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<document id="3675386A1E9967E70A03CB07F874C864" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE4FB622DD9E45CAFD17059A" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius fukiensis Balthasar S. Tarasov 2017, new combination" docType="treatment" docVersion="12" lastPageNumber="119" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="119" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="5B4F1F6430731F030AA6FD3EF33DB9B3">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<treatment id="03C187E1AE4FB622DD9E45CAFD17059A" ID-DOI="http://doi.org/10.5281/zenodo.6010814" ID-GBIF-Taxon="135106130" ID-Zenodo-Dep="6010814" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A" httpUri="http://treatment.plazi.org/id/03C187E1AE4FB622DD9E45CAFD17059A" lastPageNumber="119" pageId="18" pageNumber="119">
<subSubSection id="C372657CAE4FB622DD9E45CAFEE900B0" pageId="18" pageNumber="119" type="nomenclature">
<paragraph id="8BD736F7AE4FB622DD9E45CAFC8000EF" blockId="18.[151,859,645,704]" box="[151,859,645,671]" pageId="18" pageNumber="119">
<heading id="D09F819BAE4FB622DD9E45CAFC8000EF" bold="true" box="[151,859,645,671]" fontSize="11" level="1" pageId="18" pageNumber="119" reason="1">
<emphasis id="B91CEAE5AE4FB622DD9E45CAFC8000EF" bold="true" box="[151,859,645,671]" pageId="18" pageNumber="119">
<taxonomicName id="4C684D74AE4FB622DD9E45CAFDA700EF" ID-CoL="75NLS" authority="(Balthasar, 1960)" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[151,636,645,671]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis" status="comb. nov.">
<emphasis id="B91CEAE5AE4FB622DD9E45CAFE7A00EF" bold="true" box="[151,417,645,671]" italics="true" pageId="18" pageNumber="119">Parachorius fukiensis</emphasis>
(
<bibRefCitation id="EFF94B06AE4FB622DCB945CAFDAF00EF" author="Balthasar" box="[432,628,645,671]" pageId="18" pageNumber="119" refString="Balthasar, V. (1960) Einige neue Arten der Familie Scarabaeidae. Entomologische Blatter, 56, 88 - 94." type="journal article" year="1960">Balthasar, 1960</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE4FB622DF8D45CAFC8000EF" box="[644,859,645,671]" pageId="18" pageNumber="119">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE4FB622DD9E45E8FEE900B0" blockId="18.[151,859,645,704]" box="[151,306,679,704]" pageId="18" pageNumber="119">
(
<figureCitation id="13532A72AE4FB622DD9645E8FF3600B0" box="[159,237,679,704]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="18" pageNumber="119">Figs. 8</figureCitation>
,
<figureCitation id="13532A72AE4FB622DDF345E8FEF100B0" box="[250,298,679,704]" captionStart="FIGURE 25" captionStartId="42.[151,250,1911,1933]" captionTargetBox="[151,1433,273,1824]" captionTargetId="figure@42.[151,1436,273,1824]" captionTargetPageId="42" captionText="FIGURE 25. Distribution maps of Parachorius species. (A) Parachorius asymmetricus. (B) Parachorius bolavensis. (C) Parachorius fukiensis. (D) Parachorius fungorum. (E) Parachorius globosus. (F) Parachorius gotoi. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001967/files/figure.png" pageId="18" pageNumber="119">25C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DD9E45BFFDF70154" pageId="18" pageNumber="119" type="reference_group">
<paragraph id="8BD736F7AE4FB622DD9E45BFFDF70154" blockId="18.[151,556,752,804]" pageId="18" pageNumber="119">
<treatmentCitationGroup id="AB7811D9AE4FB622DD9E45BFFDF70154" pageId="18" pageNumber="119">
