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<document id="A86255D15D65556D45D0D3554C9BD26E" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC2F4C04F7DFFAF9FB11FEA8" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana mzuzuensis Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="D75642D2-177A-4F17-B360-60FCE204FF4E" docUuidSource="ZooBank" docVersion="3" lastPageNumber="33" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="31" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="F89CE7109281E635DBE52DE9F9864642">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="F5B887D569E38013A4095AE84BD7CB0E">Lehmann, Ingo</mods:namePart>
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<mods:namePart id="9E766EC32E54BE2E369C94B9468EF5F6">Zahiri, Reza</mods:namePart>
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<mods:namePart id="A8B59CBA1AC44A55272C5D6C62B3AFD9">Husemann, Martin</mods:namePart>
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<emphasis id="B9010172CC2F4C3AF7DFFAF9FE30FA84" bold="true" box="[151,430,1302,1329]" italics="true" pageId="30" pageNumber="31">Lukeniana mzuzuensis</emphasis>
Lehmann, Zahiri &amp; Husemann
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<figureCitation id="134EC1E5CC2F4C3AF7DFFAD4FE8EFAE0" box="[151,272,1338,1365]" captionStart="FIGURE 5" captionStartId="26.[151,250,1484,1508]" captionTargetBox="[151,1436,181,1460]" captionTargetId="figure-28@26.[151,1436,181,1460]" captionTargetPageId="26" captionText="FIGURE 5. a. Lukeniana chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; b. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; c. L. michaelgrzimeki, paratype, female, Tanzania, Arusha Region, Mount Meru, Momella; d. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave; e. L. mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; f. L. mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkhorongo." figureDoi="http://doi.org/10.5281/zenodo.7840798" httpUri="https://zenodo.org/record/7840798/files/figure.png" pageId="30" pageNumber="31">Figs 5e, 5f</figureCitation>
,
<figureCitation id="134EC1E5CC2F4C3AF654FAD4FED9FAE0" box="[284,327,1339,1365]" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="30" pageNumber="31">12a</figureCitation>
,
<figureCitation id="134EC1E5CC2F4C3AF61BFAD4FE1FFAE0" box="[339,385,1339,1365]" captionStart="FIGURE 11" captionStartId="62.[151,250,1774,1798]" captionTargetBox="[162,1436,606,1750]" captionTargetId="figure-205@62.[151,1436,597,1750]" captionTargetPageId="62" captionText="FIGURE 11. a. Zambezia madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; b. Z. jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); c. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare); d1. Lukeniana mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkho-rongo, female genitalia with ductus bursae and corpus bursae, both thinly membranous and without any structures; d2. Detail of female genitalia of L. mzuzuensis, paratype, with narrow bandson lateroventral surface of segment 8 (cf. red arrows); e. L. obliqualinea, holotype, male, Kenya, County Nairobi, Nairobi (picture taken by Kim Buckmaster, NHMUK, London (picture taken in 1996).." figureDoi="http://doi.org/10.5281/zenodo.7840814" httpUri="https://zenodo.org/record/7840814/files/figure.png" pageId="30" pageNumber="31">11d</figureCitation>
<figureCitation id="134EC1E5CC2F4C3AF6C9FAD4FE12FAE0" box="[385,396,1339,1365]" captionStart="FIGURE 1" captionStartId="9.[152,255,1793,1817]" captionTargetBox="[152,1436,777,1769]" captionTargetId="figure-310@9.[152,1436,777,1769]" captionTargetPageId="9" captionText="FIGURE 1. a. Zambezia jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); b. Z. durrellbarnesi, holotype, female, Zimbabwe, Manicaland Province, Bvumba Mountains; c. Z. darrelplowesi, holotype, female, Zimbabwe, Manicaland Province, Mutare (formerly Umtali), Hospital (today Kopje House); d. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare)." figureDoi="http://doi.org/10.5281/zenodo.7840788" httpUri="https://zenodo.org/record/7840788/files/figure.png" pageId="30" pageNumber="31">1</figureCitation>
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<figureCitation id="134EC1E5CC2F4C3AF6D0FAD4FE5EFAE0" box="[408,448,1339,1365]" captionStart="FIGURE 11" captionStartId="62.[151,250,1774,1798]" captionTargetBox="[162,1436,606,1750]" captionTargetId="figure-205@62.[151,1436,597,1750]" captionTargetPageId="62" captionText="FIGURE 11. a. Zambezia madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; b. Z. jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); c. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare); d1. Lukeniana mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkho-rongo, female genitalia with ductus bursae and corpus bursae, both thinly membranous and without any structures; d2. Detail of female genitalia of L. mzuzuensis, paratype, with narrow bandson lateroventral surface of segment 8 (cf. red arrows); e. L. obliqualinea, holotype, male, Kenya, County Nairobi, Nairobi (picture taken by Kim Buckmaster, NHMUK, London (picture taken in 1996).." figureDoi="http://doi.org/10.5281/zenodo.7840814" httpUri="https://zenodo.org/record/7840814/files/figure.png" pageId="30" pageNumber="31">11d</figureCitation>
<figureCitation id="134EC1E5CC2F4C3AF688FAD4FE50FAE0" box="[448,462,1339,1365]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1880]" captionTargetBox="[151,1436,489,1832]" captionTargetId="figure-187@12.[151,1436,489,1832]" captionTargetPageId="12" captionText="FIGURE 2. a. Lukeniana butleri, paratype, male, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; b. L. butleri, paratype, female, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; c. L. kollhorsti, holotype, male, Zimbabwe, Manicaland Province, Busi Farm; d. Zambezia diredaouaensis, holotype, male, Ethiopia, Harar Plateau, Dire Daua, today: chartered city of Dire Dawa; e. Z. madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; f. Z. madambae, paratype, female, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids." figureDoi="http://doi.org/10.5281/zenodo.7840790" httpUri="https://zenodo.org/record/7840790/files/figure.png" pageId="30" pageNumber="31">2</figureCitation>
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.
