507 lines
91 KiB
XML
507 lines
91 KiB
XML
<document id="4B6E3741FFB014163FA12870ED29D1D8" ID="10.11646/zootaxa.4258.1.1SLASH10897" ID-CLB-Dataset="33167" ID-DOI="10.11646/zootaxa.4258.1.1" ID-GBIF-Dataset="fdbe32e3-f5b6-4537-b870-6d295865da73" ID-ISSN="1175-5326" ID-Zenodo-Dep="569114" ID-ZooBank="B1F45C56-A43A-4A8C-9190-2B861A37D33C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493194184337" checkinUser="plazi" docAuthor="Judson, Mark L. I." docDate="2017" docId="03D65C0B657E3358FF333922661FFF3A" docLanguage="en" docName="zootaxa.4258.1.1SLASH10897.pdf" docOrigin="Zootaxa 4258 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybella bedosae Judson, 2017, n. sp." docType="treatment" docUuid="E40C8CA5-237F-41B3-B7E1-D108F163933E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="20" masterDocId="FFEF24736573334CFFA43E146234FFEF" masterDocTitle="A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="14" updateTime="1698439057093" updateUser="ExternalLinkService">
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<mods:title id="B73D61EB6D316B00B457E4692B554929">A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia</mods:title>
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<mods:name id="1D792949CC1A001D9207B4A09FD89C31" type="personal">
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<mods:roleTerm id="74BA1E37AA118D348602D37FC3E023E0">Author</mods:roleTerm>
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<mods:namePart id="F98E3CF21B7CC119D2982163A2BE2B81">Judson, Mark L. I.</mods:namePart>
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<mods:typeOfResource id="C644275E7A0CEDA5E6640315BC18F23F">text</mods:typeOfResource>
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<mods:title id="46AF541FABC73261F92EFC4B9E203441">Zootaxa</mods:title>
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<mods:part id="BBF6A3A29E21F1F3D8E3120931D9574A">
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<mods:date id="392B21491567C50BAA5DF4EF54EE1807">2017</mods:date>
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<mods:number id="1C45B824D6AE1ADDBA4789C139EDAA5E">4258</mods:number>
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<mods:identifier id="7A4DF1E7DD6183D33AF7B3CE5F44AFE8" type="DOI">10.11646/zootaxa.4258.1.1</mods:identifier>
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<mods:identifier id="EE8C90780073B0AB245FABD027FD1B88" type="Zenodo-Dep">569114</mods:identifier>
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<treatment id="03D65C0B657E3358FF333922661FFF3A" ID-DOI="http://doi.org/10.5281/zenodo.6023805" ID-GBIF-Taxon="128699664" ID-Zenodo-Dep="6023805" LSID="urn:lsid:zoobank.org:act:E40C8CA5-237F-41B3-B7E1-D108F163933E" httpUri="http://treatment.plazi.org/id/03D65C0B657E3358FF333922661FFF3A" lastPageId="20" lastPageNumber="20" pageId="13" pageNumber="14">
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<subSubSection id="C365BE96657E3341FF333922630AF89D" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BC0ED1D657E3341FF3339226397F8BE" blockId="13.[151,419,1846,1906]" box="[151,419,1846,1873]" pageId="13" pageNumber="14">
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<heading id="D0885A71657E3341FF3339226397F8BE" bold="true" box="[151,419,1846,1873]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B90B310F657E3341FF3339226397F8BE" bold="true" box="[151,419,1846,1873]" pageId="13" pageNumber="14">
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<taxonomicName id="4C7F969E657E3341FF333922636DF8BF" ID-CoL="32RLH" box="[151,345,1846,1872]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
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<emphasis id="B90B310F657E3341FF333922636DF8BF" bold="true" box="[151,345,1846,1872]" italics="true" pageId="13" pageNumber="14">Cybella bedosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C74657E3341FEC539236397F8BE" box="[353,419,1847,1873]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC0ED1D657E3341FF33394D630AF89D" blockId="13.[151,419,1846,1906]" box="[151,318,1881,1906]" pageId="13" pageNumber="14">
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<figureCitation id="1344F198657E3341FF33394D62D8F89D" box="[151,236,1881,1906]" captionStart="FIGURES 48 ‒ 49" captionStartId="14.[151,264,1428,1451]" captionTargetBox="[151,1435,625,1406]" captionTargetId="figure@14.[151,1436,625,1407]" captionTargetPageId="14" captionText="FIGURES 48 ‒ 49. Cybella bedosae n. sp. 48, habitus of male, dorsal view; 49, type locality, Kien Krol, Cambodia (photograph taken 23 June 2008); cave (not visible) is at base of hill." httpUri="https://zenodo.org/record/569123/files/figure.png" pageId="13" pageNumber="14">Figs 48</figureCitation>
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,
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<figureCitation id="1344F198657E3341FF5C394D630AF89D" box="[248,318,1881,1906]" captionStart-0="FIGURES 50 – 59" captionStart-1="FIGURES 60 – 69" captionStart-2="FIGURES 70 – 71" captionStart-3="FIGURES 72 – 73" captionStart-4="FIGURES 74 – 87" captionStartId-0="15.[151,264,1701,1724]" captionStartId-1="16.[151,264,1823,1846]" captionStartId-2="17.[151,264,688,711]" captionStartId-3="17.[151,264,1558,1581]" captionStartId-4="18.[151,264,1722,1745]" captionTargetBox-0="[233,1333,185,1671]" captionTargetBox-1="[219,1368,206,1763]" captionTargetBox-2="[157,1434,216,660]" captionTargetBox-3="[158,1410,786,1534]" captionTargetBox-4="[211,1365,216,1664]" captionTargetId-0="figure@15.[228,1338,181,1681]" captionTargetId-1="figure@16.[214,1376,199,1776]" captionTargetId-2="figure@17.[151,1436,193,667]" captionTargetId-3="figure@17.[153,1416,779,1545]" captionTargetId-4="figure@18.[207,1368,205,1675]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="17" captionTargetPageId-4="18" captionText-0="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." captionText-1="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." captionText-2="FIGURES 70 – 71. Cybella bedosae n. sp., male, legs. 70, right leg IV, antiaxial view; 71, right leg I, antiaxial view. Scale line 0.2 mm." captionText-3="FIGURES 72 – 73. Cybella bedosae n. sp., protonymph. 72, carapace and anterior segments (I – III) of opisthosoma, dorsal view, reticulation of tergite I only partially drawn; 73, leg coxae, ventral view. Abbreviation: pcs, primary coxal spine. Scale lines 0.1 mm." captionText-4="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri-0="https://zenodo.org/record/569124/files/figure.png" httpUri-1="https://zenodo.org/record/569125/files/figure.png" httpUri-2="https://zenodo.org/record/569126/files/figure.png" httpUri-3="https://zenodo.org/record/569127/files/figure.png" httpUri-4="https://zenodo.org/record/569128/files/figure.png" pageId="13" pageNumber="14">50‒87</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C365BE96657E3342FF3339B567BFFF17" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8BC0ED1D657E3342FF3339B567BFFF17" blockId="13.[151,1436,1953,2014]" lastBlockId="14.[151,1437,151,573]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<emphasis id="B90B310F657E3341FF3339B56325F855" bold="true" box="[151,273,1953,1978]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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Very similar to
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<taxonomicName id="4C7F969E657E3341FE6C39B6605CF855" box="[456,616,1953,1978]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
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<emphasis id="B90B310F657E3341FE6C39B6605CF855" box="[456,616,1953,1978]" italics="true" pageId="13" pageNumber="14">C. deharvengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C74657E3341FDCB39B5609BF855" box="[623,687,1953,1978]" pageId="13" pageNumber="14" rank="species">
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<emphasis id="B90B310F657E3341FDCB39B5609BF855" bold="true" box="[623,687,1953,1978]" pageId="13" pageNumber="14">n. sp.</emphasis>
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</taxonomicNameLabel>
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, but larger (e.g. male palp femur length 0.573 versus
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<quantity id="4C8740F8657E3341FAB439B562F3F831" metricMagnitude="-4" metricUnit="m" metricValue="5.075" metricValueMax="5.22" metricValueMin="4.93" pageId="13" pageNumber="14" unit="mm" value="0.5075" valueMax="0.522" valueMin="0.493">0.493–0.522 mm</quantity>
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), with chela less robust (4.8 times longer than broad, versus 4.3–4.5), most segments of legs more attenuate (e.g. femur of leg IV 2.8 times longer than broad, versus 2.0–2.1), trichobothrium
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<emphasis id="B90B310F657D3342FBC73E836648FF5F" box="[1123,1148,151,176]" italics="true" pageId="14" pageNumber="15">sb</emphasis>
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slightly proximad of pit (versus below pit), trichobothria
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<emphasis id="B90B310F657D3342FD853EA86008FF3A" box="[545,572,188,213]" italics="true" pageId="14" pageNumber="15">eb</emphasis>
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and
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<emphasis id="B90B310F657D3342FDDB3EA86090FF3A" box="[639,676,188,213]" italics="true" pageId="14" pageNumber="15">esb</emphasis>
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separated by more than one bothridial diameter (versus half a diameter), anterior genital sternite with 12 setae (versus 8), and lateral diverticulum of male atrial plate of longer.