<taxonomicName id="4C684D74AE4FB622DD9E45BFFDC70176" ID-CoL="RNHQ" authority="Balthasar, 1960: 90" authorityName="Balthasar" authorityPageNumber="90" authorityYear="1960" box="[151,540,752,774]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE4FB622DD9E45BFFE890176" box="[151,338,752,774]" italics="true" pageId="18" pageNumber="119">Cassolus fukiensis</emphasis>
<treatmentCitation id="0AC910E6AE4FB622DC5E45BFFDC70176" author="Balthasar" box="[343,540,752,774]" page="90" pageId="18" pageNumber="119" year="1960">
<bibRefCitation id="EFF94B06AE4FB622DC5E45BFFE220176" author="Balthasar" box="[343,505,752,774]" pageId="18" pageNumber="119" refString="Balthasar, V. (1960) Einige neue Arten der Familie Scarabaeidae. Entomologische Blatter, 56, 88 - 94." type="journal article" year="1960">Balthasar, 1960</bibRefCitation>
: 90
</treatmentCitation>
</taxonomicName>
<emphasis id="B91CEAE5AE4FB622DD9E4441FE810154" box="[151,346,782,804]" italics="true" pageId="18" pageNumber="119">
<taxonomicName id="4C684D74AE4FB622DD9E4441FE8F0154" ID-CoL="RNHQ" authorityName="Balthasar" authorityYear="1960" box="[151,340,782,804]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis">Cassolus fukiensis</taxonomicName>
;
</emphasis>
<treatmentCitation id="0AC910E6AE4FB622DC694441FDF70154" author="Balthasar" box="[352,556,782,804]" page="261" pageId="18" pageNumber="119" year="1963">
<bibRefCitation id="EFF94B06AE4FB622DC694441FE270154" author="Balthasar" box="[352,508,782,804]" pageId="18" pageNumber="119" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Folume 1. Tschechoslowakische Akademie der Wissenschaften, Prague, 391 pp., XXIV pls." type="book" year="1963">Balthasar 1963</bibRefCitation>
: 261
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DD9E441EFD7C011A" box="[151,679,849,874]" pageId="18" pageNumber="119" type="materials_examined">
<paragraph id="8BD736F7AE4FB622DD9E441EFD7C011A" blockId="18.[151,1437,849,2026]" box="[151,679,849,874]" pageId="18" pageNumber="119">
<materialsCitation id="3B003CAAAE4FB622DD9E441EFD7C011A" ID-GBIF-Occurrence="1658411499" box="[151,679,849,874]" country="China" location="Kuatun" pageId="18" pageNumber="119" specimenCount="1" stateProvince="Fujian Province" typeStatus="holotype">
<emphasis id="B91CEAE5AE4FB622DD9E441EFEF6011A" bold="true" box="[151,301,849,874]" pageId="18" pageNumber="119">
<typeStatus id="54D38855AE4FB622DD9E441EFF09011A" box="[151,210,849,874]" pageId="18" pageNumber="119">Type</typeStatus>
locality
</emphasis>
.
<location id="8EB7602CAE4FB622DC35441DFE4A0119" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A:8EB7602CAE4FB622DC35441DFE4A0119" box="[316,401,850,873]" country="China" name="Kuatun" pageId="18" pageNumber="119" stateProvince="Fujian Province">Kuatun</location>
,
<collectingRegion id="49ACF815AE4FB622DC95441EFD8A011A" box="[412,593,849,874]" country="China" name="Fujian" pageId="18" pageNumber="119">Fújiàn Province</collectingRegion>
,
<collectingCountry id="F37F7667AE4FB622DF55441EFD78011A" box="[604,675,849,874]" name="China" pageId="18" pageNumber="119">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DDCE443AFF0D0616" pageId="18" pageNumber="119" type="distribution">
<paragraph id="8BD736F7AE4FB622DDCE443AFB9E01A6" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE443AFEBB01FE" bold="true" box="[199,352,885,910]" pageId="18" pageNumber="119">Distribution.</emphasis>
The species is known from southeast
<collectingCountry id="F37F7667AE4FB622DE1B443AFC8E01FE" box="[786,853,885,910]" name="China" pageId="18" pageNumber="119">China</collectingCountry>
(
<collectingRegion id="49ACF815AE4FB622DE6D443AFBC601FE" box="[868,1053,885,910]" country="China" name="Fujian" pageId="18" pageNumber="119">Fújiàn Province</collectingRegion>
),