<materialsCitation id="3B1DD73DCC2F4C3AF3C7F9D8FD4EF92C" collectingDate="2009-10-17" collectionCode="R, ZFMK" collectorName="J. Murphy &amp; I. Lehmann" country="Malawi" county="Northern Region" elevation="1" location="Northern Region" municipality="Mzuzu" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Mzimba" typeStatus="paratype">
<typeStatus id="54CE63C2CC2F4C3AF3C7F9D8FA9EF9E4" box="[1167,1280,1591,1617]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7316E9CC2F4C3AF244F9D8FA06F9E4" box="[1292,1432,1591,1617]" pageId="30" pageNumber="31" type="female">two females</specimenCount>
,
<collectingCountry id="F3629DF0CC2F4C3AF7DFF9B4FF71F9C0" box="[151,239,1627,1653]" name="Malawi" pageId="30" pageNumber="31">Malawi</collectingCountry>
,
<location id="8EAA8BBBCC2F4C3AF7B3F9B4FE25F9C0" LSID="urn:lsid:plazi:treatment:03DC6C76CC2F4C04F7DFFAF9FB11FEA8:8EAA8BBBCC2F4C3AF7B3F9B4FE25F9C0" box="[251,443,1627,1653]" country="Malawi" name="Northern Region" pageId="30" pageNumber="31" stateProvince="Mzimba">Northern Region</location>
,
<collectingRegion id="49B11382CC2F4C3AF68EF9B4FD1FF9C0" box="[454,641,1627,1653]" country="Malawi" name="Mzimba" pageId="30" pageNumber="31">Mzimba District</collectingRegion>
, same locality and elevation,
<date id="FFCBFBA0CC2F4C3AF485F9B4FB12F9C0" box="[973,1164,1627,1653]" pageId="30" pageNumber="31" value="2009-10-17">
<collectingDate id="EF8F0248CC2F4C3AF485F9B4FB12F9C0" box="[973,1164,1627,1653]" pageId="30" pageNumber="31" value="2009-10-17">17 October 2009</collectingDate>
</date>
,
<collectionCode id="ED6445A5CC2F4C3AF3DFF9B4FB30F9C0" box="[1175,1198,1627,1653]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="31">R</collectionCode>
.
<collectorName id="2680B8B6CC2F4C3AF3FAF9B4FABAF9C0" box="[1202,1316,1627,1653]" pageId="30" pageNumber="31">J. Murphy</collectorName>
leg., genitalia slide numbers 05/052012
<collectorName id="2680B8B6CC2F4C3AF6A5F990FDF9F92C" box="[493,615,1663,1689]" pageId="30" pageNumber="31">I.Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2F4C3AF53CF990FD56F92C" box="[628,712,1663,1689]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B1DD73DCC2F4C3AF592F991FBFDF92C" box="[730,1123,1662,1689]" collectionCode="ZFMK" country="Malawi" location="Stuning" municipality="Dr." pageId="30" pageNumber="31" specimenCount="1" stateProvince="Mzimba" typeStatus="paratype">
55/052013
<collectingMunicipality id="6BAE471ACC2F4C3AF410F990FCE2F92C" box="[856,892,1663,1689]" pageId="30" pageNumber="31">Dr.</collectingMunicipality>
D.
<location id="8EAA8BBBCC2F4C3AF4EAF990FC67F92C" LSID="urn:lsid:plazi:treatment:03DC6C76CC2F4C04F7DFFAF9FB11FEA8:8EAA8BBBCC2F4C3AF4EAF990FC67F92C" box="[930,1017,1663,1689]" country="Malawi" municipality="Dr." name="Stuning" pageId="30" pageNumber="31" stateProvince="Mzimba">Stüning</location>
(
<collectionCode id="ED6445A5CC2F4C3AF34FF990FBC5F92C" box="[1031,1115,1663,1689]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF326F990FD85F954" collectingDate="2008-10-25" collectingDateMax="2008-10-27" collectingDateMin="2008-10-25" collectorName="I. Lehmann" country="Malawi" location="Nkho-rongo" pageId="30" pageNumber="31" specimenCount="14" specimenCount-male="14" stateProvince="Mzimba" typeStatus="paratype">
<specimenCount id="9D7316E9CC2F4C3AF326F990FB4FF92C" box="[1134,1233,1663,1689]" pageId="30" pageNumber="31" type="male">14 males</specimenCount>
from
<location id="8EAA8BBBCC2F4C3AF25BF990FA02F92C" LSID="urn:lsid:plazi:treatment:03DC6C76CC2F4C04F7DFFAF9FB11FEA8:8EAA8BBBCC2F4C3AF25BF990FA02F92C" box="[1299,1436,1663,1689]" country="Malawi" name="Nkho-rongo" pageId="30" pageNumber="31" stateProvince="Mzimba">Nkho-rongo</location>
(same elevation and collector): male,
<date id="FFCBFBA0CC2F4C3AF505F94DFC8DF908" box="[589,787,1698,1725]" pageId="30" pageNumber="31" value="2008-10-25">
<collectingDate id="EF8F0248CC2F4C3AF505F94DFC8DF908" box="[589,787,1698,1725]" pageId="30" pageNumber="31" value="2008-10-25">25 October 2008</collectingDate>
</date>
, genitalia slide number 28/102015
<collectorName id="2680B8B6CC2F4C3AF3E5F94CFAB4F908" box="[1197,1322,1699,1725]" pageId="30" pageNumber="31">I.Lehmann</collectorName>
, male,
<date id="FFCBFBA0CC2F4C3AF2C8F94CFEB1F954" pageId="30" pageNumber="31" value="2008-10-27">