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</paragraph>
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</subSubSection>
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<subSubSection id="C365BE96657D3342FF633F1061EBFEAF" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="8BC0ED1D657D3342FF633F1061EBFEAF" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="15">
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<emphasis id="B90B310F657D3342FF633F10637AFEF2" bold="true" box="[199,334,260,285]" pageId="14" pageNumber="15">Etymology.</emphasis>
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This species is named after Anne Bedos (MNHN), who helped collect the
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<typeStatus id="54C453BF657D3342FB673F1166CAFEF2" box="[1219,1278,261,285]" pageId="14" pageNumber="15">types</typeStatus>
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and who has contributed much to our knowledge of the soil and cave faunas of SE Asia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C365BE96657D3342FF633F586140FDD3" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8BC0ED1D657D3342FF633F5860BFFE3F" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="15">
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<materialsCitation id="3B17E740657D3342FF633F5866AFFE42" ID-GBIF-Occurrence="1493295184" collectingDate="2005-11-20" collectionCode="MNHN" collectorName="L. Deharveng & A. Bedos" country="Cambodia" county="Kampuchea" elevation="45" location="Kien Krol" municipality="Phnom Laang" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Kampot" typeStatus="holotype">
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<emphasis id="B90B310F657D3342FF633F586348FE8A" bold="true" box="[199,380,332,357]" pageId="14" pageNumber="15">
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<typeStatus id="54C453BF657D3342FF633F586336FE8A" box="[199,258,332,357]" pageId="14" pageNumber="15">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54C453BF657D3342FE2F3F5863CCFE8A" box="[395,504,332,357]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D792694657D3342FDAD3F596014FE8B" box="[521,544,333,356]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
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,
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<collectingCountry id="F368AD8D657D3342FD903F58609EFE8A" box="[564,682,332,357]" name="Cambodia" pageId="14" pageNumber="15">Cambodia</collectingCountry>
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,
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<collectingCounty id="62A19591657D3342FD183F586171FE8A" box="[700,837,332,357]" pageId="14" pageNumber="15">Kampuchea</collectingCounty>
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,
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<collectingRegion id="49BB23FF657D3342FCFC3F596181FE8B" box="[856,949,333,356]" country="Cambodia" name="Kampot" pageId="14" pageNumber="15">Kampot</collectingRegion>
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,
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<collectingMunicipality id="6BA47767657D3342FC633F58665AFE8A" box="[967,1134,332,357]" pageId="14" pageNumber="15">Phnom Laang</collectingMunicipality>
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,
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<location id="8EA0BBC6657D3342FB253F5866C8FE8A" LSID="urn:lsid:plazi:treatment:03D65C0B657E3358FF333922661FFF3A:8EA0BBC6657D3342FB253F5866C8FE8A" box="[1153,1276,332,357]" country="Cambodia" county="Kampuchea" municipality="Phnom Laang" name="Kien Krol" pageId="14" pageNumber="15" stateProvince="Kampot">Kien Krol</location>
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, 1042ʹ14.4ʺN 10420ʹ41.1ʺE,
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<elevation id="00520A2E657D3342FE993F7B6386FE67" box="[317,434,367,392]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="15" unit="m" value="45.0">
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<quantity id="4C8740F8657D3342FE993F7B6342FE68" box="[317,374,367,392]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="15" unit="m" value="45.0">45 m</quantity>
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a.s.l.
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</elevation>
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,
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<location id="8EA0BBC6657D3342FE1A3F7B6026FE67" LSID="urn:lsid:plazi:treatment:03D65C0B657E3358FF333922661FFF3A:8EA0BBC6657D3342FE1A3F7B6026FE67" box="[446,530,367,392]" country="Cambodia" county="Kampuchea" municipality="Phnom Laang" name="Berlese" pageId="14" pageNumber="15" stateProvince="Kampot">Berlese</location>
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extraction of soil from low passage (“boyau”) on left side of cave,
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<collectingDate id="EF853235657D3342FAA73F6462E6FE42" pageId="14" pageNumber="15" value="2005-11-20">
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<date id="FFC1CBDD657D3342FAA73F6462E6FE42" pageId="14" pageNumber="15" value="2005-11-20">20 November 2005</date>
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</collectingDate>
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, leg.
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<collectorName id="268A88CB657D3342FEB63F81639AFE42" box="[274,430,404,429]" pageId="14" pageNumber="15">L. Deharveng</collectorName>
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&
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<collectorName id="268A88CB657D3342FE733F806075FE42" box="[471,577,404,429]" pageId="14" pageNumber="15">A. Bedos</collectorName>
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(
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<collectionCode id="ED6E75D8657D3342FDF63F80609EFE43" box="[594,682,404,428]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
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Ps116-01; collectors’ number KAM05-21)
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</materialsCitation>
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. Paratype protonymph (MNHN Ps116-02), collected with holotype.
