" latitude="22.2736" longLatPrecision="1" longitude="114.1389" municipality="Hong Kong Island" name="The Peak" pageId="8" pageNumber="124" stateProvince="Hong Kong">The Peak</location>
;
<geoCoordinate id="EE7449852626B83DFE16D597077DA3ED" box="[403,590,1762,1790]" degrees="22" direction="north" minutes="16" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="24.96" value="22.2736">22°16′24.96″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFDDFD5970622A3ED" box="[602,785,1762,1790]" degrees="114" direction="east" minutes="8" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="20.04" value="114.1389">114°8′20.04″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFC9AD5910699A3EE" box="[799,938,1763,1790]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="8" pageNumber="124" unit="m" value="391.0">
<quantity id="4CB882A72626B83DFC9AD5910659A3ED" box="[799,874,1764,1790]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="8" pageNumber="124" unit="m" value="391.0">391 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFC3CD591017EA3ED" box="[953,1101,1763,1790]" pageId="8" pageNumber="124" value="2017-07-11">
<collectingDate id="EFBAF06A2626B83DFC3CD591017EA3ED" box="[953,1101,1763,1790]" pageId="8" pageNumber="124" value="2017-07-11">11 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFBDCD59101C9A3ED" box="[1113,1274,1764,1790]" pageId="8" pageNumber="124">Roger H. Lee</collectorName>
leg.; hand collection;
<collectionCode id="ED51B7872626B83DFEC6D47204A8A232" box="[323,411,1799,1825]" pageId="8" pageNumber="124">HKBM</collectionCode>
<specimenCode id="DBE687392626B83DFE20D472071AA232" box="[421,553,1799,1825]" collectionCode="HKBM" pageId="8" pageNumber="124">RHL03474</specimenCode>
[RHL5272]
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83DFD5FD47207ADA274" ID-GBIF-Occurrence="4161170304" collectingDate="2022-08-23" collectionCode="IBBL" collectorName="Shaolin Han" country="China" elevation="349" latitude="22.420198" location="Tai Po Kau Nature Reserve" longLatPrecision="1" longitude="114.1766" pageId="8" pageNumber="124" specimenCode="ANTWEB1010993" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFD5FD4720679A232" box="[730,842,1799,1825]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<location id="8E9F79992626B83DFCD2D4720195A232" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFCD2D4720195A232" box="[855,1190,1799,1825]" country="China" latitude="22.420198" longLatPrecision="1" longitude="114.1766" name="Tai Po Kau Nature Reserve" pageId="8" pageNumber="124" stateProvince="Hong Kong">Tai Po Kau Nature Reserve</location>
;
<geoCoordinate id="EE7449852626B83DFB30D4700041A232" box="[1205,1394,1797,1825]" degrees="22" direction="north" minutes="25" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="12.72" value="22.420198">22°25′12.72″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFF38D45D04B4A257" box="[189,391,1832,1860]" degrees="114" direction="east" minutes="10" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="35.76" value="114.1766">114°10′35.76″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFE1CD45F071DA257" box="[409,558,1834,1860]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="8" pageNumber="124" unit="m" value="349.0">
<quantity id="4CB882A72626B83DFE1CD45F04DAA257" box="[409,489,1834,1860]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="8" pageNumber="124" unit="m" value="349.0">349 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFDC4D45F07DDA257" box="[577,750,1834,1860]" pageId="8" pageNumber="124" value="2022-08-23">
<collectingDate id="EFBAF06A2626B83DFDC4D45F07DDA257" box="[577,750,1834,1860]" pageId="8" pageNumber="124" value="2022-08-23">23 Aug. 2022</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFD7AD45F06A5A257" box="[767,918,1834,1860]" pageId="8" pageNumber="124">Shaolin Han</collectorName>
leg.; arboreal baiting, 20 meters high;
<collectionCode id="ED51B7872626B83DFF38D43B05CCA27B" box="[189,255,1870,1896]" pageId="8" pageNumber="124">IBBL</collectionCode>
<specimenCode id="DBE687392626B83DFE8DD43804DFA274" box="[264,492,1869,1895]" collectionCode="IBBL" pageId="8" pageNumber="124">ANTWEB1010993</specimenCode>
[TPK_S1_T3]
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83DFD3BD4380791A2BD" ID-GBIF-Occurrence="4161170310" collectingDate="2022-05-23" collectionCode="IBBL" collectorName="Matthew T. Hamer &amp; Andre Ibanez" country="China" elevation="48" latitude="22.4904" location="Fanling Golf Course" longLatPrecision="1" longitude="114.1103" municipality="Fanling" pageId="8" pageNumber="124" specimenCode="ANTWEB1010975" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFD3BD438061CA274" box="[702,815,1869,1895]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382626B83DFCBBD43806A8A274" box="[830,923,1869,1895]" pageId="8" pageNumber="124">Fanling</collectingMunicipality>