<collectingCountry id="F37F7667AE4FB622D9384439FBB101FD" box="[1073,1130,886,909]" name="Laos" pageId="18" pageNumber="119">Laos</collectingCountry>
, and
<collectingCountry id="F37F7667AE4FB622D9AE443AFAD101FE" box="[1191,1290,885,910]" name="Vietnam" pageId="18" pageNumber="119">Vietnam</collectingCountry>
. The locality in
<collectingRegion id="49ACF815AE4FB622DDBD44D6FEBD01C2" box="[180,358,921,946]" country="China" name="Fujian" pageId="18" pageNumber="119">Fújiàn province</collectingRegion>
is the northernmost one among all the species in the genus
<taxonomicName id="4C684D74AE4FB622D90144D6FB4F01C2" box="[1032,1172,921,946]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE4FB622D90144D6FB4F01C2" box="[1032,1172,921,946]" italics="true" pageId="18" pageNumber="119">Parachorius</emphasis>
</taxonomicName>
. Given the large extent of occurrence, this species is probably widespread in Indochina and southern
<collectingCountry id="F37F7667AE4FB622DEF244F2FB9901A6" box="[1019,1090,957,982]" name="China" pageId="18" pageNumber="119">China</collectingCountry>
.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE44AEFF0D0616" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE44AEFE43018A" bold="true" box="[199,408,993,1018]" pageId="18" pageNumber="119">Taxonomic notes.</emphasis>
The known localities of this species are quite distantly located from each other, which is uncommon for the members of
<taxonomicName id="4C684D74AE4FB622DF01434AFD4F066E" box="[520,660,1029,1054]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE4FB622DF01434AFD4F066E" box="[520,660,1029,1054]" italics="true" pageId="18" pageNumber="119">Parachorius</emphasis>
</taxonomicName>
. Therefore, this distributional pattern may suggest that
<taxonomicName id="4C684D74AE4FB622D8124349FA47066E" authorityName="Balthasar" authorityYear="1960" box="[1307,1436,1029,1054]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE4FB622D8124349FA47066E" box="[1307,1436,1029,1054]" italics="true" pageId="18" pageNumber="119">P. fukiensis</emphasis>
</taxonomicName>
represents a complex of similar species and requires additional material and a more detailed study to resolve the issue.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DDCE433EFEDE0686" pageId="18" pageNumber="119" type="diagnosis">
<paragraph id="8BD736F7AE4FB622DDCE433EFEDE0686" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE433EFE9A06FA" bold="true" box="[199,321,1137,1162]" pageId="18" pageNumber="119">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="4C684D74AE4FB622DFB8433DFC9E06FA" authorityName="S. Tarasov" authorityYear="2017" box="[689,837,1137,1162]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="bolavensis">
<emphasis id="B91CEAE5AE4FB622DFB8433DFC9E06FA" box="[689,837,1137,1162]" italics="true" pageId="18" pageNumber="119">P. bolavensis</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="4C684D74AE4FB622D908433DFB4E06FA" authorityName="S. Tarasov" authorityYear="2017" box="[1025,1173,1137,1162]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="bolavensis">
<emphasis id="B91CEAE5AE4FB622D908433DFB4E06FA" box="[1025,1173,1137,1162]" italics="true" pageId="18" pageNumber="119">P. bolavensis</emphasis>
</taxonomicName>
in aedeagal parameres emarginate superiorly (
<emphasis id="B91CEAE5AE4FB622DC9243D9FE6506DD" box="[411,446,1174,1197]" italics="true" pageId="18" pageNumber="119">i.e.</emphasis>
, dorsally in lateral view,
<figureCitation id="13532A72AE4FB622DFD043DAFC8806DE" box="[729,851,1173,1198]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="18" pageNumber="119">Fig. 8AC</figureCitation>
), rounded aedeagal apex, and metafemoral posterior margin not rimmed.