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</paragraph>
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<paragraph id="8BC0ED1D657D3342FF633FC86140FDD3" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="15">
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<emphasis id="B90B310F657D3342FF633FC863B9FE1A" bold="true" box="[199,397,476,501]" pageId="14" pageNumber="15">
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<collectionCode id="ED6E75D8657D3342FF633FC86330FE1A" box="[199,260,476,501]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="14" pageNumber="15">DNA</collectionCode>
|
||
sequences.
|
||
</emphasis>
|
||
Partial sequences, obtained by non-destructive extraction from the palp of the
|
||
<typeStatus id="54C453BF657D3342FA9F3FC967A8FE1A" box="[1339,1436,477,501]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
protonymph (voucher no. MNHN-JAD72), were deposited in GenBank for 18S RNA (accession no. JN018313.1) and 28S RNA (accession no. JN018410.1) by
|
||
<bibRefCitation id="EFEE90EC657D3342FD383C306144FDD3" box="[668,880,548,573]" pageId="14" pageNumber="15" refString="Arabi, J., Judson, M. L. I., Deharveng, L., Lourenco, W. R., Cruaud, C. & Hassanin, A. (2012) Nucleotide composition of CO 1 sequences in Chelicerata (Arthropoda): detecting new mitogenomic rearrangements. Journal of Molecular Evolution, 74, 81 - 95." type="journal article">
|
||
Arabi
|
||
<emphasis id="B90B310F657D3342FD473C31612FFDD2" box="[739,795,548,573]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF00BD95657D3342FF333B8060FAFA26" httpUri="https://zenodo.org/record/569123/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1435,625,1406]" targetPageId="14">
|
||
<paragraph id="8BC0ED1D657D3342FF333B8060FAFA26" blockId="14.[151,1436,1428,1481]" pageId="14" pageNumber="15">
|
||
<emphasis id="B90B310F657D3342FF333B806367FA45" bold="true" box="[151,339,1428,1451]" pageId="14" pageNumber="15">FIGURES 48‒49.</emphasis>
|
||
<taxonomicName id="4C7F969E657D3342FEFD3B806034FA45" box="[345,512,1428,1450]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FEFD3B806034FA45" box="[345,512,1428,1450]" italics="true" pageId="14" pageNumber="15">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FDA23B81600BFA45" box="[518,575,1429,1450]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FDA23B81600BFA45" bold="true" box="[518,575,1429,1450]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
48, habitus of male, dorsal view; 49, type locality, Kien Krol, Cambodia (photograph taken 23 June 2008); cave (not visible) is at base of hill.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C365BE96657D335FFF633BE46604F8E5" lastPageId="19" lastPageNumber="20" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8BC0ED1D657D3342FF633BE4638EF89E" blockId="14.[151,1436,1520,1977]" pageId="14" pageNumber="15">
|
||
<emphasis id="B90B310F657D3342FF633BE46385F9E6" bold="true" box="[199,433,1520,1545]" pageId="14" pageNumber="15">Description of male</emphasis>
|
||
(limited to characters showing differences from
|
||
<taxonomicName id="4C7F969E657D3342FC723BE56641F9E6" box="[982,1141,1520,1545]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FC723BE56641F9E6" box="[982,1141,1520,1545]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FBD83BE4668FF9E6" box="[1148,1211,1520,1545]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FBD83BE4668FF9E6" bold="true" box="[1148,1211,1520,1545]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
). Tergal and sternal chaetotaxy not determined due to accumulation of exocuticle and debris, but where visible numbers of setae similar to those seen in
|
||
<taxonomicName id="4C7F969E657D3342FEEB382D63DAF9BE" box="[335,494,1592,1617]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FEEB382D63DAF9BE" box="[335,494,1592,1617]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FE52382C6002F9BE" box="[502,566,1592,1617]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FE52382C6002F9BE" bold="true" box="[502,566,1592,1617]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Coxa I with 1 primary and 10–12 secondary spines around pit (
|
||
<figureCitation id="1344F198657D3342FAAF382C67BBF9BE" box="[1291,1423,1592,1617]" captionStart="FIGURES 50 – 59" captionStartId="15.[151,264,1701,1724]" captionTargetBox="[233,1333,185,1671]" captionTargetId="figure@15.[228,1338,181,1681]" captionTargetPageId="15" captionText="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="14" pageNumber="15">Figs 52, 55</figureCitation>
|
||
). Coxal setae (
|
||
<figureCitation id="1344F198657D3342FE833848634FF99A" box="[295,379,1628,1653]" captionStart="FIGURES 50 – 59" captionStartId="15.[151,264,1701,1724]" captionTargetBox="[233,1333,185,1671]" captionTargetId="figure@15.[228,1338,181,1681]" captionTargetPageId="15" captionText="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="14" pageNumber="15">Fig. 52</figureCitation>
|
||
): P 3 on manducatory process + 32 (total 35),
|
||
<date id="FFC1CBDD657D3342FCDA384961AAF99A" box="[894,926,1629,1653]" pageId="14" pageNumber="15">I 8</date>
|
||
,
|
||
<date id="FFC1CBDD657D3342FC0E384961E5F99A" box="[938,977,1629,1653]" pageId="14" pageNumber="15">II 9</date>
|
||
(1 lateral),
|
||
<date id="FFC1CBDD657D3342FBF7384966B7F99B" box="[1107,1155,1629,1652]" pageId="14" pageNumber="15">III 7</date>
|
||
(1 lateral), 10 (1 lateral). Anterior genital sternite (
|
||
<figureCitation id="1344F198657D3342FE123894603EF976" box="[438,522,1664,1689]" captionStart="FIGURES 50 – 59" captionStartId="15.[151,264,1701,1724]" captionTargetBox="[233,1333,185,1671]" captionTargetId="figure@15.[228,1338,181,1681]" captionTargetPageId="15" captionText="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="14" pageNumber="15">Fig. 52</figureCitation>
|
||
) with 12 setae, posterior genital sternite (
|
||
<figureCitation id="1344F198657D3342FC413894660CF976" box="[997,1080,1664,1689]" captionStart="FIGURES 50 – 59" captionStartId="15.[151,264,1701,1724]" captionTargetBox="[233,1333,185,1671]" captionTargetId="figure@15.[228,1338,181,1681]" captionTargetPageId="15" captionText="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="14" pageNumber="15">Fig. 52</figureCitation>
|
||
) with 14 setae (2+10+2). Male genitalia (
|
||
<figureCitation id="1344F198657D3342FEA238B06363F952" box="[262,343,1700,1725]" captionStart="FIGURES 50 – 59" captionStartId="15.[151,264,1701,1724]" captionTargetBox="[233,1333,185,1671]" captionTargetId="figure@15.[228,1338,181,1681]" captionTargetPageId="15" captionText="FIGURES 50 – 59. Cybella bedosae n. sp., male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II ‒ III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right lgs omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations: ap, apodeme; da, dorsal apodeme; ejca, ejaculatory canal atrium; gp, gland pore; ic, intercoxal canal; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pcs, primary coxal spine; scs, secondary coxal spine; sp, spiracle; sps, spatulate seta; sr, sclerotized ring; tp, posteromedian tecta of palp coxae; tr, trachea; w, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53 ‒ 59)." httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="14" pageNumber="15">Fig. 58</figureCitation>
|
||
) very similar to those of
|
||