<collectingDate id="EF8F0248CC2F4C3AF2C8F94CFEB1F954" pageId="30" pageNumber="31" value="2008-10-27">27 October 2008</collectingDate>
</date>
(authors collection)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF56DF928FC35F954" box="[549,939,1735,1761]" collectingDate="2009-10-12" collectionCode="ZMA" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Mzimba" typeStatus="paratype">
<specimenCount id="9D7316E9CC2F4C3AF56DF928FD12F954" box="[549,652,1735,1761]" pageId="30" pageNumber="31" type="male">one male</specimenCount>
,
<date id="FFCBFBA0CC2F4C3AF5DEF928FCCEF954" box="[662,848,1735,1761]" pageId="30" pageNumber="31" value="2009-10-12">
<collectingDate id="EF8F0248CC2F4C3AF5DEF928FCCEF954" box="[662,848,1735,1761]" pageId="30" pageNumber="31" value="2009-10-12">12 October 2009</collectingDate>
</date>
(
<collectionCode id="ED6445A5CC2F4C3AF416F928FC3CF954" box="[862,930,1735,1761]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="30" pageNumber="31">ZMA</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D7316E9CC2F4C3AF4FDF928FBA9F954" box="[949,1079,1735,1761]" pageId="30" pageNumber="31" type="male">three males</specimenCount>
,
<date id="FFCBFBA0CC2F4C3AF309F928FB60F954" box="[1089,1278,1735,1761]" pageId="30" pageNumber="31" value="2009-10-29">29 October 2009</date>
,
<materialsCitation id="3B1DD73DCC2F4C3AF240F928FCC4F8B0" collectionCode="ZMFK" country="Malawi" location="Lehmann" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Mzimba" typeStatus="paratype">
<specimenCount id="9D7316E9CC2F4C3AF240F928FAF2F954" box="[1288,1388,1735,1761]" pageId="30" pageNumber="31" type="male">one male</specimenCount>
dissected, genitalia slide number 05/092014 I.
<location id="8EAA8BBBCC2F4C3AF5CCF904FD70F8B0" LSID="urn:lsid:plazi:treatment:03DC6C76CC2F4C04F7DFFAF9FB11FEA8:8EAA8BBBCC2F4C3AF5CCF904FD70F8B0" box="[644,750,1771,1797]" country="Malawi" name="Lehmann" pageId="30" pageNumber="31" stateProvince="Mzimba">Lehmann</location>
(
<collectionCode id="ED6445A5CC2F4C3AF5B5F904FCCFF8B0" box="[765,849,1771,1797]" pageId="30" pageNumber="31">ZMFK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF42DF904FB61F8B0" box="[869,1279,1771,1797]" collectingDate="2009-10-31" collectionCode="ZMA" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Mzimba" typeStatus="paratype">
<specimenCount id="9D7316E9CC2F4C3AF42DF904FC45F8B0" box="[869,987,1771,1797]" pageId="30" pageNumber="31" type="male">two males</specimenCount>
,
<date id="FFCBFBA0CC2F4C3AF4AEF904FB3AF8B0" box="[998,1188,1771,1797]" pageId="30" pageNumber="31" value="2009-10-31">
<collectingDate id="EF8F0248CC2F4C3AF4AEF904FB3AF8B0" box="[998,1188,1771,1797]" pageId="30" pageNumber="31" value="2009-10-31">31 October 2009</collectingDate>
</date>
(
<collectionCode id="ED6445A5CC2F4C3AF3FBF904FB69F8B0" box="[1203,1271,1771,1797]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="30" pageNumber="31">ZMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF243F904FE38F89C" collectingDate="2011-11-07" collectionCode="NRM" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Mzimba" typeStatus="paratype">
<specimenCount id="9D7316E9CC2F4C3AF243F904FAEAF8B0" box="[1291,1396,1771,1797]" pageId="30" pageNumber="31" type="male">one male</specimenCount>
,
<date id="FFCBFBA0CC2F4C3AF2C8F904FED4F89C" pageId="30" pageNumber="31" value="2011-11-07">
<collectingDate id="EF8F0248CC2F4C3AF2C8F904FED4F89C" pageId="30" pageNumber="31" value="2011-11-07">07 November 2011</collectingDate>
</date>
(
<collectionCode id="ED6445A5CC2F4C3AF610F8E0FE00F89C" box="[344,414,1807,1833]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="30" pageNumber="31" type="Museum">NRM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF6F8F8E0FACBF89C" box="[432,1365,1807,1833]" collectingDate="2011-11-08" collectionCode="NHMUK" collectorName="Lehmann" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Mzimba" typeStatus="paratype">
male,
<date id="FFCBFBA0CC2F4C3AF6BBF8E0FD55F89C" box="[499,715,1807,1833]" pageId="30" pageNumber="31" value="2011-11-08">
<collectingDate id="EF8F0248CC2F4C3AF6BBF8E0FD55F89C" box="[499,715,1807,1833]" pageId="30" pageNumber="31" value="2011-11-08">08 November 2011</collectingDate>
</date>
, genitalia slide number 14/092014 I.