,
<location id="8E9F79992626B83DFC2CD4380196A274" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFC2CD4380196A274" box="[937,1189,1869,1895]" country="China" latitude="22.4904" longLatPrecision="1" longitude="114.1103" municipality="Fanling" name="Fanling Golf Course" pageId="8" pageNumber="124" stateProvince="Hong Kong">Fanling Golf Course</location>
;
<geoCoordinate id="EE7449852626B83DFB31D43E0041A27B" box="[1204,1394,1867,1896]" degrees="22" direction="north" minutes="29" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="25.44" value="22.4904">22°29′25.44″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFF38D41A0446A298" box="[189,373,1903,1931]" degrees="114" direction="east" minutes="6" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="37.08" value="114.1103">114°6′37.08″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFE06D4050732A299" box="[387,513,1904,1931]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="124" unit="m" value="48.0">
<quantity id="4CB882A72626B83DFE06D40504F3A298" box="[387,448,1904,1931]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="124" unit="m" value="48.0">48 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFD95D4050780A299" box="[528,691,1904,1931]" pageId="8" pageNumber="124" value="2022-05-23">
<collectingDate id="EFBAF06A2626B83DFD95D4050780A299" box="[528,691,1904,1931]" pageId="8" pageNumber="124" value="2022-05-23">23 May 2022</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFD44D4050692A298" box="[705,929,1904,1931]" pageId="8" pageNumber="124">Matthew T. Hamer</collectorName>
and
<collectorName id="26B54A942626B83DFC5ED405014FA299" box="[987,1148,1904,1930]" pageId="8" pageNumber="124">André Ibáñez</collectorName>
leg.; ground baiting;
<collectionCode id="ED51B7872626B83DFF38D4E105CCA2BD" box="[189,255,1940,1966]" pageId="8" pageNumber="124">IBBL</collectionCode>
<specimenCode id="DBE687392626B83DFE86D4E604DBA2BD" box="[259,488,1939,1966]" collectionCode="IBBL" pageId="8" pageNumber="124">ANTWEB1010975</specimenCode>
[FGE1T2B2-1]
</materialsCitation>
•
<materialsCitation id="3B28251F2626B83CFD3FD4E1068BA430" ID-GBIF-Occurrence="4161170308" collectingDate="2022-06-24" collectingDateMax="2022-07-01" collectingDateMin="2022-06-24" collectionCode="IBBL" collectorName="Matthew T. Hamer &amp; Andre Ibanez" country="China" elevation="84" lastPageId="9" lastPageNumber="125" latitude="22.3713" location="Girl Guides Pok Hong Campsite" longLatPrecision="1" longitude="114.196304" pageId="8" pageNumber="124" specimenCode="ANTWEB1010987" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2626B83DFD3FD4E10615A2BE" box="[698,806,1939,1966]" pageId="8" pageNumber="124" type="worker">1 worker</specimenCount>
;
<location id="8E9F79992626B83DFCB5D4E6019DA2BE" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992626B83DFCB5D4E6019DA2BE" box="[816,1198,1939,1966]" country="China" latitude="22.3713" longLatPrecision="1" longitude="114.196304" name="Girl Guides Pok Hong Campsite" pageId="8" pageNumber="124" stateProvince="Hong Kong">Girl Guides Pok Hong Campsite</location>
;
<geoCoordinate id="EE7449852626B83DFB3CD4E70041A2BD" box="[1209,1394,1938,1966]" degrees="22" direction="north" minutes="22" orientation="latitude" pageId="8" pageNumber="124" precision="1" seconds="16.68" value="22.3713">22°22′16.68″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852626B83DFF38D4C004B7A2C2" box="[189,388,1973,2001]" degrees="114" direction="east" minutes="11" orientation="longitude" pageId="8" pageNumber="124" precision="1" seconds="46.68" value="114.196304">114°11′46.68″ E</geoCoordinate>
;
<elevation id="006DC8712626B83DFE11D4C20724A2C2" box="[404,535,1975,2001]" metricMagnitude="1" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="124" unit="m" value="84.0">
<quantity id="4CB882A72626B83DFE11D4C204E0A2C2" box="[404,467,1975,2001]" metricMagnitude="1" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="124" unit="m" value="84.0">84 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822626B83DFDADD4C20628A2C2" box="[552,795,1975,2001]" pageId="8" pageNumber="124" value="2022-06-24" valueMax="2022-07-01" valueMin="2022-06-24">
<collectingDate id="EFBAF06A2626B83DFDADD4C20628A2C2" box="[552,795,1975,2001]" pageId="8" pageNumber="124" value="2022-06-24" valueMax="2022-07-01" valueMin="2022-06-24">24 Jun.–1 Jul. 2022</collectingDate>
</date>
;
<collectorName id="26B54A942626B83DFCAFD4C2013CA2C2" box="[810,1039,1975,2001]" pageId="8" pageNumber="124">Matthew T. Hamer</collectorName>
and
<collectorName id="26B54A942626B83DFBC8D4C201DCA2C2" box="[1101,1263,1975,2001]" pageId="8" pageNumber="124">André Ibáñez</collectorName>
leg.;
<collectingMethod id="520157552626B83CFABDD4C204F7A430" lastPageId="9" lastPageNumber="125" pageId="8" pageNumber="124">flight interception vane trap</collectingMethod>