<taxonomicName id="4C7F969E657D3342FDCB38B16139F952" box="[623,781,1700,1725]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FDCB38B16139F952" box="[623,781,1700,1725]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FCB038B06167F952" box="[788,851,1700,1725]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FCB038B06167F952" bold="true" box="[788,851,1700,1725]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but comparison of dorsal apodeme, lateral rods and atrial plate hampered by differences in orientation in the specimens examined (this depending on the degree to which the genital plates are open or closed); lateral sacs longer and thinner than those of
|
||
<taxonomicName id="4C7F969E657D3342FB0338F9677DF8EA" box="[1191,1353,1772,1797]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FB0338F9677DF8EA" box="[1191,1353,1772,1797]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FAF638F867A0F8EA" box="[1362,1428,1772,1797]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FAF638F867A0F8EA" bold="true" box="[1362,1428,1772,1797]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; lateral pockets of atrial plate slightly longer than those of
|
||
<taxonomicName id="4C7F969E657D3342FCE1390561DCF8C6" box="[837,1000,1808,1833]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FCE1390561DCF8C6" box="[837,1000,1808,1833]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FC5739046601F8C6" box="[1011,1077,1808,1833]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FC5739046601F8C6" bold="true" box="[1011,1077,1808,1833]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; posterior plate with 1 pair of modified basal setae, which seems to be more rounded at the tip than those of
|
||
<taxonomicName id="4C7F969E657D3342FBBC3921668DF8A2" box="[1048,1209,1844,1869]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F657D3342FBBC3921668DF8A2" box="[1048,1209,1844,1869]" italics="true" pageId="14" pageNumber="15">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657D3342FB6539206735F8A2" box="[1217,1281,1844,1869]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B90B310F657D3342FB6539206735F8A2" bold="true" box="[1217,1281,1844,1869]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and 2 small, simple setae on each side.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D657D335DFF63396861A2F8F3" blockId="14.[151,1436,1520,1977]" lastBlockId="17.[151,1437,1687,2000]" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
|
||
Chelicera (
|
||
<figureCitation id="1344F198657D3342FEE4396863F4F87A" box="[320,448,1916,1941]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="14" pageNumber="15">Figs 60–64</figureCitation>
|
||
) with 6‒7 small setae on palm, in addition to the 5 large setae. Galeal seta (
|
||
<emphasis id="B90B310F657D3342FAA1396A672AF87A" box="[1285,1310,1918,1941]" italics="true" pageId="14" pageNumber="15">gs</emphasis>
|
||
) 0.85 from base, extending well beyond tip of spinneret (by nearly half length of hair). Spinneret (
|
||
<figureCitation id="1344F198657D3342FBC539B466D5F856" box="[1121,1249,1952,1977]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="14" pageNumber="15">Figs 61‒62</figureCitation>
|
||
) slightly thicker than in
|
||
<taxonomicName id="4C7F969E6562335DFF54388C63A5F95F" box="[240,401,1687,1712]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F6562335DFF54388C63A5F95F" box="[240,401,1687,1712]" italics="true" pageId="17" pageNumber="18">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746562335DFE3E388363EFF95F" box="[410,475,1687,1712]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B90B310F6562335DFE3E388363EFF95F" bold="true" box="[410,475,1687,1712]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but still tubular (i.e. not blade-shaped), length 16 µm. Serrula exterior with 19–20 blades, plus small distally-directed apical process. Serrula interior with 17 internal ducts; apically with a distallydirected process, followed by two rows of three, low, rounded mounds; basal blades forming a velum. Rallum with 1 (right chelicera) or 2 (left chelicera;
|
||
<figureCitation id="1344F1986562335DFDE7391760A7F8F3" box="[579,659,1795,1820]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="17" pageNumber="18">Fig. 63</figureCitation>
|
||
) blades, length 23 µm.
|
||
</paragraph>
|
||
<caption id="DF00BD95657C3343FF3338B16393F800" httpUri="https://zenodo.org/record/569124/files/figure.png" pageId="15" pageNumber="16" targetBox="[233,1333,185,1671]" targetPageId="15">
|
||
<paragraph id="8BC0ED1D657C3343FF3338B16393F800" blockId="15.[151,1436,1701,2031]" pageId="15" pageNumber="16">
|
||
<emphasis id="B90B310F657C3343FF3338B16360F954" bold="true" box="[151,340,1701,1724]" pageId="15" pageNumber="16">FIGURES 50–59.</emphasis>
|
||
<taxonomicName id="4C7F969E657C3343FEF838B26031F953" box="[348,517,1702,1724]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F657C3343FEF838B26031F953" box="[348,517,1702,1724]" italics="true" pageId="15" pageNumber="16">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C74657C3343FDA838B36072F953" box="[524,582,1703,1724]" pageId="15" pageNumber="16" rank="species">
|
||
<emphasis id="B90B310F657C3343FDA838B36072F953" bold="true" box="[524,582,1703,1724]" pageId="15" pageNumber="16">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male. 50, anterior part of carapace, dorsal view (setae omitted); 51, tapeta of right eyes, ventral (internal) view; 52, leg coxae and median parts of sternites II‒III, ventral view; 53, intercoxal canal between posterior parts of palp coxae (membrane torn distally); 54, manducatory process of right palp coxa, ventral view (lyrifissure abnormally doubled); 55, primary spine and adjacent secondary spines of right coxa I; 56, left spiracle and adjacent apodeme of segment III, ventral view; 57, membranous part of intercoxal canal between coxa I and palp coxae (latter removed), ventral view; 58, genitalia, ventral view (right
|
||
<emphasis id="B90B310F657C3343FD95392B607AF8BA" box="[561,590,1855,1877]" italics="true" pageId="15" pageNumber="16">lgs</emphasis>
|
||
omitted; part of anterior wall of genital atrium removed on left of drawing); 59, tergite (upper) and sternite (lower) of segment XII, ventral view. Abbreviations:
|
||
<emphasis id="B90B310F657C3343FC50394B6639F89B" box="[1012,1037,1887,1908]" italics="true" pageId="15" pageNumber="16">ap</emphasis>
|
||
, apodeme;
|
||
<emphasis id="B90B310F657C3343FB28394A6691F89B" box="[1164,1189,1886,1908]" italics="true" pageId="15" pageNumber="16">da</emphasis>
|
||
, dorsal apodeme;
|
||
<emphasis id="B90B310F657C3343FACF394B67A1F89B" box="[1387,1429,1887,1908]" italics="true" pageId="15" pageNumber="16">ejca</emphasis>
|
||
, ejaculatory canal atrium;
|
||
<emphasis id="B90B310F657C3343FE3F396A6380F87C" box="[411,436,1918,1939]" italics="true" pageId="15" pageNumber="16">gp</emphasis>
|
||
, gland pore;
|
||
<emphasis id="B90B310F657C3343FD9F396A6079F87C" box="[571,589,1918,1939]" italics="true" pageId="15" pageNumber="16">ic</emphasis>
|
||
, intercoxal canal;
|
||
<emphasis id="B90B310F657C3343FCAF396A611CF87C" box="[779,808,1918,1939]" italics="true" pageId="15" pageNumber="16">inv</emphasis>
|
||
, invagination at side of atrial plate;
|
||
<emphasis id="B90B310F657C3343FB3E3968668CF87D" box="[1178,1208,1916,1938]" italics="true" pageId="15" pageNumber="16">lgs</emphasis>