<collectorName id="2680B8B6CC2F4C3AF32DF8E0FB51F89C" box="[1125,1231,1807,1833]" pageId="30" pageNumber="31">Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2F4C3AF394F8E0FAD3F89C" box="[1244,1357,1807,1833]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF217F8E0FC52F8F8" collectingDate="2012-10-31" collectionCode="ZMUC" collectorName="Lehmann" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Mzimba" typeStatus="paratype">
male,
<date id="FFCBFBA0CC2F4C3AF7DFF8DCFEC8F8F8" box="[151,342,1843,1869]" pageId="30" pageNumber="31" value="2012-10-31">
<collectingDate id="EF8F0248CC2F4C3AF7DFF8DCFEC8F8F8" box="[151,342,1843,1869]" pageId="30" pageNumber="31" value="2012-10-31">31 October 2012</collectingDate>
</date>
, genitalia slide number 22/072013 I.
<collectorName id="2680B8B6CC2F4C3AF5BDF8DCFCC1F8F8" box="[757,863,1843,1869]" pageId="30" pageNumber="31">Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2F4C3AF425F8DCFC5AF8F8" box="[877,964,1843,1869]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="30" pageNumber="31">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF49FF8DCFDC1F8C4" collectingDate="2013-11-03" collectionCode="SMNS" collectorName="Lehmann" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Mzimba" typeStatus="paratype">
male,
<date id="FFCBFBA0CC2F4C3AF353F8DCFB68F8F8" box="[1051,1270,1843,1869]" pageId="30" pageNumber="31" value="2013-11-03">
<collectingDate id="EF8F0248CC2F4C3AF353F8DCFB68F8F8" box="[1051,1270,1843,1869]" pageId="30" pageNumber="31" value="2013-11-03">03 November 2013</collectingDate>
</date>
, genitalia slide number 21/092014 I.
<collectorName id="2680B8B6CC2F4C3AF6C4F8B8FE68F8C4" box="[396,502,1879,1905]" pageId="30" pageNumber="31">Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2F4C3AF54DF8B8FDC9F8C4" box="[517,599,1879,1905]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="30" pageNumber="31">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC2F4C3AF524F8B8FF75F820" collectingDate="2013-11-08" collectionCode="ZMH" collectorName="Lehmann" country="Malawi" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Mzimba" typeStatus="paratype">
male,
<date id="FFCBFBA0CC2F4C3AF5F9F8B8FC13F8C4" box="[689,909,1879,1905]" pageId="30" pageNumber="31" value="2013-11-08">
<collectingDate id="EF8F0248CC2F4C3AF5F9F8B8FC13F8C4" box="[689,909,1879,1905]" pageId="30" pageNumber="31" value="2013-11-08">08 November 2013</collectingDate>
</date>
, genitalia slide number 15/092014 I.
<collectorName id="2680B8B6CC2F4C3AF27AF8B8FA02F8C4" box="[1330,1436,1879,1905]" pageId="30" pageNumber="31">Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2F4C3AF7D7F894FF7DF820" box="[159,227,1915,1941]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="30" pageNumber="31" type="Museum">ZMH</collectionCode>
)
</materialsCitation>
; male,
<date id="FFCBFBA0CC2F4C3AF673F894FE69F820" box="[315,503,1915,1941]" pageId="30" pageNumber="31" value="1997-10-29">29 October 1997</date>
genitalia slide number 18/092012 I. Lehmann (authors collection).
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2F4C3BF78FF871FE57F9BC" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="8BCADD60CC2F4C3BF78FF871FEB5FF4C" blockId="30.[151,1437,1554,2013]" lastBlockId="31.[151,1437,150,2013]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="B9010172CC2F4C3AF78FF871FEC7F80D" bold="true" box="[199,345,1950,1976]" pageId="30" pageNumber="31">Description.</emphasis>
Male.
<emphasis id="B9010172CC2F4C3AF6E6F870FE75F80C" box="[430,491,1951,1977]" italics="true" pageId="30" pageNumber="31">Head</emphasis>
: Cream-coloured and mixed with some sepia, shiny, hair-like scales; long, dense hair-like scales between eyes; eyes brown with large black patches; antenna 0.50 length of forewing, bipectinate, with branches 4.5 width of shaft, branches covered with cream-coloured scales laterally, shaft covered with creamcoloured scales dorsally; antennal tips slightly spatulate with two long scales, bending towards apex; labial palpi cream-ochre.