;
<collectionCode id="ED51B7872627B83CFE56D27C0726A430" box="[467,533,265,291]" pageId="9" pageNumber="125">IBBL</collectionCode>
<specimenCode id="DBE687392627B83CFD99D27C0633A430" box="[540,768,265,291]" collectionCode="IBBL" pageId="9" pageNumber="125">ANTWEB1010987</specimenCode>
[GGPH2V2-1]
</materialsCitation>
•
<materialsCitation id="3B28251F2627B83CFC50D27C070BA47A" ID-GBIF-Occurrence="4161170301" collectingDate="2018-03-16" collectionCode="IBBL" collectorName="R. Cheung &amp; M. Law" country="China" elevation="244" latitude="22.343102" location="Tsing Yi Peak" longLatPrecision="1" longitude="114.0999" municipality="Tsing Yi" pageId="9" pageNumber="125" specimenCode="ANTWEB1016704" specimenCount="1" specimenCount-worker="1" stateProvince="Hong Kong" typeStatus="paratype">
<specimenCount id="9D46E4CB2627B83CFC50D27C0176A430" box="[981,1093,265,291]" pageId="9" pageNumber="125" type="worker">1 worker</specimenCount>
;
<collectingMunicipality id="6B9BB5382627B83CFBD7D27C0189A430" box="[1106,1210,265,291]" pageId="9" pageNumber="125">Tsing Yi</collectingMunicipality>
,
<location id="8E9F79992627B83CFB42D27C0041A430" LSID="urn:lsid:plazi:treatment:03E99E542626B838FE58D1E701D0A7B1:8E9F79992627B83CFB42D27C0041A430" box="[1223,1394,265,291]" country="China" latitude="22.343102" longLatPrecision="1" longitude="114.0999" municipality="Tsing Yi" name="Tsing Yi Peak" pageId="9" pageNumber="125" stateProvince="Hong Kong">Tsing Yi Peak</location>
;
<geoCoordinate id="EE7449852627B83CFF38D25E0445A454" box="[189,374,299,327]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="125" precision="1" seconds="35.16" value="22.343102">22°20′35.16″ N</geoCoordinate>
,
<geoCoordinate id="EE7449852627B83CFE05D25E0706A454" box="[384,565,299,327]" degrees="114" direction="east" minutes="5" orientation="longitude" pageId="9" pageNumber="125" precision="1" seconds="59.64" value="114.0999">114°5′59.64″ E</geoCoordinate>
;
<elevation id="006DC8712627B83CFDC5D25907F4A455" box="[576,711,300,327]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="9" pageNumber="125" unit="m" value="244.0">
<quantity id="4CB882A72627B83CFDC5D25907BAA454" box="[576,649,300,327]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="9" pageNumber="125" unit="m" value="244.0">244 m</quantity>
a.s.l.
</elevation>
;
<date id="FFFE09822627B83CFD56D2590642A455" box="[723,881,300,327]" pageId="9" pageNumber="125" value="2018-03-16">
<collectingDate id="EFBAF06A2627B83CFD56D2590642A455" box="[723,881,300,327]" pageId="9" pageNumber="125" value="2018-03-16">16 Mar. 2018</collectingDate>
</date>
;
<collectorName id="26B54A942627B83CFCF9D25806C4A455" box="[892,1015,300,327]" pageId="9" pageNumber="125">R. Cheung</collectorName>
and
<collectorName id="26B54A942627B83CFBABD25801BAA454" box="[1070,1161,301,327]" pageId="9" pageNumber="125">M. Law</collectorName>
leg.;
<collectorName id="26B54A942627B83CFB4DD2590018A455" box="[1224,1323,300,326]" pageId="9" pageNumber="125">Winkler</collectorName>
;
<collectionCode id="ED51B7872627B83CFAB0D2580044A454" box="[1333,1399,301,327]" pageId="9" pageNumber="125">IBBL</collectionCode>
<specimenCode id="DBE687392627B83CFF38D23A0492A47A" box="[189,417,335,361]" collectionCode="IBBL" pageId="9" pageNumber="125">ANTWEB1016704</specimenCode>
[TYP S1-R]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A7CC92627B83EFF38D2E506ADA76A" lastPageId="11" lastPageNumber="127" pageId="9" pageNumber="125" type="description">
<paragraph id="8BFF2F422627B83CFF38D2E5047DA4B9" blockId="9.[189,334,400,426]" box="[189,334,400,426]" pageId="9" pageNumber="125">
<heading id="D0B7982E2627B83CFF38D2E5047DA4B9" bold="true" box="[189,334,400,426]" fontSize="11" level="3" pageId="9" pageNumber="125" reason="3">
<emphasis id="B934F3502627B83CFF38D2E5047DA4B9" bold="true" box="[189,334,400,426]" pageId="9" pageNumber="125">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422627B83CFF38D2B505CBA7C3" blockId="9.[188,1399,447,720]" pageId="9" pageNumber="125">
<smallCapsWord id="8D19B99E2627B83CFF38D2B5045DA4CB" baselines="467,467" box="[189,366,448,474]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="9" pageNumber="125">MEASUREMENTS</smallCapsWord>
(n = 9).
<typeStatus id="54FB91E02627B83CFE53D2CA0770A4CA" box="[470,579,447,473]" pageId="9" pageNumber="125" type="holotype">Holotype</typeStatus>
(n = 1); CL 0.55; CW 0.54; CWb 0.50; SL 0.4; WL 0.65; SPST 0.23; PEL 0.29; PPL 0.15; PEH 0.16; PPH 0.16; PW 0.34; SPBA 0.12; SPTI 0.17; PEW 0.13; PPW 0.18; ATL 0.62; HS 0.52; SI 80.6; CI 91.58; SBI 23.2;
<collectingRegion id="4984E1A02627B83CFD7DD1730611A733" box="[760,802,518,544]" country="Philippines" name="Pasay" pageId="9" pageNumber="125">PSI</collectingRegion>
35.02; PWI 145.6; PLI 194.63; TL 2.26.