|
||
, lateral genital sac;
|
||
<emphasis id="B90B310F657C3343FA21396867A2F87D" box="[1413,1430,1916,1938]" italics="true" pageId="15" pageNumber="16">ly</emphasis>
|
||
, lyrifissure;
|
||
<emphasis id="B90B310F657C3343FEA839886301F85E" box="[268,309,1948,1969]" italics="true" pageId="15" pageNumber="16">mgs</emphasis>
|
||
, median genital sac;
|
||
<emphasis id="B90B310F657C3343FDB539886007F85E" box="[529,563,1948,1969]" italics="true" pageId="15" pageNumber="16">pcs</emphasis>
|
||
, primary coxal spine;
|
||
<emphasis id="B90B310F657C3343FCB83988610FF85E" box="[796,827,1948,1969]" italics="true" pageId="15" pageNumber="16">scs</emphasis>
|
||
, secondary coxal spine;
|
||
<emphasis id="B90B310F657C3343FB9E39886664F85E" box="[1082,1104,1948,1969]" italics="true" pageId="15" pageNumber="16">sp</emphasis>
|
||
, spiracle;
|
||
<emphasis id="B90B310F657C3343FB18398866E8F85E" box="[1212,1244,1948,1969]" italics="true" pageId="15" pageNumber="16">sps</emphasis>
|
||
, spatulate seta;
|
||
<emphasis id="B90B310F657C3343FA27398867A2F85E" box="[1411,1430,1948,1969]" italics="true" pageId="15" pageNumber="16">sr</emphasis>
|
||
, sclerotized ring;
|
||
<emphasis id="B90B310F657C3343FEE439AF6360F83F" box="[320,340,1979,2000]" italics="true" pageId="15" pageNumber="16">tp</emphasis>
|
||
, posteromedian tecta of palp coxae;
|
||
<emphasis id="B90B310F657C3343FD6039AF60E0F83F" box="[708,724,1979,2000]" italics="true" pageId="15" pageNumber="16">tr</emphasis>
|
||
, trachea;
|
||
<emphasis id="B90B310F657C3343FC9239AF6171F83F" box="[822,837,1979,2000]" italics="true" pageId="15" pageNumber="16">w</emphasis>
|
||
, wall of coxa in cross-section. Scale lines 0.1 mm (Figs 50, 52), 0.05 (Figs 51, 53‒59).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00BD956563335CFF33390B67B5F801" httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="16" pageNumber="17" targetBox="[219,1368,206,1763]" targetPageId="16">
|
||
<paragraph id="8BC0ED1D6563335CFF33390B67B5F801" blockId="16.[151,1436,1823,2030]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90B310F6563335CFF33390B6362F8DA" bold="true" box="[151,342,1823,1846]" pageId="16" pageNumber="17">FIGURES 60–69.</emphasis>
|
||
<taxonomicName id="4C7F969E6563335CFEFB3934603EF8D9" box="[351,522,1824,1846]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6563335CFEFB3934603EF8D9" box="[351,522,1824,1846]" italics="true" pageId="16" pageNumber="17">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746563335CFDB039346064F8DA" box="[532,592,1824,1845]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B90B310F6563335CFDB039346064F8DA" bold="true" box="[532,592,1824,1845]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations:
|
||
<emphasis id="B90B310F6563335CFADA396867A1F87D" box="[1406,1429,1916,1938]" italics="true" pageId="16" pageNumber="17">dx</emphasis>
|
||
, duplex trichobothria,
|
||
<emphasis id="B90B310F6563335CFED0398F63BFF85F" box="[372,395,1947,1968]" italics="true" pageId="16" pageNumber="17">pc</emphasis>
|
||
, coupled sensilla;
|
||
<emphasis id="B90B310F6563335CFDE8398F6056F85F" box="[588,610,1947,1968]" italics="true" pageId="16" pageNumber="17">pr</emphasis>
|
||
, distal process of serrula exterior;
|
||
<emphasis id="B90B310F6563335CFC6C398F61C2F85F" box="[968,1014,1947,1968]" italics="true" pageId="16" pageNumber="17">s.ext</emphasis>
|
||
, serrula exterior;
|
||
<emphasis id="B90B310F6563335CFB0A398F66EDF85F" box="[1198,1241,1947,1968]" italics="true" pageId="16" pageNumber="17">s.int</emphasis>
|
||
, serrula interior;
|
||
<emphasis id="B90B310F6563335CFA29398F67A2F85F" box="[1421,1430,1947,1968]" italics="true" pageId="16" pageNumber="17">r</emphasis>
|
||
, rallum. Other abbreviations (
|
||
<emphasis id="B90B310F6563335CFE1939AD63E1F820" box="[445,469,1977,1999]" italics="true" pageId="16" pageNumber="17">eb</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFE4639AD6030F820" box="[482,516,1977,1999]" italics="true" pageId="16" pageNumber="17">esb</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFDB539AE6019F820" box="[529,557,1978,1999]" italics="true" pageId="16" pageNumber="17">est</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFD9E39AE6078F820" box="[570,588,1978,1999]" italics="true" pageId="16" pageNumber="17">et</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFDFE39AD605AF820" box="[602,622,1977,1999]" italics="true" pageId="16" pageNumber="17">ib</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFDDE39AD60A3F820" box="[634,663,1977,1999]" italics="true" pageId="16" pageNumber="17">isb</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFD0039AE6088F820" box="[676,700,1978,1999]" italics="true" pageId="16" pageNumber="17">ist</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFD6E39AE60ECF820" box="[714,728,1978,1999]" italics="true" pageId="16" pageNumber="17">it</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFD4039AD60C5F820" box="[740,753,1977,1999]" italics="true" pageId="16" pageNumber="17">b</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFD5A39AD6120F820" box="[766,788,1977,1999]" italics="true" pageId="16" pageNumber="17">sb</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFC8639AE6107F820" box="[802,819,1978,1999]" italics="true" pageId="16" pageNumber="17">st</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6563335CFCE439AE6173F820" box="[832,839,1978,1999]" italics="true" pageId="16" pageNumber="17">t</emphasis>
|
||
) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65‒66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61‒63, 67), 0.01 (Fig. 69) (figures 61‒63 share same scale line).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00BD956562335DFF333CA462DFFD0A" httpUri="https://zenodo.org/record/569126/files/figure.png" pageId="17" pageNumber="18" targetBox="[157,1434,216,660]" targetPageId="17">
|
||
<paragraph id="8BC0ED1D6562335DFF333CA462DFFD0A" blockId="17.[151,1435,688,741]" pageId="17" pageNumber="18">
|
||
<emphasis id="B90B310F6562335DFF333CA46360FD2A" bold="true" box="[151,340,688,711]" pageId="17" pageNumber="18">FIGURES 70–71.</emphasis>
|
||
<taxonomicName id="4C7F969E6562335DFEFE3CA46036FD29" box="[346,514,688,710]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6562335DFEFE3CA46036FD29" box="[346,514,688,710]" italics="true" pageId="17" pageNumber="18">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746562335DFDAD3CA56076FD29" box="[521,578,689,710]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B90B310F6562335DFDAD3CA56076FD29" bold="true" box="[521,578,689,710]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male, legs. 70, right leg IV, antiaxial view; 71, right leg I, antiaxial view. Scale line 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00BD956562335DFF3338026316F985" httpUri="https://zenodo.org/record/569127/files/figure.png" pageId="17" pageNumber="18" targetBox="[158,1410,786,1534]" targetPageId="17">
|
||
<paragraph id="8BC0ED1D6562335DFF3338026316F985" blockId="17.[151,1436,1558,1642]" pageId="17" pageNumber="18">
|
||
<emphasis id="B90B310F6562335DFF3338026363F9C3" bold="true" box="[151,343,1558,1581]" pageId="17" pageNumber="18">FIGURES 72–73.</emphasis>
|
||