</paragraph>
<paragraph id="8BCADD60CC2E4C3BF78FFEECFCC7FDD4" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FFEECFE86FEA8" box="[199,280,259,285]" italics="true" pageId="31" pageNumber="32">Thorax</emphasis>
: Patagia and tegulae with shiny and long hair-like scales of cream-ochre mixed with sepia; a small crest of cream-coloured scales mixed with ivory-yellow on metathorax. Hindlegs yellow ochre with fine hair-like scales, shiny; one pair of narrow, long tibial spurs present, outer spur
<emphasis id="B9010172CC2E4C3BF41BFEA4FCEAFED1" box="[851,884,331,356]" italics="true" pageId="31" pageNumber="32">ca.</emphasis>
<quantity id="4C8D7085CC2E4C3BF433FEA4FC4DFED0" box="[891,979,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="31" pageNumber="32" unit="mm" value="1.1">1.1 mm</quantity>
, inner spur
<emphasis id="B9010172CC2E4C3BF31FFEA4FBE7FED1" box="[1111,1145,331,356]" italics="true" pageId="31" pageNumber="32">ca.</emphasis>
<quantity id="4C8D7085CC2E4C3BF3C8FEA4FB47FED0" box="[1152,1241,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="mm" value="0.9">0.9 mm</quantity>
. Forewing length of
<typeStatus id="54CE63C2CC2E4C3BF7F0FE80FE84FE3C" box="[184,282,367,393]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
13.0 mm (wingspan 29.0 mm). Forewing upperside ochre and contrasting; costal margin and entire forewing with sepia coloured striae; termen with small lunules of sepia; terminal line and subterminal line of sepia extending from near apex to dorsum; CuA
<subScript id="17F1DF25CC2E4C3BF534FE26FD1AFE62" attach="left" box="[636,644,457,471]" fontSize="6" pageId="31" pageNumber="32">2</subScript>
very broad and ivory-yellow, edged sepia above; all remaining veins ochre and hence not distinctly coloured except 1A+2A sepia (less visible in females); cilia long,
<quantity id="4C8D7085CC2E4C3BF3A0FE34FADFFE40" box="[1256,1345,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="32" unit="mm" value="1.2">1.2 mm</quantity>
, creamcoloured, shiny. Underside of forewing roughly scaled, ivory-yellow, glossy; costal margin with few striae of dark ochre; terminal and subterminal lines concolorous with striae. Hindwing upperside and cilia ivory-yellow, glossy, cilia long,
<quantity id="4C8D7085CC2E4C3BF646FDA8FEF9FDD4" box="[270,359,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="32" unit="mm" value="1.2">1.2 mm</quantity>
; underside as in forewing but without striae.
</paragraph>
<paragraph id="8BCADD60CC2E4C3BF78FFD84FEF7FC58" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FFD84FEAFFD30" box="[199,305,619,645]" italics="true" pageId="31" pageNumber="32">Abdomen</emphasis>
: Mainly olive and cream-coloured, shiny; abdominal tuft short, one-fourth abdominal length. Genitalia (
<figureCitation id="134EC1E5CC2E4C3BF78AFD60FE81FD1C" box="[194,287,655,681]" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="31" pageNumber="32">Fig. 12a</figureCitation>
) with uncus lobes bearing rounded tips, both short and long setae ventrally, basal edge of uncus bent at middle (ventral view); gnathos arms long, 1.3 longer than basal width of valva; valvae ovoid, broad at base, costa with few setae; sacculus with a few long setae; weakly-sclerotized projection densely setose, with rectangular tip, and slightly longer than single thorn-like process below; the latter hollow and with few scattered setae; median sector of valva with some long setae on inner side; an elongate emargination extending between weakly-sclerotized projection and thorn-like process, 4050% of length of valva; ventral side of valva slightly bent at middle. Saccus short, stoutly finger-shaped, opposite saccus a broader vinculum forming a small chitinized rectangular plate. Juxta as long as saccus with two acuminate tips, each with a short process, between tips a deep emargination 90% of length of juxta. Phallus long, 1.2 as long as width of valva, not trumpet-like, bent at middle, weakly bilobate with a cleft at each end.
</paragraph>
<paragraph id="8BCADD60CC2E4C3BF78FFC18FD71FBEC" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FFC18FEBBFBA4" bold="true" box="[199,293,1015,1041]" pageId="31" pageNumber="32">Female.</emphasis>
<emphasis id="B9010172CC2E4C3BF666FC18FEF5FBA4" box="[302,363,1015,1041]" italics="true" pageId="31" pageNumber="32">Head</emphasis>
: Essentially as described for male, except antenna 0.31 length of forewing, unipectinate, with branches 1.5 width of shaft, shaft scaled dorsally, branches not scaled; antennal tips very short, not bending towards apex; forewing length
<quantity id="4C8D7085CC2E4C3BF6DCFBD1FE66FBEC" box="[404,504,1086,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="31" pageNumber="32" unit="mm" value="16.5">16.5 mm</quantity>
(wingspan 36.0 mm).
</paragraph>
<paragraph id="8BCADD60CC2E4C3BF78FFB8CFB3EFB70" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FFB8CFE86FBC8" box="[199,280,1123,1149]" italics="true" pageId="31" pageNumber="32">Thorax</emphasis>
: Essentially as described for male, except, forewing upperside lighter and less contrasting than in male. Both terminal and subterminal lines more faded than in males. Subterminal line not slightly bent towards base of wing at outer half of M
<subScript id="17F1DF25CC2E4C3BF6D4FB52FE37FB7E" attach="left" box="[412,425,1213,1227]" fontSize="6" pageId="31" pageNumber="32">2.</subScript>
Distinctness of sepia coloration of 1A+2A less visible than in male.