<typeStatus id="54FB91E02627B83CFA81D1730045A733" box="[1284,1398,518,544]" pageId="9" pageNumber="125" type="paratype">Paratypes</typeStatus>
(n = 8); HL 0.51–0.55; CW 0.5–0.55; CWb 0.44–0.5; SL 0.35–0.41; WL 0.61–0.69; SPST 0.23–0.29; PEL 0.25–0.32; PPL 0.12–0.17; PEH 0.14–0.18; PPH 0.15–0.16; PW 0.3–0.34; SPBA 0.1–0.17; SPTI 0.11–0.18; PEW 0.11–0.12; PPW 0.16–0.18; ATL 0.46–0.58; HS 0.47–0.53; SI 75.71–87.67; CI 86.22–92.04; SBI 21.1–37.66;
<collectingRegion id="4984E1A02627B83CFDDDD1E607B1A7BE" box="[600,642,659,685]" country="Philippines" name="Pasay" pageId="9" pageNumber="125">PSI</collectingRegion>
36.99–41.84; PWI 140.16–151.35; PLI 160–229.41; TL 1.99– 2.22.
</paragraph>
<paragraph id="8BFF2F422627B83CFF38D18D0173A15D" blockId="9.[189,1400,759,1103]" pageId="9" pageNumber="125">
<smallCapsWord id="8D19B99E2627B83CFF38D18D05CFA603" baselines="779,779" box="[189,252,760,786]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="9" pageNumber="125">HEAD</smallCapsWord>
. In full face view, head subquadrate, longer than broad (CI 86.22–92.04), with weakly convex sides and occipital margin, rounded occipital corners. Clypeus widely inserted between antennal lobes; anterior margin weakly convex and angulate medially; three clypeal carinae present. Mandible broadly triangular, masticatory margin with five teeth, apical most tooth larger than preceding teeth. Frontal carinae weak but moderately long, extends from the antennal insertions to the area of the vertex; frontal lobes present. Antenna with 12 segments terminating in an incrassate three-segmented club; apical segment longer and broader than following segments. Scape of medium length (SI 78.57–87.67), terminating before posterior corners of the head. Eye convex; located medially on head and extending laterally beyond the cephalic capsule. In lateral view, eyes composed of 9–10 ommatidia across the longest width. In dorsal view, occipital carina present but weakly developed.
</paragraph>
<paragraph id="8BFF2F422627B83CFF38D70307C6A095" blockId="9.[187,1400,1141,1414]" pageId="9" pageNumber="125">
<smallCapsWord id="8D19B99E2627B83CFF38D7030409A19D" baselines="1161,1161" box="[189,314,1142,1168]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesosoma" pageId="9" pageNumber="125">MESOSOMA</smallCapsWord>
. In dorsal view, mesosoma widest at the middle portion of pronotum; humeri widely rounded; mesosoma weakly tapering posteriorly, reaching a minimum width at the anterior part to propodeum. Promesonotal suture absent dorsally but present laterally. Metanotal groove absent. In lateral view, promesonotum convex and follow by a distinct concavity at mesonotum, forming a weak promesonotal dome. Propodeal spiracle circular. Propodeal lobe round. Propodeal spines well-developed, long, and slightly downward curved toward the end; spines longer than the distance between their bases (SPST 0.23–0.29; SBPA 0.10–0.17), spines feebly diverge postero-laterally from the dorsal view. In lateral view, propodeal declivity subtly concave.
</paragraph>
<paragraph id="8BFF2F422627B83CFF38D6DB066FA322" blockId="9.[187,1399,1453,1585]" pageId="9" pageNumber="125">
<smallCapsWord id="8D19B99E2627B83CFF38D6DB040AA0D5" baselines="1473,1473" box="[189,313,1454,1480]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metasoma" pageId="9" pageNumber="125">METASOMA</smallCapsWord>
. In lateral view, petiole subtriangular, longer than high. Anterior face of petiole distinctly longer than posterior face; node with acute apex. Postpetiole short and convex. In dorsal view, postpetiole subquadrate; distinctly wider than petiole. Gaster wider than postpetiole; first gastral tergite long, as long as mesosoma; anterolateral corners obtusely angled.
</paragraph>
<paragraph id="8BFF2F422627B83EFF38D52D04FAA455" blockId="9.[188,1401,1624,1968]" lastBlockId="11.[189,1398,265,326]" lastPageId="11" lastPageNumber="127" pageId="9" pageNumber="125">
<smallCapsWord id="8D19B99E2627B83CFF38D52D0433A362" baselines="1643,1644" box="[189,256,1624,1650]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Setae" pageId="9" pageNumber="125">SETAE</smallCapsWord>
. In full face view, mandible dorsum with well-spaced sub-decumbent pilosity; anterior clypeal margin with two long and tapering setae on either side of clypeal median. Several sub-erect setae present on anterior clypeal margin, directed towards clypeal median. From clypeal dorsum to cephalic dorsum, covered with sparse, stout and erect setae that are spaced roughly equidistantly. Scapes and subsequent antennal segments with sub-decumbent to semi-erected pilosity. In lateral view, ventral part of head with scare erect to semi-erect pilosity, intertwined with stout-erect setae. Mesosoma with long, erect setae arranged in series of transverse rows, normally not more than five pairs. Mesosoma dorsum also with sparse, short decumbent and appressed setae between the long, erect, setae pairs; of propodeal spines also with a single pair of setae sub-apically. Posterior face of petiole dorsum with a few pairs of erect setae, anterior face lacking setae. Postpetiole dorsum with a few pairs of erect setae. Gastral tergite with scarce erect stout setae in varying length arranged in loose rows. Femur and tibia with short and appressed pubescence.