<taxonomicName id="4C7F969E6562335DFEC43803603FF9C2" box="[352,523,1559,1581]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6562335DFEC43803603FF9C2" box="[352,523,1559,1581]" italics="true" pageId="17" pageNumber="18">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746562335DFDB138036065F9C3" box="[533,593,1559,1580]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B90B310F6562335DFDB138036065F9C3" bold="true" box="[533,593,1559,1580]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, protonymph. 72, carapace and anterior segments (I–III) of opisthosoma, dorsal view, reticulation of tergite I only partially drawn; 73, leg coxae, ventral view. Abbreviation:
|
||
<emphasis id="B90B310F6562335DFBFF38236648F9A3" box="[1115,1148,1591,1612]" italics="true" pageId="17" pageNumber="18">pcs</emphasis>
|
||
, primary coxal spine. Scale lines 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0ED1D6562335DFF633933666DF83F" blockId="17.[151,1437,1687,2000]" pageId="17" pageNumber="18">
|
||
Palp as illustrated (
|
||
<figureCitation id="1344F1986562335DFE0239336018F8AF" box="[422,556,1831,1856]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="17" pageNumber="18">Figs 65–69</figureCitation>
|
||
). Fixed chelal finger with 12 chemosensory setae in antiaxial row. Movable finger with 14–16 long (22–24 µm), spatulate, chemosensory setae in pit (
|
||
<figureCitation id="1344F1986562335DFC7A395F6607F88B" box="[990,1075,1867,1892]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="17" pageNumber="18">Fig. 67</figureCitation>
|
||
). Trichobothria (
|
||
<figureCitation id="1344F1986562335DFB54395F6745F88B" box="[1264,1393,1867,1892]" captionStart="FIGURES 60 – 69" captionStartId="16.[151,264,1823,1846]" captionTargetBox="[219,1368,206,1763]" captionTargetId="figure@16.[214,1376,199,1776]" captionTargetPageId="16" captionText="FIGURES 60 – 69. Cybella bedosae n. sp., male. 60, left chelicera, antiaxial view; 61, distal end of movable finger of left chelicera, antiaxial view; 62, spinneret of left chelicera, ventral view; 63, rallum of left chelicera, antiaxial view; 64, left chelicera, dorsal view; 65, left palp, minus chela, dorsal view; 66, left chela, dorsal view; 67, chemosensory setae of movable finger of left chela, antiaxial view; 68, left chela, antiaxial view; 69, distal sensillum of movable finger Abbreviations: dx, duplex trichobothria, pc, coupled sensilla; pr, distal process of serrula exterior; s. ext, serrula exterior; s. int, serrula interior; r, rallum. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines 0.2 mm (Figs 65 ‒ 66), 0.1 mm (Figs 60, 64, 68), 0.05 mm (Figs 61 ‒ 63, 67), 0.01 (Fig. 69) (figures 61 ‒ 63 share same scale line)." httpUri="https://zenodo.org/record/569125/files/figure.png" pageId="17" pageNumber="18">Figs 66, 68</figureCitation>
|
||
)
|
||
<emphasis id="B90B310F6562335DFA25395F67AFF88B" box="[1409,1435,1867,1892]" italics="true" pageId="17" pageNumber="18">eb</emphasis>
|
||
and
|
||
<emphasis id="B90B310F6562335DFF63397B62D8F867" box="[199,236,1903,1928]" italics="true" pageId="17" pageNumber="18">esb</emphasis>
|
||
separated by 1.2–1.8 bothridial diameters (absolute distance 17–22 µm);
|
||
<emphasis id="B90B310F6562335DFB8F397B6670F867" box="[1067,1092,1903,1928]" italics="true" pageId="17" pageNumber="18">sb</emphasis>
|
||
slightly proximad of posterior edge of pit. Fixed finger with 30 teeth: apodens plus 22 marginal, 6 paraxial and basal tuberculate tooth. Movable finger with 28 teeth: apodens plus 21 marginal, 7 paraxial and basal tuberculate tooth.
|
||
</paragraph>
|
||
<caption id="DF00BD956561335EFF3338AE63FEF808" httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="18" pageNumber="19" targetBox="[211,1365,216,1664]" targetPageId="18">
|
||
<paragraph id="8BC0ED1D6561335EFF3338AE63FEF808" blockId="18.[151,1437,1722,2023]" pageId="18" pageNumber="19">
|
||
<emphasis id="B90B310F6561335EFF3338AE6362F920" bold="true" box="[151,342,1722,1745]" pageId="18" pageNumber="19">FIGURES 74–87.</emphasis>
|
||
<taxonomicName id="4C7F969E6561335EFEFB38AE603DF93F" box="[351,521,1722,1744]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6561335EFEFB38AE603DF93F" box="[351,521,1722,1744]" italics="true" pageId="18" pageNumber="19">Cybella bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746561335EFDB638AF607AF93F" box="[530,590,1723,1744]" pageId="18" pageNumber="19" rank="species">
|
||
<emphasis id="B90B310F6561335EFDB638AF607AF93F" bold="true" box="[530,590,1723,1744]" pageId="18" pageNumber="19">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (
|
||
<emphasis id="B90B310F6561335EFAC53922675AF8A4" box="[1377,1390,1846,1867]" italics="true" pageId="18" pageNumber="19">p</emphasis>
|
||
1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations:
|
||
<emphasis id="B90B310F6561335EFE9839816365F845" box="[316,337,1941,1962]" italics="true" pageId="18" pageNumber="19">cs</emphasis>
|
||
, chemosensory seta;
|
||
<emphasis id="B90B310F6561335EFDE539806066F845" box="[577,594,1940,1962]" italics="true" pageId="18" pageNumber="19">ly</emphasis>
|
||
, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76‒77, 83, 86‒87), 0.05 mm (Figs 74‒75, 78‒82, 85; Figs 75 and 78‒82 share same scale line), 0.01 mm (Fig. 84).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0ED1D6560335FFF633E8C6005FF5F" blockId="19.[151,1437,151,2018]" box="[199,561,151,176]" pageId="19" pageNumber="20">
|
||
Legs as illustrated (
|
||
<figureCitation id="1344F1986560335FFE063E836011FF5F" box="[418,549,151,176]" captionStart="FIGURES 70 – 71" captionStartId="17.[151,264,688,711]" captionTargetBox="[157,1434,216,660]" captionTargetId="figure@17.[151,1436,193,667]" captionTargetPageId="17" captionText="FIGURES 70 – 71. Cybella bedosae n. sp., male, legs. 70, right leg IV, antiaxial view; 71, right leg I, antiaxial view. Scale line 0.2 mm." httpUri="https://zenodo.org/record/569126/files/figure.png" pageId="19" pageNumber="20">Figs 70‒71</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633EA9635CFE67" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">Measurements (in mm, standard ratios in parentheses). Body (contracted) 1.82×1.25 (1.5). Carapace 0.659×0.47 (1.4). Chelicera 0.234×0.141 (1.7), palm 0.159 (1.1), movable finger 0.094. Palp femur 0.573×0.298 (1.8), patella 0.453×0.178 (2.5), chela 0.665×0.138 (4.8), palm 0.162 (1.2), movable finger 0.474 (2.9). Leg I femur 0.256×0.079 (3.3), patella 0.214×0.090 (2.4), tibia 0.212×0.062 (3.4), tarsus 0.304×0.051 (5.9). Leg IV trochanter 0.289×0.140 (2.1), femur 0.246×0.087 (2.8), patella 0.298×0.110 (2.7), tibia 0.426×0.063 (6.8), tarsus 0.418×0.052 (8.1).</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633F80671EFD22" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90B310F6560335FFF633F806038FE42" bold="true" box="[199,524,404,429]" pageId="19" pageNumber="20">Description of protonymph</emphasis>
|
||
. Sclerotized parts pale straw-yellow, except tips of cheliceral fingers and teeth of chela, which are reddish-brown. Carapace (
|
||
<figureCitation id="1344F1986560335FFD2D3FA360E9FE3F" box="[649,733,439,464]" captionStart="FIGURES 72 – 73" captionStartId="17.[151,264,1558,1581]" captionTargetBox="[158,1410,786,1534]" captionTargetId="figure@17.[153,1416,779,1545]" captionTargetPageId="17" captionText="FIGURES 72 – 73. Cybella bedosae n. sp., protonymph. 72, carapace and anterior segments (I – III) of opisthosoma, dorsal view, reticulation of tergite I only partially drawn; 73, leg coxae, ventral view. Abbreviation: pcs, primary coxal spine. Scale lines 0.1 mm." httpUri="https://zenodo.org/record/569127/files/figure.png" pageId="19" pageNumber="20">Fig. 72</figureCitation>