</paragraph>
<paragraph id="8BCADD60CC2E4C3BF78FFB20FE57F9BC" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FFB20FEAFFB5C" box="[199,305,1231,1257]" italics="true" pageId="31" pageNumber="32">Abdomen</emphasis>
. Genitalia (
<figureCitation id="134EC1E5CC2E4C3BF6F0FB20FDBEFB5C" box="[440,544,1231,1257]" captionStart="FIGURE 11" captionStartId="62.[151,250,1774,1798]" captionTargetBox="[162,1436,606,1750]" captionTargetId="figure-205@62.[151,1436,597,1750]" captionTargetPageId="62" captionText="FIGURE 11. a. Zambezia madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; b. Z. jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); c. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare); d1. Lukeniana mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkho-rongo, female genitalia with ductus bursae and corpus bursae, both thinly membranous and without any structures; d2. Detail of female genitalia of L. mzuzuensis, paratype, with narrow bandson lateroventral surface of segment 8 (cf. red arrows); e. L. obliqualinea, holotype, male, Kenya, County Nairobi, Nairobi (picture taken by Kim Buckmaster, NHMUK, London (picture taken in 1996).." figureDoi="http://doi.org/10.5281/zenodo.7840814" httpUri="https://zenodo.org/record/7840814/files/figure.png" pageId="31" pageNumber="32">Figs 11d</figureCitation>
<figureCitation id="134EC1E5CC2E4C3BF569FB20FDB2FB5C" box="[545,556,1231,1257]" captionStart="FIGURE 1" captionStartId="9.[152,255,1793,1817]" captionTargetBox="[152,1436,777,1769]" captionTargetId="figure-310@9.[152,1436,777,1769]" captionTargetPageId="9" captionText="FIGURE 1. a. Zambezia jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); b. Z. durrellbarnesi, holotype, female, Zimbabwe, Manicaland Province, Bvumba Mountains; c. Z. darrelplowesi, holotype, female, Zimbabwe, Manicaland Province, Mutare (formerly Umtali), Hospital (today Kopje House); d. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare)." figureDoi="http://doi.org/10.5281/zenodo.7840788" httpUri="https://zenodo.org/record/7840788/files/figure.png" pageId="31" pageNumber="32">1</figureCitation>
,
<figureCitation id="134EC1E5CC2E4C3BF571FB20FDFCFB5C" box="[569,610,1231,1257]" captionStart="FIGURE 11" captionStartId="62.[151,250,1774,1798]" captionTargetBox="[162,1436,606,1750]" captionTargetId="figure-205@62.[151,1436,597,1750]" captionTargetPageId="62" captionText="FIGURE 11. a. Zambezia madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; b. Z. jennyhuntae, holotype, male, Zimbabwe, Manicaland Province, Mutare (formerly Umtali); c. Metarbelodes umtaliana, holotype, male, Zimbabwe, Manicaland Province, Old Umtali or Umtali (today Mutare); d1. Lukeniana mzuzuensis, paratype, female, Malawi, Northern Region, Mzuzu, Nkho-rongo, female genitalia with ductus bursae and corpus bursae, both thinly membranous and without any structures; d2. Detail of female genitalia of L. mzuzuensis, paratype, with narrow bandson lateroventral surface of segment 8 (cf. red arrows); e. L. obliqualinea, holotype, male, Kenya, County Nairobi, Nairobi (picture taken by Kim Buckmaster, NHMUK, London (picture taken in 1996).." figureDoi="http://doi.org/10.5281/zenodo.7840814" httpUri="https://zenodo.org/record/7840814/files/figure.png" pageId="31" pageNumber="32">11d</figureCitation>
<figureCitation id="134EC1E5CC2E4C3BF52AFB20FDEFFB5C" box="[610,625,1231,1257]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1880]" captionTargetBox="[151,1436,489,1832]" captionTargetId="figure-187@12.[151,1436,489,1832]" captionTargetPageId="12" captionText="FIGURE 2. a. Lukeniana butleri, paratype, male, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; b. L. butleri, paratype, female, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; c. L. kollhorsti, holotype, male, Zimbabwe, Manicaland Province, Busi Farm; d. Zambezia diredaouaensis, holotype, male, Ethiopia, Harar Plateau, Dire Daua, today: chartered city of Dire Dawa; e. Z. madambae, holotype, male, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids; f. Z. madambae, paratype, female, Botswana, Chobe District Council, Kasane, Chobe River, Chobe Rapids." figureDoi="http://doi.org/10.5281/zenodo.7840790" httpUri="https://zenodo.org/record/7840790/files/figure.png" pageId="31" pageNumber="32">2</figureCitation>
) with papillae anales broad, 8-shaped in posterior-dorsal view, densely covered with long and short setae. Segment 8 with few scattered setae along posterior margin and two narrow lateroventral sclerotized plates or bands; inner plate with an acuminate end, outer plate with a rectangular end; broader plate covering
<emphasis id="B9010172CC2E4C3BF609FAD4FEFDFAE1" box="[321,355,1339,1364]" italics="true" pageId="31" pageNumber="32">ca.</emphasis>
35% of width of ventral side of segment 8. Dorso-anterior margin of abdominal plate without an emargination. Posterior apophyses straight, almost twice as broad at base as width of anterior apophyses, the broader part extending to half of length of anterior apophyses, posteriorly with a narrow thorn-like projection and a uneven dorsal edge. Anterior apophyses slightly sinuate, slightly longer than posterior apophyses. Ductus and corpus bursae broad, thinly membranous, without any distinct characters; corpus bursae rounded in shape, 1.2 as long as anterior apophyses.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2E4C3BF78FF9FDFD34F820" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BCADD60CC2E4C3BF78FF9FDFD34F820" blockId="31.[151,1437,150,2013]" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FF9FDFEDCF999" bold="true" box="[199,322,1554,1580]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