</paragraph>
<caption id="DF3F7FCA2624B83FFF38D4DE079AA2FB" ID-DOI="http://doi.org/10.5281/zenodo.8155433" ID-Zenodo-Dep="8155433" httpUri="https://zenodo.org/record/8155433/files/figure.png" pageId="10" pageNumber="126" startId="10.[189,232,1963,1989]" targetBox="[393,1194,265,1922]" targetPageId="10">
<paragraph id="8BFF2F422624B83FFF38D4DE079AA2FB" blockId="10.[189,1399,1963,2025]" pageId="10" pageNumber="126">
<emphasis id="B934F3502624B83FFF38D4DE043EA2D6" bold="true" box="[189,269,1963,1989]" pageId="10" pageNumber="126">Fig. 3.</emphasis>
<taxonomicName id="4C4054C12624B83FFE93D4DE0619A2D6" authority="Lee, Hamer &amp; Guenard" authorityName="Lee, Hamer &amp; Guenard" authorityYear="2023" box="[278,810,1963,1990]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="126" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502624B83FFE93D4DE0734A2D6" box="[278,519,1963,1989]" italics="true" pageId="10" pageNumber="126">Temnothorax haveni</emphasis>
Lee, Hamer &amp; Guénard
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2624B83FFCB6D4D906A2A2D5" box="[819,913,1964,1990]" pageId="10" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (RHL03433).
<emphasis id="B934F3502624B83FFB3DD4DE01FFA2D6" bold="true" box="[1208,1228,1963,1989]" pageId="10" pageNumber="126">A</emphasis>
. Lateral view.
<emphasis id="B934F3502624B83FFF3AD4BB05E2A2FB" bold="true" box="[191,209,1998,2024]" pageId="10" pageNumber="126">B</emphasis>
. Dorsal view.
<emphasis id="B934F3502624B83FFEFED4BB04A3A2FB" bold="true" box="[379,400,1998,2024]" pageId="10" pageNumber="126">C</emphasis>
. Head in full face view.
</paragraph>
</caption>
<paragraph id="8BFF2F422625B83EFF38D21B004BA707" blockId="11.[188,1400,366,533]" pageId="11" pageNumber="127">
<smallCapsWord id="8D19B99E2625B83EFF38D21B040BA495" baselines="385,385" box="[189,312,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sculpture" pageId="11" pageNumber="127">SCULPTURE</smallCapsWord>
. In full face view, mandibles overlain by very weak lateral striae. Majority of clypeus dorsum smooth other than short, longitudinal carinae that begin at the clypeal anterior border. Dorsum of head, from clypeus to posterior head corners, glabrate. Dorsal pronotum, mesonotum and propodeum glabrate. Pronotum, mesonotum and propodeum overlain with faint but weak lateral striae laterally. Petiole and postpetiole dorsally and laterally weekly punctate-recticulate. Gaster comparatively smooth and shining.
</paragraph>
<paragraph id="8BFF2F422625B83EFF38D14906ADA76A" blockId="11.[189,1399,572,633]" pageId="11" pageNumber="127">
<smallCapsWord id="8D19B99E2625B83EFF38D149042BA747" baselines="591,591" box="[189,280,572,598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="11" pageNumber="127">COLOUR</smallCapsWord>
. Core body concolorous ochreous-yellow. Setae across whole of body yellowish white. Gaster with dark brown patches laterally from dorsal and lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C35A7CC92625B839FF38D1D50769A7FF" lastPageId="12" lastPageNumber="128" pageId="11" pageNumber="127" type="discussion">
<paragraph id="8BFF2F422625B83EFF38D1D50477A7A9" blockId="11.[189,324,672,698]" box="[189,324,672,698]" pageId="11" pageNumber="127">
<heading id="D0B7982E2625B83EFF38D1D50477A7A9" bold="true" box="[189,324,672,698]" fontSize="11" level="3" pageId="11" pageNumber="127" reason="3">
<emphasis id="B934F3502625B83EFF38D1D50477A7A9" bold="true" box="[189,324,672,698]" pageId="11" pageNumber="127">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422625B839FF38D1BA0769A7FF" blockId="11.[188,1400,719,922]" lastBlockId="12.[188,1399,265,749]" lastPageId="12" lastPageNumber="128" pageId="11" pageNumber="127">
<taxonomicName id="4C4054C12625B83EFF38D1BA049FA7FA" authority="Hamer &amp; Lee &amp; Guénard, 2023" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[189,428,719,745]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="127" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502625B83EFF38D1BA049FA7FA" box="[189,428,719,745]" italics="true" pageId="11" pageNumber="127">Temnothorax haveni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2625B83EFE31D1A5073DA7F9" box="[436,526,720,746]" pageId="11" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
would key out to
<taxonomicName id="4C4054C12625B83EFD66D1A506ADA7FA" box="[739,926,719,745]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="127" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B934F3502625B83EFD66D1A506ADA7FA" box="[739,926,719,745]" italics="true" pageId="11" pageNumber="127">T. zhejiangensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFD152B32625B83EFC41D1BA01AAA7FA" author="Zhou S. &amp; Huang J. &amp; Yu D. &amp; Liu Z." box="[964,1177,719,745]" pageId="11" pageNumber="127" pagination="7 - 26" refId="ref10425" refString="Zhou S., Huang J., Yu D. &amp; Liu Z. 2010. Eight new species and three newly recorded species of the ant genus Temnothorax Mayr (Hymenoptera: Formicidae) from the Chinese mainland, with a key. Sociobiology 56 (1): 7 - 26." type="journal article" year="2010">
Zhou
<emphasis id="B934F3502625B83EFB8CD1A50177A7FA" box="[1033,1092,719,745]" italics="true" pageId="11" pageNumber="127">et al.</emphasis>
(2010)
</bibRefCitation>
and shares several morphological characters. These characters include the presence of erect setae on the mesosomal dorsum, a pair of long slightly downcurved propodeal spines, humeri rounded in dorsal view, a short petiole peduncle and a petiole that is longer than high in lateral view. However, various characters differ, including the sculpture on the head dorsum, lateral and dorsum mesosoma being predominately glabrate in
<taxonomicName id="4C4054C12625B83EFF5FD0F50477A68A" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[218,324,895,921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="127" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502625B83EFF5FD0F50477A68A" box="[218,324,895,921]" italics="true" pageId="11" pageNumber="127">T. haveni</emphasis>
</taxonomicName>
rather than punctate in
<taxonomicName id="4C4054C12625B83EFDD6D0F5063EA68A" box="[595,781,895,921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="127" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B934F3502625B83EFDD6D0F5063EA68A" box="[595,781,895,921]" italics="true" pageId="11" pageNumber="127">T. zhejiangensis</emphasis>
</taxonomicName>
. The mesosomal outline differs greatly between both species with a convex promesonotum followed by a distinct concavity forming a weak promesonotal dome in
<taxonomicName id="4C4054C12622B839FEAED25804A9A455" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[299,410,300,326]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FEAED25804A9A455" box="[299,410,300,326]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
but only slightly convex across its whole length in
<taxonomicName id="4C4054C12622B839FB98D25801EFA455" box="[1053,1244,300,326]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B934F3502622B839FB98D25801EFA455" box="[1053,1244,300,326]" italics="true" pageId="12" pageNumber="128">T. zhejiangensis</emphasis>
</taxonomicName>
. The petiole peduncle is narrower and slightly longer in
<taxonomicName id="4C4054C12622B839FD3FD2250617A47A" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[698,804,335,361]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FD3FD2250617A47A" box="[698,804,335,361]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C4054C12622B839FC06D225010DA47A" box="[899,1086,335,361]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B934F3502622B839FC06D225010DA47A" box="[899,1086,335,361]" italics="true" pageId="12" pageNumber="128">T. zhejiangensis</emphasis>
</taxonomicName>
being broader and shorter. The petiole node in
<taxonomicName id="4C4054C12622B839FE35D206072EA49F" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[432,541,370,396]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FE35D206072EA49F" box="[432,541,370,396]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
has an acute dorsal apex within
<taxonomicName id="4C4054C12622B839FC2DD2060157A49F" box="[936,1124,370,396]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B934F3502622B839FC2DD2060157A49F" box="[936,1124,370,396]" italics="true" pageId="12" pageNumber="128">T. zhejiangensis</emphasis>
</taxonomicName>
is subtriangular with a narrowly rounded dorsum. Similarly,
<taxonomicName id="4C4054C12622B839FDFCD2E207D6A4A3" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[633,741,406,432]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FDFCD2E207D6A4A3" box="[633,741,406,432]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
might be mistaken for
<taxonomicName id="4C4054C12622B839FC7CD2E201B9A4A3" box="[1017,1162,407,432]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="ruginosus">
<emphasis id="B934F3502622B839FC7CD2E201B9A4A3" box="[1017,1162,407,432]" italics="true" pageId="12" pageNumber="128">T. ruginosus</emphasis>
</taxonomicName>
both species can be differentiated by size (
<taxonomicName id="4C4054C12622B839FE46D2CF0753A4C0" authority="WL" authorityName="WL" box="[451,608,441,467]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FE46D2CF071EA4C0" box="[451,557,441,467]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
WL
</taxonomicName>
0.61–0.69;
<taxonomicName id="4C4054C12622B839FD6CD2CF069EA4C0" authority="WL" authorityName="WL" box="[745,941,441,467]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="ruginosus">
<emphasis id="B934F3502622B839FD6CD2CF0648A4C0" box="[745,891,442,467]" italics="true" pageId="12" pageNumber="128">T. ruginosus</emphasis>
WL
</taxonomicName>
(ML in
<bibRefCitation id="EFD152B32622B839FB89D2CC01D9A4C0" author="Zhou S. &amp; Huang J. &amp; Yu D. &amp; Liu Z." box="[1036,1258,441,467]" pageId="12" pageNumber="128" pagination="7 - 26" refId="ref10425" refString="Zhou S., Huang J., Yu D. &amp; Liu Z. 2010. Eight new species and three newly recorded species of the ant genus Temnothorax Mayr (Hymenoptera: Formicidae) from the Chinese mainland, with a key. Sociobiology 56 (1): 7 - 26." type="journal article" year="2010">
Zhou
<emphasis id="B934F3502622B839FBD5D2CF01B7A4C0" box="[1104,1156,441,467]" italics="true" pageId="12" pageNumber="128">et al</emphasis>
. (2010))
</bibRefCitation>
1.80–1.84), as well as head and mesosomal sculpturing and the mesosomal outline.
<taxonomicName id="4C4054C12622B839FC72D2A901D6A4E5" box="[1015,1253,476,502]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FC72D2A901D6A4E5" box="[1015,1253,476,502]" italics="true" pageId="12" pageNumber="128">Temnothorax haveni</emphasis>
</taxonomicName>
may also be mistaken for
<taxonomicName id="4C4054C12622B839FEDCD17504EDA70A" authority="Hamer &amp; Lee &amp; Guénard, 2023" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[345,478,511,537]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="barrettoi" status="sp. nov.">
<emphasis id="B934F3502622B839FEDCD17504EDA70A" box="[345,478,511,537]" italics="true" pageId="12" pageNumber="128">T. barrettoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2622B839FE63D1750770A709" box="[486,579,512,538]" pageId="12" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
, however, both species can be differentiated by the glabrate sculpture and more scarce erect stout setae over the body of
<emphasis id="B934F3502622B839FC8AD15606B1A72F" box="[783,898,546,572]" italics="true" pageId="12" pageNumber="128">
<taxonomicName id="4C4054C12622B839FC8AD156064EA72F" box="[783,893,546,572]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">T. haveni</taxonomicName>
.
</emphasis>
We believe the above characters distinctly differentiate
<taxonomicName id="4C4054C12622B839FED3D13204F0A773" box="[342,451,582,608]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FED3D13204F0A773" box="[342,451,582,608]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
from its congeneric species due to their uniqueness and consistency across all specimens examined. In fact, the lack of sculpture, particularly on the head, combined with distinct mesopropodeal depression, makes
<taxonomicName id="4C4054C12622B839FDE5D1F807FEA7B5" box="[608,717,652,678]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FDE5D1F807FEA7B5" box="[608,717,652,678]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
morphologically distinct amongst Chinese
<taxonomicName id="4C4054C12622B839FB5DD1F9005CA7B5" authorityName="Mayr" authorityYear="1861" box="[1240,1391,652,678]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B934F3502622B839FB5DD1F9005CA7B5" box="[1240,1391,652,678]" italics="true" pageId="12" pageNumber="128">Temnothorax</emphasis>
</taxonomicName>
. Further description of species of
<taxonomicName id="4C4054C12622B839FDBFD1DA07E7A7DA" authorityName="Mayr" authorityYear="1861" box="[570,724,687,713]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B934F3502622B839FDBFD1DA07E7A7DA" box="[570,724,687,713]" italics="true" pageId="12" pageNumber="128">Temnothorax</emphasis>
</taxonomicName>
from other regions of Southeast Asia were examined with no species satisfying all characters.
</paragraph>
</subSubSection>
<caption id="DF3F7FCA2625B83EFF39D4FD06FCA2FB" ID-DOI="http://doi.org/10.5281/zenodo.8155435" ID-Zenodo-Dep="8155435" httpUri="https://zenodo.org/record/8155435/files/figure.png" pageId="11" pageNumber="127" startId="11.[188,231,1928,1954]" targetBox="[189,1398,992,1886]" targetPageId="11">
<paragraph id="8BFF2F422625B83EFF39D4FD06FCA2FB" blockId="11.[188,1399,1928,2025]" pageId="11" pageNumber="127">
<emphasis id="B934F3502625B83EFF39D4FD043DA2B1" bold="true" box="[188,270,1928,1954]" pageId="11" pageNumber="127">Fig. 4.</emphasis>
<taxonomicName id="4C4054C12625B83EFE9FD4FD0605A2B1" authority="Lee, Hamer &amp; Guenard" authorityName="Lee, Hamer &amp; Guenard" authorityYear="2023" box="[282,822,1928,1954]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="127" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502625B83EFE9FD4FD073FA2B1" box="[282,524,1928,1954]" italics="true" pageId="11" pageNumber="127">Temnothorax haveni</emphasis>
Lee, Hamer &amp; Guénard
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2625B83EFCC4D4FD0692A2B1" box="[833,929,1928,1954]" pageId="11" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (RHL03433), morphological characters.
<emphasis id="B934F3502625B83EFECFD4DE046CA2D6" bold="true" box="[330,351,1963,1989]" pageId="11" pageNumber="127">A</emphasis>
. Petiole and postpetiole in lateral view.
<emphasis id="B934F3502625B83EFCECD4DE064EA2D6" bold="true" box="[873,893,1963,1989]" pageId="11" pageNumber="127">B</emphasis>
. Petiole and postpetiole in dorsal view.
<emphasis id="B934F3502625B83EFF38D4BB05E1A2FB" bold="true" box="[189,210,1998,2024]" pageId="11" pageNumber="127">C</emphasis>
. Mesosoma in dorsal view.
<emphasis id="B934F3502625B83EFD9FD4BB071CA2FB" bold="true" box="[538,559,1998,2024]" pageId="11" pageNumber="127">D</emphasis>
. First gastral tergite in dorsal view.
</paragraph>
</caption>
<subSubSection id="C35A7CC92622B838FF38D06101D0A7B1" lastPageId="13" lastPageNumber="129" pageId="12" pageNumber="128" type="biology_ecology">
<paragraph id="8BFF2F422622B839FF38D061044DA63D" blockId="12.[189,382,788,814]" box="[189,382,788,814]" pageId="12" pageNumber="128">
<heading id="D0B7982E2622B839FF38D061044DA63D" bold="true" box="[189,382,788,814]" fontSize="11" level="3" pageId="12" pageNumber="128" reason="3">
<emphasis id="B934F3502622B839FF38D061044DA63D" bold="true" box="[189,382,788,814]" pageId="12" pageNumber="128">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF2F422622B838FF38D03706BFA4E5" blockId="12.[187,1400,834,1002]" lastBlockId="13.[188,1399,265,503]" lastPageId="13" lastPageNumber="129" pageId="12" pageNumber="128">
<taxonomicName id="4C4054C12622B839FF38D037049CA64F" authority="Hamer &amp; Lee &amp; Guénard, 2023" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[189,431,834,860]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502622B839FF38D037049CA64F" box="[189,431,834,860]" italics="true" pageId="12" pageNumber="128">Temnothorax haveni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2622B839FE3FD0360724A64E" box="[442,535,835,861]" pageId="12" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
has been collected from semi-open to closed canopy secondary forests throughout the territory of
<collectingCountry id="F3576FD22622B839FE7DD01307B5A693" box="[504,646,870,896]" name="Hong Kong" pageId="12" pageNumber="128">Hong Kong</collectingCountry>
. Specimens are mostly found within leaf litter samples but have also been attracted to ground baiting. One worker of
<taxonomicName id="4C4054C12622B839FCBDD0FF0691A6B0" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[824,930,905,931]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502622B839FCBDD0FF0691A6B0" box="[824,930,905,931]" italics="true" pageId="12" pageNumber="128">T. haveni</emphasis>
</taxonomicName>
was collected twenty metres above the ground in a tree from an arboreal bait sample within a secondary forest. An additional specimen was hand collected along a waist high handrail and another from a flight interception trap (vane trap) hung from a tree at head height. These samples may indicate
<taxonomicName id="4C4054C12623B838FD7DD27F0650A430" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[760,867,265,291]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="129" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="B934F3502623B838FD7DD27F0650A430" box="[760,867,265,291]" italics="true" pageId="13" pageNumber="129">T. haveni</emphasis>
</taxonomicName>
forages on shrubs or understorey vegetation, as well as within trees and could therefore be a predominately arboreal species, which may also forage occasionally on the forest floor. However, it is difficult to rule out individuals falling from plants due to unintentional vegetation interaction by samplers, which may explain specimens from Winkler samples and ground hand collection. Moreover, the lack of any whole nest samples from the mostly ground based sampling effort in
<collectingCountry id="F3576FD22623B838FE14D2CF072AA4C7" box="[401,537,442,468]" name="Hong Kong" pageId="13" pageNumber="129">Hong Kong</collectingCountry>
(e.g., Winklers), indicates this species may not nest in leaf litter, with only singletons found and no reproductive caste thus far collected.
</paragraph>
<caption id="DF3F7FCA2622B839FF38D4DE07A3A2FA" ID-DOI="http://doi.org/10.5281/zenodo.8155437" ID-Zenodo-Dep="8155437" httpUri="https://zenodo.org/record/8155437/files/figure.png" pageId="12" pageNumber="128" startId="12.[189,232,1963,1989]" targetBox="[219,1391,1101,1915]" targetPageId="12">
<paragraph id="8BFF2F422622B839FF38D4DE07A3A2FA" blockId="12.[189,1398,1963,2025]" pageId="12" pageNumber="128">
<emphasis id="B934F3502622B839FF38D4DE043EA2D6" bold="true" box="[189,269,1963,1989]" pageId="12" pageNumber="128">Fig. 5.</emphasis>
Distribution map of
<taxonomicName id="4C4054C12622B839FD89D4DE0604A2D6" authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" box="[524,823,1963,1990]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B934F3502622B839FD89D4DE0795A2D6" box="[524,678,1963,1989]" italics="true" pageId="12" pageNumber="128">Temnothorax</emphasis>
Mayr, 1861
</taxonomicName>
records from Hong Kong. The darker the green areas, the greater the tree canopy cover.
</paragraph>
</caption>
<paragraph id="8BFF2F422623B838FF38D16B01D0A7B1" blockId="13.[189,1399,542,674]" pageId="13" pageNumber="129">
While infrequently collected within
<collectingCountry id="F3576FD22623B838FDF2D16B063BA72B" box="[631,776,542,568]" name="Hong Kong" pageId="13" pageNumber="129">Hong Kong</collectingCountry>
, perhaps due to limited sampling towards arboreal species at this point, the species appears relatively widespread, being found on most larger islands and continental parts of the SAR (
<figureCitation id="137B33C72623B838FDA4D1110759A76C" box="[545,618,612,639]" captionStart="Fig" captionStartId="12.[189,232,1963,1989]" captionTargetBox="[219,1391,1101,1915]" captionTargetId="figure-310@12.[189,1398,1067,1922]" captionTargetPageId="12" captionText="Fig. 5. Distribution map of Temnothorax Mayr, 1861 records from Hong Kong. The darker the green areas, the greater the tree canopy cover." figureDoi="http://doi.org/10.5281/zenodo.8155437" httpUri="https://zenodo.org/record/8155437/files/figure.png" pageId="13" pageNumber="129">Fig. 5</figureCitation>
).
<taxonomicName id="4C4054C12623B838FD07D1110641A76D" authority="Hamer &amp; Lee &amp; Guénard, 2023" authorityName="Hamer &amp; Lee &amp; Guénard" authorityYear="2023" box="[642,882,612,638]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="129" phylum="Arthropoda" rank="species" species="haveni" status="sp. nov.">
<emphasis id="B934F3502623B838FD07D1110641A76D" box="[642,882,612,638]" italics="true" pageId="13" pageNumber="129">Temnothorax haveni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074E2B2623B838FCFCD11006E6A76C" box="[889,981,613,639]" pageId="13" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
is thus expected to be found in the nearby province of Guangdong which shares a similar climate and habitats as
<collectingCountry id="F3576FD22623B838FBD7D1FD01ECA7B1" box="[1106,1247,648,674]" name="Hong Kong" pageId="13" pageNumber="129">Hong Kong</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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