<taxonomicName id="4C684D74AE4FB622DC8843F6FDA306A2" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[385,632,1209,1234]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="B91CEAE5AE4FB622DC8843F6FDA306A2" box="[385,632,1209,1234]" italics="true" pageId="18" pageNumber="119">Parachorius fukiensis</emphasis>
</taxonomicName>
can be separated from the rest of
<taxonomicName id="4C684D74AE4FB622DEF643F6FB5006A2" box="[1023,1163,1209,1234]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE4FB622DEF643F6FB5006A2" box="[1023,1163,1209,1234]" italics="true" pageId="18" pageNumber="119">Parachorius</emphasis>
</taxonomicName>
species by the shape of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DDCE424EFF210562" pageId="18" pageNumber="119" type="description">
<paragraph id="8BD736F7AE4FB622DDCE424EFB1C076A" blockId="18.[151,1437,849,2026]" box="[199,1223,1281,1306]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE424EFE82076A" bold="true" box="[199,345,1281,1306]" pageId="18" pageNumber="119">Description.</emphasis>
<emphasis id="B91CEAE5AE4FB622DC56424EFE44076A" box="[351,415,1281,1306]" italics="true" pageId="18" pageNumber="119">Body.</emphasis>
Body black, length
<quantity id="4C909B12AE4FB622DF88424EFCD00769" box="[641,779,1281,1306]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.6" metricValueMin="3.5" pageId="18" pageNumber="119" unit="mm" value="4.05" valueMax="4.6" valueMin="3.5">3.54.6 mm</quantity>
. Elytra uniformly colored, mostly dark.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE426AFBAD074E" blockId="18.[151,1437,849,2026]" box="[199,1142,1317,1342]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE426AFED0074E" box="[199,267,1317,1342]" italics="true" pageId="18" pageNumber="119">Head.</emphasis>
Clypeus with two teeth, outer margin of each tooth distinctly notched basally.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE4205FB5807F6" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE4205FEE60711" box="[199,317,1354,1377]" italics="true" pageId="18" pageNumber="119">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 12 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE42DEFD1B07D9" blockId="18.[151,1437,849,2026]" box="[199,704,1425,1450]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE42DEFECE07DA" box="[199,277,1425,1450]" italics="true" pageId="18" pageNumber="119">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE42F9FD9407BE" blockId="18.[151,1437,849,2026]" box="[199,591,1461,1486]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE42F9FEC907BD" box="[199,274,1462,1485]" italics="true" pageId="18" pageNumber="119">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE4295FBD004D6" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE4295FED80781" box="[199,259,1498,1521]" italics="true" pageId="18" pageNumber="119">Legs.</emphasis>
Profemur with anterior margin produced in angle medially or with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth normal, notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, or dilated in apical third of metafemur and produced in angle. Metatibia slightly sinuate, inner margin not denticulate, produced in small angle apically.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE41FDFBBD04BA" blockId="18.[151,1437,849,2026]" box="[199,1126,1713,1738]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE41FDFEB904B9" box="[199,354,1714,1737]" italics="true" pageId="18" pageNumber="119">Metasternum.</emphasis>
Transverse basal ridge of metasternum reduced and extremely short.
</paragraph>
<paragraph id="8BD736F7AE4FB622DDCE419AFF210562" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DDCE419AFEE0049E" box="[199,315,1749,1774]" italics="true" pageId="18" pageNumber="119">Aedeagus.</emphasis>
Parameres symmetrical, emarginate superiorly (
<emphasis id="B91CEAE5AE4FB622DE7C4199FC43049D" box="[885,920,1750,1773]" italics="true" pageId="18" pageNumber="119">i.e.</emphasis>
, dorsally in lateral view,
<figureCitation id="13532A72AE4FB622D9C6419AFA95049E" box="[1231,1358,1749,1774]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="18" pageNumber="119">Fig. 8AC</figureCitation>
), apex rounded.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE4FB622DDCE4052FD17059A" pageId="18" pageNumber="119" type="materials_examined">
<paragraph id="8BD736F7AE4FB622DDCE4052FD17059A" blockId="18.[151,1437,849,2026]" pageId="18" pageNumber="119">
<materialsCitation id="3B003CAAAE4FB622DDCE4052FDB4052A" ID-GBIF-Occurrence="1658411497" collectingDate="1938-04-29" collectorName="L. Klapperich &amp; Fukien &amp; Balth" country="China" county="Material" elevation="2300" latitude="27.4" location="Br." municipality="Kuatun" pageId="18" pageNumber="119" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEAE5AE4FB622DDCE4052FDFF0546" bold="true" box="[199,548,1821,1846]" pageId="18" pageNumber="119">
<collectingCounty id="62B64E7BAE4FB622DDCE4052FEF40546" box="[199,303,1821,1846]" pageId="18" pageNumber="119">Material</collectingCounty>
examined.