The closest related species in the genus
<taxonomicName id="4C75A6E3CC2E4C3BF443F9FCFC1FF998" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[779,897,1555,1581]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B9010172CC2E4C3BF443F9FCFC1FF998" box="[779,897,1555,1581]" italics="true" pageId="31" pageNumber="32">Lukeniana</emphasis>
</taxonomicName>
is unknown. On the basis of the contrasting colouration of the forewing, with its broad, white CuA
<subScript id="17F1DF25CC2E4C3BF5A2F9A6FD6CF9E2" attach="left" box="[746,754,1609,1623]" fontSize="6" pageId="31" pageNumber="32">2</subScript>
,
<taxonomicName id="4C75A6E3CC2E4C3BF44AF9D8FC00F9E5" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[770,926,1591,1616]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2E4C3BF44AF9D8FC00F9E5" box="[770,926,1591,1616]" italics="true" pageId="31" pageNumber="32">L. mzuzuensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C75A6E3CC2E4C3BF357F9D8FA83F9E4" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1055,1309,1591,1617]" class="Insecta" family="Metarbelidae" genus="Zambezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jennyhuntae">
<emphasis id="B9010172CC2E4C3BF357F9D8FA83F9E4" box="[1055,1309,1591,1617]" italics="true" pageId="31" pageNumber="32">Zambezia jennyhuntae</emphasis>
</taxonomicName>
, indicating that the wing colouration cannot be used alone in the field do distinguish species of both genera, particularly where species of both genera occur sympatric in the Zambezian regional centre of endemism. For example,
<taxonomicName id="4C75A6E3CC2E4C3BF249F990FA02F92D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1281,1436,1663,1688]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2E4C3BF249F990FA02F92D" box="[1281,1436,1663,1688]" italics="true" pageId="31" pageNumber="32">L. mzuzuensis</emphasis>
</taxonomicName>
can be easily distinguished by the CuP vein of the forewing present only as a weak, discontinuous fold, rather than the strong, continuous, tubular fold of
<taxonomicName id="4C75A6E3CC2E4C3BF501F928FD6EF954" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[585,752,1735,1761]" class="Insecta" family="Metarbelidae" genus="Zambezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jennyhuntae">
<emphasis id="B9010172CC2E4C3BF501F928FD6EF954" box="[585,752,1735,1761]" italics="true" pageId="31" pageNumber="32">Z. jennyhuntae</emphasis>
</taxonomicName>
. The male genitalia of
<taxonomicName id="4C75A6E3CC2E4C3BF4BEF928FB0CF955" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1014,1170,1735,1760]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2E4C3BF4BEF928FB0CF955" box="[1014,1170,1735,1760]" italics="true" pageId="31" pageNumber="32">L. mzuzuensis</emphasis>
</taxonomicName>
have at least four diagnostic characters: (i) the ventral side of the valva is slightly bent at middle; (ii) the emargination of the valva is long, 4050% of the length of valva; (iii) the thorn-like process is well developed; and (iv) the sacculus is broad, at least one-third the width of the valva. The female post-abdominal structure of
<taxonomicName id="4C75A6E3CC2E4C3BF486F8DCFBF4F8F9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[974,1130,1843,1868]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2E4C3BF486F8DCFBF4F8F9" box="[974,1130,1843,1868]" italics="true" pageId="31" pageNumber="32">L. mzuzuensis</emphasis>
</taxonomicName>
has two diagnostic characters: (i) a latero-ventral plate covers only 35% of the ventral side of segment 8; and (ii) the posterior apophyses have a short thorn-like process at their posterior end.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2E4C04F78FF871FE77FF4C" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BCADD60CC2E4C04F78FF871FE7CFF60" blockId="31.[151,1437,150,2013]" lastBlockId="32.[151,1436,151,285]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B9010172CC2E4C3BF78FF871FEFFF80D" bold="true" box="[199,353,1950,1976]" pageId="31" pageNumber="32">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC2E4C3BF622F870FDF9F80D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[362,615,1951,1977]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC2E4C3BF622F870FDF9F80D" box="[362,615,1951,1977]" italics="true" pageId="31" pageNumber="32">Lukeniana mzuzuensis</emphasis>
</taxonomicName>
is known from Nkhorongo in northern
<collectingCountry id="F3629DF0CC2E4C3BF378F870FB17F80C" box="[1072,1161,1951,1977]" name="Malawi" pageId="31" pageNumber="32">Malawi</collectingCountry>
. Nkhorongo is a suburb of Mzuzu (elevation
<quantity id="4C8D7085CC2E4C3BF6C0F82DFDAAF868" box="[392,564,1986,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.289" metricValueMax="1.323" metricValueMin="1.255" pageId="31" pageNumber="32" unit="m" value="1289.0" valueMax="1323.0" valueMin="1255.0">
<geoCoordinate id="EE41BBA7CC2E4C3BF6C0F82DFE59F868" box="[392,455,1986,2013]" degrees="1,255" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="55" value="1.255">1,255</geoCoordinate>
<geoCoordinate id="EE41BBA7CC2E4C3BF68EF82DFD8DF868" box="[454,531,1986,2013]" degrees="1,323" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="55" value="-1.323">1,323</geoCoordinate>
m
</quantity>
) located
<emphasis id="B9010172CC2E4C3BF5D7F82CFD5FF869" box="[671,705,1987,2012]" italics="true" pageId="31" pageNumber="32">ca.</emphasis>