|
||
) distinctly broadened posteriorly; less elongate than in adult; anteromedian projections small, anterolateral corners of carapace more rounded than in adult; two pairs of lyrifissures; two pairs of eyes with reflective tapeta, anterior pair sessile, posterior pair weakly raised, both pairs less prominent than in adult. Tergite I (
|
||
<figureCitation id="1344F1986560335FFDF93C306086FDD3" box="[605,690,548,573]" captionStart="FIGURES 72 – 73" captionStartId="17.[151,264,1558,1581]" captionTargetBox="[158,1410,786,1534]" captionTargetId="figure@17.[153,1416,779,1545]" captionTargetPageId="17" captionText="FIGURES 72 – 73. Cybella bedosae n. sp., protonymph. 72, carapace and anterior segments (I – III) of opisthosoma, dorsal view, reticulation of tergite I only partially drawn; 73, leg coxae, ventral view. Abbreviation: pcs, primary coxal spine. Scale lines 0.1 mm." httpUri="https://zenodo.org/record/569127/files/figure.png" pageId="19" pageNumber="20">Fig. 72</figureCitation>
|
||
) with same general form as in adult, but presence or absence of separate lateral plate could not be determined due to weak sclerotization; tergites II–VIII divided (II and VIII weakly so); setae difficult to count, but most half-tergites with about 6 setae; denticulation present on tergites VIII– IX. Pleural membranes with two folds, as in adult, lyrifissures (orientated dorsoventrally) on dorsal fold; only 1 gland pore seen (anteriorly on dorsal fold), but others might be present; no setae observed; plates absent.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633CC360C3FC4B" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
Palp coxa not as elongate as in adult, median maxillary lyrifissure present, with same form and position as that of adult; no lyrifissure distad of foramen. Coxa I with 1 primary and 9–10 secondary spines flanking pit (
|
||
<figureCitation id="1344F1986560335FFA9E3CE867BAFCFB" box="[1338,1422,764,789]" captionStart="FIGURES 72 – 73" captionStartId="17.[151,264,1558,1581]" captionTargetBox="[158,1410,786,1534]" captionTargetId="figure@17.[153,1416,779,1545]" captionTargetPageId="17" captionText="FIGURES 72 – 73. Cybella bedosae n. sp., protonymph. 72, carapace and anterior segments (I – III) of opisthosoma, dorsal view, reticulation of tergite I only partially drawn; 73, leg coxae, ventral view. Abbreviation: pcs, primary coxal spine. Scale lines 0.1 mm." httpUri="https://zenodo.org/record/569127/files/figure.png" pageId="19" pageNumber="20">Fig. 73</figureCitation>
|
||
); no pores in pit. Coxae III reduced in size, reaching closer to midline than in adult, overlapped by coxae II. Palp coxa with 12 setae (including 3 on manducatory process), leg coxae each with 3 setae. Sternites IV–X divided, setae 0: 2: 2+2: 3+3: 3+3: 4+4: 4+4: 4+4: 4+4: 4 (2 flanking anal tubercle): 2 (slightly closer together than those of tergite XI); denticulate granulation on IX–XI.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633DBB66B9FBDA" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
Chelicera (
|
||
<figureCitation id="1344F1986560335FFEE63DBB63F0FC27" box="[322,452,943,968]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Figs 74‒75</figureCitation>
|
||
) with distinction between smooth basal part and distal reticulate part of palm less well marked than in adult; three large (probably
|
||
<emphasis id="B90B310F6560335FFDDE3DC060B9FC02" box="[634,653,980,1005]" italics="true" pageId="19" pageNumber="20">ls</emphasis>
|
||
,
|
||
<emphasis id="B90B310F6560335FFD3E3DC06087FC02" box="[666,691,980,1005]" italics="true" pageId="19" pageNumber="20">sb</emphasis>
|
||
and
|
||
<emphasis id="B90B310F6560335FFD4C3DC160CBFC03" box="[744,767,981,1004]" italics="true" pageId="19" pageNumber="20">es</emphasis>
|
||
) and four small setae on palm; movable finger without seta; rallum of 2 blades (
|
||
<figureCitation id="1344F1986560335FFED33DE363F8FBFF" box="[375,460,1015,1040]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 75</figureCitation>
|
||
), as in adult; spinneret (
|
||
<figureCitation id="1344F1986560335FFD443DE36107FBFF" box="[736,819,1015,1040]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 74</figureCitation>
|
||
) simple, much longer (30 µm) than in adult, recurved apically. Serrula exterior with 10 blades plus apical process. Serrula interior with 10 ducts.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633A2B62D2FA29" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
Palp (
|
||
<figureCitation id="1344F1986560335FFEAB3A2B63A3FBB7" box="[271,407,1087,1112]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Figs 76‒84</figureCitation>
|
||
) similar to that of adult, trochanter with well developed horn, femur with moderate anterobasal projection. Chela with three trichobothria on fixed finger and one on movable finger. Trichobothrium
|
||
<date id="FFC1CBDD6560335FFA2C3A7167A8FB6E" box="[1416,1436,1125,1153]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90B310F6560335FFA2C3A7167A0FB93" box="[1416,1428,1125,1148]" italics="true" pageId="19" pageNumber="20">x</emphasis>
|
||
1
|
||
</date>
|
||
inserted dorsally in shallow depression, hair straight and directed backwards, length 44 µm; trichobothrium
|
||
<emphasis id="B90B310F6560335FFA303A9867A8FB4C" box="[1428,1436,1164,1187]" italics="true" pageId="19" pageNumber="20">t</emphasis>
|
||
distinctly crooked (
|
||
<figureCitation id="1344F1986560335FFED33ABA63FAFB28" box="[375,462,1198,1223]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 83</figureCitation>
|
||
); hairs of other trichobothria (
|
||
<emphasis id="B90B310F6560335FFC963AA46178FB28" box="[818,844,1200,1223]" italics="true" pageId="19" pageNumber="20">ist</emphasis>
|
||
and
|
||
<emphasis id="B90B310F6560335FFC2F3AA461AFFB28" box="[907,923,1200,1223]" italics="true" pageId="19" pageNumber="20">it</emphasis>
|
||
) lost or broken. Fixed finger with 21 teeth: apodens plus 14 marginal (irregular distally), 5 paraxial and basal tuberculate tooth. Movable finger with 21 teeth: apodens plus 15 marginal, 4 paraxial and basal tuberculate tooth. Tuberculate tooth of fixed finger bearing seta at base. Fixed finger with 3 lyrifissures: one on each side at base and 1 dorsal, just in front of trichobothrium
|
||
<emphasis id="B90B310F6560335FFA213B0867A0FADC" box="[1413,1428,1308,1331]" italics="true" pageId="19" pageNumber="20">it</emphasis>
|
||
. Movable finger with 1 ventral lyrifissure, situated between
|
||
<emphasis id="B90B310F6560335FFC9D3B546175FAB8" box="[825,833,1344,1367]" italics="true" pageId="19" pageNumber="20">t</emphasis>
|
||
and pit; a single spot sensillum situated immediately behind lyrifissure (
|
||
<figureCitation id="1344F1986560335FFECE3B76638AFA94" box="[362,446,1378,1403]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 80</figureCitation>
|
||
). Movable finger with 5 clavate chemosensory setae in a pit that is shallower than that of adult (
|
||
<figureCitation id="1344F1986560335FFF7A3B926355FA70" box="[222,353,1414,1439]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Figs 79‒80</figureCitation>