<typeStatus id="54D38855AE4FB622DCBE4052FDFF0546" box="[439,548,1821,1846]" pageId="18" pageNumber="119" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F37F7667AE4FB622DF3A4052FDA70546" box="[563,636,1821,1846]" name="China" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DF3A4052FDA70546" bold="true" box="[563,636,1821,1846]" pageId="18" pageNumber="119">China</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE4FB622DF824052FD1F0546" bold="true" box="[651,708,1821,1846]" pageId="18" pageNumber="119">male</emphasis>
, &quot;
<collectingMunicipality id="6BB3AC8DAE4FB622DFD34051FCF40546" box="[730,815,1822,1846]" pageId="18" pageNumber="119">Kuatun</collectingMunicipality>
(
<elevation id="0045D1C4AE4FB622DE374051FC4B0546" box="[830,912,1822,1846]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="119" unit="m" value="2300.0">
<quantity id="4C909B12AE4FB622DE374051FC4B0546" box="[830,912,1822,1846]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="119" unit="m" value="2300.0">2300m</quantity>
</elevation>
)
<geoCoordinate id="EE5C5030AE4FB622DE974051FC350546" box="[926,1006,1822,1846]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="555" value="27.4">27.40n</geoCoordinate>
.
<location id="8EB7602CAE4FB622DEF04051FBC00545" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A:8EB7602CAE4FB622DEF04051FBC00545" box="[1017,1051,1822,1845]" country="China" county="Material" latitude="27.4" municipality="Kuatun" name="Br." pageId="18" pageNumber="119">Br.</location>
117.40ö.
<collectorName id="269D5321AE4FB622D9804051FAFF0546" box="[1161,1316,1821,1846]" pageId="18" pageNumber="119">L. Klapperich</collectorName>
<collectingDate id="EF92E9DFAE4FB622D8254051FA470546" box="[1324,1436,1822,1846]" pageId="18" pageNumber="119" value="1938-04-29">
<date id="FFD61037AE4FB622D8254051FA470546" box="[1324,1436,1822,1846]" pageId="18" pageNumber="119" value="1938-04-29">29.4.1938</date>
</collectingDate>
(
<collectorName id="269D5321AE4FB622DD97400EFF34052A" box="[158,239,1857,1882]" pageId="18" pageNumber="119">Fukien</collectorName>
) /
<taxonomicName id="4C684D74AE4FB622DC1A400EFE3D052A" authority="Tarasov, 2017" authorityName="Tarasov" authorityYear="2017" box="[275,486,1857,1882]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis" status="sp. nov.">Cassolus fukiensis</taxonomicName>
<taxonomicNameLabel id="A22F579EAE4FB622DCF9400DFDFE0529" box="[496,549,1858,1881]" pageId="18" pageNumber="119" rank="species">n.sp.</taxonomicNameLabel>
<collectorName id="269D5321AE4FB622DF26400EFDB4052A" box="[559,623,1857,1882]" pageId="18" pageNumber="119">Balth</collectorName>
</materialsCitation>
.
<materialsCitation id="3B003CAAAE4FB622DF74400EFDE3050E" ID-GBIF-Occurrence="1658411513" collectionCode="NMPC" country="China" latitude="27.666666" location="Holotpus" longLatPrecision="1235" longitude="117.666664" pageId="18" pageNumber="119" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D38855AE4FB622DF74400EFD29052A" box="[637,754,1857,1882]" pageId="18" pageNumber="119" type="holotype">Holotypus</typeStatus>
/
<location id="8EB7602CAE4FB622DE04400EFCAF052A" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A:8EB7602CAE4FB622DE04400EFCAF052A" box="[781,884,1857,1882]" country="China" latitude="27.666666" longLatPrecision="1235" longitude="117.666664" name="Holotpus" pageId="18" pageNumber="119">Holotpus</location>
<taxonomicName id="4C684D74AE4FB622DE77400EFAB30529" authority="Balthasar S. Tarasov" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[894,1384,1857,1882]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="fukiensis">Parachorius fukiensis Balthasar S. Tarasov</taxonomicName>
det. 2011&quot;,
<geoCoordinate id="EE5C5030AE4FB622DDE74029FE97050E" box="[238,332,1894,1918]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="925" value="27.666666">27°40'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE4FB622DC564029FE1E050E" box="[351,453,1894,1918]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="925" value="117.666664">117°40'E</geoCoordinate>
(
<collectionCode id="ED79AE32AE4FB622DCD34029FDEB050E" box="[474,560,1894,1918]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="18" pageNumber="119">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE4FB622DF40402AFB1D05D2" ID-GBIF-Occurrence="1658411504" collectingDate="2008-08-03" collectingDateMax="2008-08-16" collectingDateMin="2008-08-03" collectorName="S. Tarasov" country="Laos" elevation="1100" latitude="19.288834" location="Plain of Jars" longLatPrecision="12" longitude="103.15283" pageId="18" pageNumber="119" specimenCount="1" specimenCount-male="1" stateProvince="Xieng Khouang" typeStatus="holotype">
<collectingCountry id="F37F7667AE4FB622DF40402AFD58050E" box="[585,643,1893,1918]" name="Laos" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622DF40402AFD58050E" bold="true" box="[585,643,1893,1918]" pageId="18" pageNumber="119">Laos</emphasis>
</collectingCountry>
:
<specimenCount id="9D6EFD7EAE4FB622DF91402AFD35050E" box="[664,750,1893,1918]" pageId="18" pageNumber="119" type="male">
<emphasis id="B91CEAE5AE4FB622DF91402AFD35050E" bold="true" box="[664,750,1893,1918]" pageId="18" pageNumber="119">1 male</emphasis>
</specimenCount>
,
<collectingRegion id="49ACF815AE4FB622DE0B402AFC60050E" box="[770,955,1893,1918]" country="Laos" name="Xiangkhouang" pageId="18" pageNumber="119">Xieng Khouang</collectingRegion>
Province, Phonsavan,
<location id="8EB7602CAE4FB622D9C7402AFABD050D" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A:8EB7602CAE4FB622D9C7402AFABD050D" box="[1230,1382,1893,1918]" country="Laos" latitude="19.288834" longLatPrecision="12" longitude="103.15283" name="Plain of Jars" pageId="18" pageNumber="119" stateProvince="Xieng Khouang">Plain of Jars</location>
N3,
<geoCoordinate id="EE5C5030AE4FB622DD9E40C5FEC205D2" box="[151,281,1930,1954]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="9" value="19.288834">19°17.33'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE4FB622DC2C40C5FE7F05D2" box="[293,420,1930,1954]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="9" value="103.15283">103°9.17'E</geoCoordinate>
,
<elevation id="0045D1C4AE4FB622DCA640C5FDDD05D1" box="[431,518,1930,1954]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="119" unit="m" value="1100.0">
<quantity id="4C909B12AE4FB622DCA640C5FDDD05D1" box="[431,518,1930,1954]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="119" unit="m" value="1100.0">1100 m</quantity>
</elevation>
, disturb montane forest, on
<collectingDate id="EF92E9DFAE4FB622DE4940C6FC3E05D2" box="[832,997,1929,1954]" pageId="18" pageNumber="119" value="2008-08-03" valueMax="2008-08-16" valueMin="2008-08-03">
<date id="FFD61037AE4FB622DE4940C6FC3E05D2" box="[832,997,1929,1954]" pageId="18" pageNumber="119" value="2008-08-03" valueMax="2008-08-16" valueMin="2008-08-03">316/viii/2008</date>
</collectingDate>
(
<collectorName id="269D5321AE4FB622DEFE40C5FBB505D2" box="[1015,1134,1930,1954]" pageId="18" pageNumber="119">S. Tarasov</collectorName>
) (CST)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE4FB622D9D840C6FBC405B6" ID-GBIF-Occurrence="1658411512" collectingDate="1982-06-12" collectionCode="ZIN" collectorName="L. Medvedev" country="Vietnam" latitude="14.333333" location="Vietnam" longLatPrecision="1288" longitude="108.6" pageId="18" pageNumber="119" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F37F7667AE4FB622D9D840C6FAEE05D2" box="[1233,1333,1929,1954]" name="Vietnam" pageId="18" pageNumber="119">
<emphasis id="B91CEAE5AE4FB622D9D840C6FAEE05D2" bold="true" box="[1233,1333,1929,1954]" pageId="18" pageNumber="119">Vietnam</emphasis>
</collectingCountry>
:
<specimenCount id="9D6EFD7EAE4FB622D84F40C6FA4F05D2" box="[1350,1428,1929,1954]" pageId="18" pageNumber="119" type="male">
<emphasis id="B91CEAE5AE4FB622D84F40C6FA4F05D2" bold="true" box="[1350,1428,1929,1954]" pageId="18" pageNumber="119">1 male</emphasis>
</specimenCount>
, Gialai Contum, Buonluoi,
<geoCoordinate id="EE5C5030AE4FB622DCCC40E1FDFF05B6" box="[453,548,1966,1990]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="925" value="14.333333">14°20'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE4FB622DF3940E2FD4105B6" box="[560,666,1965,1990]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="925" value="108.6">108°36'E</geoCoordinate>
,
<collectingDate id="EF92E9DFAE4FB622DFAE40E2FCFB05B6" box="[679,800,1965,1990]" pageId="18" pageNumber="119" value="1982-06-12">
<date id="FFD61037AE4FB622DFAE40E2FCFB05B6" box="[679,800,1965,1990]" pageId="18" pageNumber="119" value="1982-06-12">12/vi/1982</date>
</collectingDate>
(
<collectorName id="269D5321AE4FB622DE3B40E1FC1105B6" box="[818,970,1965,1990]" pageId="18" pageNumber="119">L. Medvedev</collectorName>
) (
<collectionCode id="ED79AE32AE4FB622DEEA40E1FBCC05B6" box="[995,1047,1966,1990]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="119">ZIN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE4FB622D92240E2FD13059A" ID-GBIF-Occurrence="1658411502" collectingDate="1963-03-27" collectionCode="ZIN" collectorName="O. Kabakov" country="Vietnam" elevation="300" latitude="17.466667" location="Donghoi" longLatPrecision="1278" longitude="106.583336" pageId="18" pageNumber="119" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="9D6EFD7EAE4FB622D92240E2FB5E05B6" box="[1067,1157,1965,1990]" pageId="18" pageNumber="119" type="male">
<emphasis id="B91CEAE5AE4FB622D92240E2FB5E05B6" bold="true" box="[1067,1157,1965,1990]" pageId="18" pageNumber="119">2 males</emphasis>
</specimenCount>
, SW
<location id="8EB7602CAE4FB622D9CE40E2FAF705B6" LSID="urn:lsid:plazi:treatment:03C187E1AE4FB622DD9E45CAFD17059A:8EB7602CAE4FB622D9CE40E2FAF705B6" box="[1223,1324,1965,1990]" country="Vietnam" latitude="17.466667" longLatPrecision="1278" longitude="106.583336" name="Donghoi" pageId="18" pageNumber="119">Donghoi</location>
,
<geoCoordinate id="EE5C5030AE4FB622D83140E1FA4C05B6" box="[1336,1431,1966,1990]" direction="north" orientation="latitude" pageId="18" pageNumber="119" precision="925" value="17.466667">17°28'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE4FB622DD9E409EFEDB059A" box="[151,256,2001,2026]" direction="east" orientation="longitude" pageId="18" pageNumber="119" precision="925" value="106.583336">106°35'E</geoCoordinate>
,
<elevation id="0045D1C4AE4FB622DC05409DFE8E0599" box="[268,341,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="119" unit="m" value="300.0">
<quantity id="4C909B12AE4FB622DC05409DFE8E0599" box="[268,341,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="119" unit="m" value="300.0">300 m</quantity>
</elevation>
,
<collectingDate id="EF92E9DFAE4FB622DC68409EFE07059A" box="[353,476,2001,2026]" pageId="18" pageNumber="119" value="1963-03-27">
<date id="FFD61037AE4FB622DC68409EFE07059A" box="[353,476,2001,2026]" pageId="18" pageNumber="119" value="1963-03-27">27/iii/1963</date>
</collectingDate>
(
<collectorName id="269D5321AE4FB622DCE5409DFDAE059A" box="[492,629,2001,2026]" pageId="18" pageNumber="119">O. Kabakov</collectorName>
) (
<collectionCode id="ED79AE32AE4FB622DF85409DFD1B059A" box="[652,704,2002,2026]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="119">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>