<quantity id="4C8D7085CC2E4C3BF582F82CFC8DF868" box="[714,787,1987,2013]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="31" pageNumber="32" unit="km" value="38.0">38 km</quantity>
west of the
<collectingCountry id="F3629DF0CC2E4C3BF4ECF82CFC64F868" box="[932,1018,1987,2013]" name="Malawi" pageId="31" pageNumber="32">Malawi</collectingCountry>
Rift as part of the EARS and in the Zambezian regional centre of endemism (
<emphasis id="B9010172CC114C04F530FF78FD28FF05" box="[632,694,151,176]" italics="true" pageId="32" pageNumber="33">sensu</emphasis>
<bibRefCitation id="EFE4A091CC114C04F588FF78FCD9FF04" author="White, F." box="[704,839,151,177]" pageId="32" pageNumber="33" refId="ref83858" refString="White, F. (1983) The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco / AETFAT / UNSO Vegetation Map of Africa. Natural Resources Research No. XX. Unesco, Paris, 356 pp." type="book" year="1983">White 1983</bibRefCitation>
) (
<figureCitation id="134EC1E5CC114C04F429FF78FC5BFF04" box="[865,965,151,177]" captionStart="FIGURE 20" captionStartId="72.[151,250,1115,1139]" captionTargetBox="[151,1436,181,1091]" captionTargetId="figure-28@72.[151,1436,181,1091]" captionTargetPageId="72" captionText="FIGURE 20. a. Habitat of Lukeniana enaiposha with Rothschilds Giraffes, Giraffa camelopardalis rothschildi Lydekker (1903) graze in a woodland site at 1.786 m elevation within Lake Nakuru National Park (Kenya) ca. 1 km south of the lake dominated by tall (&gt; 15 m) and young trees of Acacia xanthophloea Benth. (Fabaceae: Mimosoideae) mixed with shrubs of Rhus natalensis Bernh. (Anacardiaceae) (right) and Solanum incanum L. (Solanaceae) (foreground). Picture taken by Shimoni Lehmann in March 2016. b. Between 1600 and 2000 m Lower montane Cassipourea forest sensu Hemp (2006b) covers the slopes of Mount Meru (4.566 m). The picture shows an undisturbed forest of this type comprising trees of Ficus thonningii Blume (Moraceae), Olea capensis L. (Oleaceae) and Ekebergia capensis Sparrm. (Meliaceae) with smaller trees of Vepris simplicifolia (Verd.) Mziray (Rutaceae) and Casearia battiscombei R.E. Fries (Flacourtiaceae). Ferns are present on the ground indicating a wetter site with species of Pteris dentata Forssk. (Pteridaceae) and Asplenium aethiopicum (Burm. f.) Bech. (Aspleniaceae). Note that Lukeniana timdavenporti occurs in a similar habitat on Mount Elgon (Kenya/Uganda), but is not closely related to L. andreashempi and L. michaelgrzimeki, that both occur on Mount Meru. Picture by Dr. Andreas Hemp. c. Habitat of Lukeniana mzuzuensis: wetter, ca. 50 years old Zambezian miombo woodland sensu White (1983) at Nkhorongo (Malawi) with diverse structure, high species diversity, e.g. leguminous trees. The treelet with broad leaves in the bottom right corner is Uapaca nitida Müll. Arg. (Euphorbiaceae); in the centre left is U. kirkiana Müll. Arg.; background trees are Brachystegia utilis Burtt Davy &amp; Hutch., B. floribunda Benth., B. spiciformis Benth. (Fabaceae: Caesalpiniaceae), Uapaca sansibarica Pax., Julbernardia paniculata (Benth.) Troupin (Caesalpiniaceae), Dalbergia melanoxylon Guill. &amp; Perr. (Papilionaceae). In the top right centre is Cussonia arborea Hochst. (Araliaceae) with light green leaves. Picture by Raymond Murphy (Nkhorongo), May 2016. d. Road ca. 23 km southeast Mbala (Zambia), close to the habitat of Lukeniana hausmanni, on the Southern African Plateau with northern wet miombo sensu Chidumayo (1997); Brachystegia-Isoberlinia-Julbernardia association mixed with Albizia antunesiana Harms (Fabaceae: Mimosoideae). Picture by Jürgen Lenz (Leipzig), October 2009." figureDoi="http://doi.org/10.5281/zenodo.7840837" httpUri="https://zenodo.org/record/7840837/files/figure.png" pageId="32" pageNumber="33">Fig. 20c</figureCitation>
).
<taxonomicName id="4C75A6E3CC114C04F493FF78FB47FF05" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[987,1241,151,177]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC114C04F493FF78FB47FF05" box="[987,1241,151,177]" italics="true" pageId="32" pageNumber="33">Lukeniana mzuzuensis</emphasis>
</taxonomicName>
is considered an Afromontane linking species.
</paragraph>
<paragraph id="8BCADD60CC114C04F78FFF31FE77FF4C" blockId="32.[151,1436,151,285]" box="[199,489,222,249]" pageId="32" pageNumber="33">
<emphasis id="B9010172CC114C04F78FFF31FEB4FF4D" bold="true" box="[199,298,222,248]" pageId="32" pageNumber="33">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC114C04F78FFEECFB11FEA8" box="[199,1167,259,285]" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8BCADD60CC114C04F78FFEECFB11FEA8" blockId="32.[151,1436,151,285]" box="[199,1167,259,285]" pageId="32" pageNumber="33">
<emphasis id="B9010172CC114C04F78FFEECFED0FEA8" bold="true" box="[199,334,259,285]" pageId="32" pageNumber="33">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC114C04F61DFEECFDCEFEA9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[341,592,259,285]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mzuzuensis">
<emphasis id="B9010172CC114C04F61DFEECFDCEFEA9" box="[341,592,259,285]" italics="true" pageId="32" pageNumber="33">Lukeniana mzuzuensis</emphasis>
</taxonomicName>
is named after the
<typeStatus id="54CE63C2CC114C04F46EFEECFCC8FEA8" box="[806,854,259,285]" pageId="32" pageNumber="33">type</typeStatus>
locality of Mzuzu,
<collectingCountry id="F3629DF0CC114C04F37AFEECFB15FEA8" box="[1074,1163,259,285]" name="Malawi" pageId="32" pageNumber="33">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>