|
||
). Proximal sensillum
|
||
<emphasis id="B90B310F6560335FFDF03B9C6056FA70" box="[596,610,1416,1439]" italics="true" pageId="19" pageNumber="20">p</emphasis>
|
||
1 (
|
||
<figureCitation id="1344F1986560335FFDDE3B9260FFFA70" box="[634,715,1414,1439]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 84</figureCitation>
|
||
) situated about a third from base of movable finger (
|
||
<figureCitation id="1344F1986560335FFABA3B926745FA70" box="[1310,1393,1414,1439]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 83</figureCitation>
|
||
);
|
||
<emphasis id="B90B310F6560335FFA223B9C67A0FA70" box="[1414,1428,1416,1439]" italics="true" pageId="19" pageNumber="20">p</emphasis>
|
||
2 absent.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF633BC6608FF9DD" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
Legs (
|
||
<figureCitation id="1344F1986560335FFEAF3BC563BFFA05" box="[267,395,1489,1514]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Figs 85–87</figureCitation>
|
||
) similar to those of adult (apart from usual differences), except that the trochanter of leg IV is relatively smaller and all femora lack the dorsal lyrifissure and its depression; femur of leg IV with large dorsoparaxial lyrifissure at base (
|
||
<figureCitation id="1344F1986560335FFE69380D602AF9DD" box="[461,542,1561,1586]" captionStart="FIGURES 74 – 87" captionStartId="18.[151,264,1722,1745]" captionTargetBox="[211,1365,216,1664]" captionTargetId="figure@18.[207,1368,205,1675]" captionTargetPageId="18" captionText="FIGURES 74 – 87. Cybella bedosae n. sp., protonymph, appendages. 74, right chelicera, antiaxial view; 75, rallum of right chelicera, paraxial view; 76, chela of right palp, dorsal view; 77, right palp, minus chela, dorsal view; 78, tip of fixed finger, dorsal view, setae omitted; 79, chemosensory setae of movable finger of right chela, ventral view; 80, chemosensory setae of movable finger of right chela, antiaxial view; 81, tip of fixed finger of right chela, paraxial view, setae omitted; 82, tip of movable finger of right chela, paraxial view, setae omitted; 83, right chela, antiaxial view; 84, proximal sensillum (p 1) of movable chelal finger, antiaxial view; 85, femur of right leg I, paraxial view, showing paraxial lyrifissure; 86, right leg I, antiaxial view (scaly ornamentation only partly indicated on femur, omitted on other segments); 87, right leg IV, antiaxial view. Abbreviations: cs, chemosensory seta; ly, lyrifissure. Other abbreviations are notations of cheliceral setae and chelal trichobothria. Scale lines 0.1 mm (Figs 76 ‒ 77, 83, 86 ‒ 87), 0.05 mm (Figs 74 ‒ 75, 78 ‒ 82, 85; Figs 75 and 78 ‒ 82 share same scale line), 0.01 mm (Fig. 84)." httpUri="https://zenodo.org/record/569128/files/figure.png" pageId="19" pageNumber="20">Fig. 85</figureCitation>
|
||
) (as in adult).
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D6560335FFF63382A6604F8E5" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">Measurements (in mm, standard ratios in parentheses). Body 0.85×0.54 (1.6). Carapace 0.309×0.315 (0.98). Chelicera 0.129×0.082 (1.6), palm 0.092 (1.1), movable finger 0.062 (0.7×palm), spinneret 0.030 (0.5×movable finger). Palp femur 0.249×0.168 (1.5), patella 0.197×0.101 (1.9), chela 0.364×0.088 (4.1), palm 0.090 (1.0), movable finger 0.278 (3.1). Leg I trochanter 0.095×0.064 (1.5), femur 0.118×0.045 (2.6), patella 0.093×0.052 (1.8), tibia 0.093×0.048 (2.0), tarsus 0.173×0.041 (4.2). Leg IV trochanter 0.128×0.069 (1.8), femur 0.095×0.051 (1.9), patella 0.136×0.057 (2.4), tibia 0.166×0.045 (3.7), tarsus 0.230×0.042 (4.5).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365BE9665603358FF633901661FFF3A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="8BC0ED1D6560335FFF63390161EDF875" blockId="19.[151,1437,151,2018]" pageId="19" pageNumber="20">
|
||
<emphasis id="B90B310F6560335FFF633901630FF8C1" bold="true" box="[199,315,1813,1838]" pageId="19" pageNumber="20">Remarks.</emphasis>
|
||
An attempt by Anne Bedos and the author to collect more material of
|
||
<taxonomicName id="4C7F969E6560335FFBFE390266E3F8C1" box="[1114,1239,1813,1838]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6560335FFBFE390266E3F8C1" box="[1114,1239,1813,1838]" italics="true" pageId="19" pageNumber="20">C. bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746560335FFB7A3901672AF8C1" box="[1246,1310,1813,1838]" pageId="19" pageNumber="20" rank="species">
|
||
<emphasis id="B90B310F6560335FFB7A3901672AF8C1" bold="true" box="[1246,1310,1813,1838]" pageId="19" pageNumber="20">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
at the
|
||
<typeStatus id="54C453BF6560335FFACF390267A8F8C1" box="[1387,1436,1814,1838]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality in
|
||
<date id="FFC1CBDD6560335FFEB6392E63B2F8BD" box="[274,390,1850,1874]" pageId="19" pageNumber="20" value="2008-06">June 2008</date>
|
||
was thwarted by dense brush outside the cave, which had grown since the previous visit in 2005, due to the cessation of grazing activity at the site. Berlese extractions of litter and soil samples taken just outside the cave in 2005 and nearby in 2008 did not produce any feaellids.
|
||
</paragraph>
|
||
<paragraph id="8BC0ED1D65603358FF6339B1661FFF3A" blockId="19.[151,1437,151,2018]" lastBlockId="20.[151,1436,151,213]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
Although
|
||
<taxonomicName id="4C7F969E6560335FFE9C39B26382F851" box="[312,438,1957,1982]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
|
||
<emphasis id="B90B310F6560335FFE9C39B26382F851" box="[312,438,1957,1982]" italics="true" pageId="19" pageNumber="20">C. bedosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746560335FFE1939B163C9F851" box="[445,509,1957,1982]" pageId="19" pageNumber="20" rank="species">
|
||
<emphasis id="B90B310F6560335FFE1939B163C9F851" bold="true" box="[445,509,1957,1982]" pageId="19" pageNumber="20">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F969E6560335FFD9139B260E0F851" box="[565,724,1957,1982]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
|
||
<emphasis id="B90B310F6560335FFD9139B260E0F851" box="[565,724,1957,1982]" italics="true" pageId="19" pageNumber="20">C. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2388C746560335FFD7939B16129F851" box="[733,797,1957,1982]" pageId="19" pageNumber="20" rank="species">
|
||
<emphasis id="B90B310F6560335FFD7939B16129F851" bold="true" box="[733,797,1957,1982]" pageId="19" pageNumber="20">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are very similar and known from only a few specimens, they appear to be isolated geographically, occurring in disjunct limestone hills about
|
||
<quantity id="4C8740F86560335FFBE239DD66B9F80D" box="[1094,1165,1993,2018]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="km" value="60.0">60 km</quantity>
|
||
apart.
|
||
<taxonomicName id="4C7F969E6560335FFB7C39DD677FF80D" box="[1240,1355,1993,2018]" class="Arachnida" family="Feaellidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Feaellidae</taxonomicName>
|
||
appear to have very poor powers of dispersal and show high levels of endemism, so it seems reasonable to infer that two closely related species are involved, rather than a single variable or clinal species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |