369 lines
39 KiB
XML
369 lines
39 KiB
XML
<document id="537CBD81EB280E013B86E55F7EB3EFB4" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE43B610DD9E44E7FB1F03A0" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius newthayerae Tarasov 2017, new species" docType="treatment" docUuid="5DE8A2B0-44BC-43A0-B38F-B3483509F42F" docUuidSource="ZooBank" docVersion="12" lastPageNumber="132" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="131" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="2ABEFE03C4798A14922003B5A0EB52C0">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="93B0121A89A7EDC628B650A5A3C159CA">Tarasov, Sergei</mods:namePart>
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<mods:title id="DC01010E10E1DF0AF99ACE63BCD489E9">Zootaxa</mods:title>
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<mods:part id="F8F16AE626063CCF0B54E39FD7BA9BF3">
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<mods:date id="8434E8B49B51DA7C0F92248DFC311AA2">2017</mods:date>
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<mods:number id="0A2473F8DA8F74782C5414E296CF5A2A">2017-10-04</mods:number>
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<mods:number id="6FFDC740CE05E6CB66C8F02C77F1090E">4329</mods:number>
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<treatment id="03C187E1AE43B610DD9E44E7FB1F03A0" ID-DOI="http://doi.org/10.5281/zenodo.6010834" ID-GBIF-Taxon="135106138" ID-Zenodo-Dep="6010834" LSID="urn:lsid:zoobank.org:act:5DE8A2B0-44BC-43A0-B38F-B3483509F42F" httpUri="http://treatment.plazi.org/id/03C187E1AE43B610DD9E44E7FB1F03A0" lastPageId="32" lastPageNumber="132" pageId="30" pageNumber="131">
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<subSubSection id="C372657CAE43B62EDD9E44E7FEE60194" pageId="30" pageNumber="131" type="nomenclature">
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<paragraph id="8BD736F7AE43B62EDD9E44E7FD1401B2" blockId="30.[151,965,936,1032]" box="[151,719,936,962]" pageId="30" pageNumber="131">
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<heading id="D09F819BAE43B62EDD9E44E7FD1401B2" bold="true" box="[151,719,936,962]" fontSize="11" level="1" pageId="30" pageNumber="131" reason="1">
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<emphasis id="B91CEAE5AE43B62EDD9E44E7FD1401B2" bold="true" box="[151,719,936,962]" pageId="30" pageNumber="131">
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<taxonomicName id="4C684D74AE43B62EDD9E44E7FDE301B2" ID-CoL="6TNLW" authority="Tarasov" authorityName="Tarasov" authorityYear="2017" box="[151,568,936,962]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae" status="sp. nov.">
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<emphasis id="B91CEAE5AE43B62EDD9E44E7FE1601B2" bold="true" box="[151,461,936,962]" italics="true" pageId="30" pageNumber="131">Parachorius newthayerae</emphasis>
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Tarasov
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</taxonomicName>
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<taxonomicNameLabel id="A22F579EAE43B62EDF3644E7FD1401B2" box="[575,719,936,962]" pageId="30" pageNumber="131" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD736F7AE43B62EDD9E4484FEE60194" blockId="30.[151,965,936,1032]" box="[151,317,971,996]" pageId="30" pageNumber="131">
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(
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<figureCitation id="13532A72AE43B62EDD964484FF270194" box="[159,252,971,996]" captionStart="FIGURE 17" captionStartId="31.[151,250,1575,1597]" captionTargetBox="[151,1416,301,1538]" captionTargetId="figure@31.[151,1436,301,1553]" captionTargetPageId="31" captionText="FIGURE 17. Morphology of Parachorius newthayerae. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype)." httpUri="https://zenodo.org/record/1001951/files/figure.png" pageId="30" pageNumber="131">Figs. 17</figureCitation>
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,
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<figureCitation id="13532A72AE43B62EDC014484FEEE0194" box="[264,309,971,996]" captionStart="FIGURE 26" captionStartId="43.[151,250,1905,1927]" captionTargetBox="[151,1436,265,1825]" captionTargetId="figure@43.[151,1436,265,1825]" captionTargetPageId="43" captionText="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001969/files/figure.png" pageId="30" pageNumber="131">26F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62EDD9E44A0FC1E0678" box="[151,965,1007,1032]" pageId="30" pageNumber="131" type="description">
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<paragraph id="8BD736F7AE43B62EDD9E44A0FC1E0678" blockId="30.[151,965,936,1032]" box="[151,965,1007,1032]" pageId="30" pageNumber="131">urn:lsid:zoobank.org:act:5DE8A2B0-44BC-43A0-B38F-B3483509F42F</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62EDD9E4378FDE6063F" box="[151,573,1079,1104]" pageId="30" pageNumber="131" type="materials_examined">
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<paragraph id="8BD736F7AE43B62EDD9E4378FDE6063F" blockId="30.[151,1436,1079,2004]" box="[151,573,1079,1104]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDD9E4378FEF60620" bold="true" box="[151,301,1079,1104]" pageId="30" pageNumber="131">Type locality</emphasis>
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. Bolaven plateau, Laos.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62EDDCE4314FEC80774" pageId="30" pageNumber="131" type="etymology">
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<paragraph id="8BD736F7AE43B62EDDCE4314FEC80774" blockId="30.[151,1436,1079,2004]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE4314FE950604" bold="true" box="[199,334,1115,1140]" pageId="30" pageNumber="131">Etymology.</emphasis>
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The species is named after two great entomologists, Alfred Newton and Margaret Thayer, who collected the largest series of specimens of this new species in
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<collectingCountry id="F37F7667AE43B62EDE5143CFFC4906E7" box="[856,914,1152,1175]" name="Laos" pageId="30" pageNumber="131">Laos</collectingCountry>
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. Alfred and Margaret kindly hosted me during a visit to the Field Museum where the largest part of this taxonomic work on
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<taxonomicName id="4C684D74AE43B62ED92843ECFB7606CC" box="[1057,1197,1187,1212]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE43B62ED92843ECFB7606CC" box="[1057,1197,1187,1212]" italics="true" pageId="30" pageNumber="131">Parachorius</emphasis>
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</taxonomicName>
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was conducted. The epithet “
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<taxonomicName id="4C684D74AE43B62EDDF04388FE530690" authorityName="Tarasov" authorityYear="2017" box="[249,392,1223,1248]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae">
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<emphasis id="B91CEAE5AE43B62EDDF04388FE530690" box="[249,392,1223,1248]" italics="true" pageId="30" pageNumber="131">newthayerae</emphasis>
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</taxonomicName>
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” is the combination of their last names. This species name should be treated as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62EDDCE4240FD5D07C8" pageId="30" pageNumber="131" type="distribution">
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<paragraph id="8BD736F7AE43B62EDDCE4240FB9F0757" blockId="30.[151,1436,1079,2004]" box="[199,1092,1295,1320]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE4240FEBB0758" bold="true" box="[199,352,1295,1320]" pageId="30" pageNumber="131">Distribution.</emphasis>
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This species seems endemic to Bolaven plateau in southern
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<collectingCountry id="F37F7667AE43B62ED90F425FFB9B0757" box="[1030,1088,1296,1319]" name="Laos" pageId="30" pageNumber="131">Laos</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BD736F7AE43B62EDDCE427CFD5D07C8" blockId="30.[151,1436,1079,2004]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE427CFE43073C" bold="true" box="[199,408,1331,1356]" pageId="30" pageNumber="131">Taxonomic notes.</emphasis>
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In females of
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<taxonomicName id="4C684D74AE43B62EDF36427BFD30073C" authorityName="Tarasov" authorityYear="2017" box="[575,747,1331,1356]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae">
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<emphasis id="B91CEAE5AE43B62EDF36427BFD30073C" box="[575,747,1331,1356]" italics="true" pageId="30" pageNumber="131">P. newthayerae</emphasis>
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</taxonomicName>
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the secondary sexual characters of legs modification are not expressed, although anterior angles of pronotum can be very slightly angulate. As only the secondary sexual characters reliably separate
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<taxonomicName id="4C684D74AE43B62EDCD94233FDA007E4" authorityName="Tarasov" authorityYear="2017" box="[464,635,1403,1428]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae">
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<emphasis id="B91CEAE5AE43B62EDCD94233FDA007E4" box="[464,635,1403,1428]" italics="true" pageId="30" pageNumber="131">P. newthayerae</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C684D74AE43B62EDFC84233FC8707E3" box="[705,860,1404,1427]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="maruyamai">
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<emphasis id="B91CEAE5AE43B62EDFC84233FC8707E3" box="[705,860,1404,1427]" italics="true" pageId="30" pageNumber="131">P. maruyamai</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C684D74AE43B62EDE624233FC3607E4" box="[875,1005,1404,1428]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="fingorum">
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<emphasis id="B91CEAE5AE43B62EDE624233FC3607E4" box="[875,1005,1404,1428]" italics="true" pageId="30" pageNumber="131">P. fingorum</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C684D74AE43B62ED9254233FB7407E4" authorityName="Harold" authorityYear="1873" box="[1068,1199,1403,1428]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="thomsoni">
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<emphasis id="B91CEAE5AE43B62ED9254233FB7407E4" box="[1068,1199,1403,1428]" italics="true" pageId="30" pageNumber="131">P. thomsoni</emphasis>
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</taxonomicName>
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, the identification of
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<taxonomicName id="4C684D74AE43B62EDD9E42EFFE9A07C8" authorityName="Tarasov" authorityYear="2017" box="[151,321,1439,1464]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae">
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<emphasis id="B91CEAE5AE43B62EDD9E42EFFE9A07C8" box="[151,321,1439,1464]" italics="true" pageId="30" pageNumber="131">P. newthayerae</emphasis>
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</taxonomicName>
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using females is ambiguous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62EDDCE428CFABD0453" pageId="30" pageNumber="131" type="diagnosis">
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<paragraph id="8BD736F7AE43B62EDDCE428CFABD0453" blockId="30.[151,1436,1079,2004]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE428CFE9A07AC" bold="true" box="[199,321,1475,1500]" pageId="30" pageNumber="131">Diagnosis.</emphasis>
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This species is most closely related to
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<emphasis id="B91CEAE5AE43B62EDFF8428BFC4807AB" box="[753,915,1476,1499]" italics="true" pageId="30" pageNumber="131">
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<taxonomicName id="4C684D74AE43B62EDFF8428BFC5407AB" box="[753,911,1476,1499]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="maruyamai">P. maruyamai</taxonomicName>
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.
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</emphasis>
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Both species differ from all other
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<taxonomicName id="4C684D74AE43B62ED819428CFA4707AC" box="[1296,1436,1475,1500]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE43B62ED819428CFA4707AC" box="[1296,1436,1475,1500]" italics="true" pageId="30" pageNumber="131">Parachorius</emphasis>
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</taxonomicName>
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by sharped frontal angles of pronotum along with strongly curved metatibia in males.
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<taxonomicName id="4C684D74AE43B62ED97E42A8FA400470" authorityName="Tarasov" authorityYear="2017" box="[1143,1435,1511,1536]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="newthayerae">
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<emphasis id="B91CEAE5AE43B62ED97E42A8FA400470" box="[1143,1435,1511,1536]" italics="true" pageId="30" pageNumber="131">Parachorius newthayerae</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C684D74AE43B62EDC2D4143FE650453" box="[292,446,1548,1571]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="131" phylum="Arthropoda" rank="species" species="maruyamai">
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<emphasis id="B91CEAE5AE43B62EDC2D4143FE650453" box="[292,446,1548,1571]" italics="true" pageId="30" pageNumber="131">P. maruyamai</emphasis>
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</taxonomicName>
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by dilated posterior margin of metafemur and angulate lateral margin of pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE43B62FDDCE4160FB7C02A5" lastPageId="31" lastPageNumber="132" pageId="30" pageNumber="131" type="description">
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<paragraph id="8BD736F7AE43B62EDDCE4160FBEF0438" blockId="30.[151,1436,1079,2004]" box="[199,1076,1583,1608]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE4160FE820438" bold="true" box="[199,345,1583,1608]" pageId="30" pageNumber="131">Description.</emphasis>
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<emphasis id="B91CEAE5AE43B62EDC564160FE440438" box="[351,415,1583,1608]" italics="true" pageId="30" pageNumber="131">Body.</emphasis>
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Body black, length 7.8–8.0 mm. Elytra uniformly colored.
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</paragraph>
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<paragraph id="8BD736F7AE43B62EDDCE411CFB7A041C" blockId="30.[151,1436,1079,2004]" box="[199,1185,1619,1644]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE411CFED0041C" box="[199,267,1619,1644]" italics="true" pageId="30" pageNumber="131">Head.</emphasis>
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Clypeus with two indistinct teeth, outer margin of each tooth not notched basally.
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</paragraph>
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<paragraph id="8BD736F7AE43B62EDDCE4137FADA04C4" blockId="30.[151,1436,1079,2004]" pageId="30" pageNumber="131">
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<emphasis id="B91CEAE5AE43B62EDDCE4137FEE604FF" box="[199,317,1656,1679]" italics="true" pageId="30" pageNumber="131">Pronotum.</emphasis>
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Pronotum distinctly angulate laterally. Frontal angles sharp. Disc covered with punctures separated by 0.5–2.0 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
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</paragraph>
|
||
<paragraph id="8BD736F7AE43B62EDDCE41F0FD1B04A7" blockId="30.[151,1436,1079,2004]" box="[199,704,1727,1752]" pageId="30" pageNumber="131">
|
||
<emphasis id="B91CEAE5AE43B62EDDCE41F0FECE04A8" box="[199,277,1727,1752]" italics="true" pageId="30" pageNumber="131">Elytra.</emphasis>
|
||
Elytra covered with sparse punctures.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE43B62EDDCE41ABFD94048C" blockId="30.[151,1436,1079,2004]" box="[199,591,1763,1788]" pageId="30" pageNumber="131">
|
||
<emphasis id="B91CEAE5AE43B62EDDCE41ABFEC9048B" box="[199,274,1764,1787]" italics="true" pageId="30" pageNumber="131">Wings.</emphasis>
|
||
Wings normally developed.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE43B62EDDCE4047FC8805A4" blockId="30.[151,1436,1079,2004]" pageId="30" pageNumber="131">
|
||
<emphasis id="B91CEAE5AE43B62EDDCE4047FED8056F" box="[199,259,1800,1823]" italics="true" pageId="30" pageNumber="131">Legs.</emphasis>
|
||
Profemur with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth almost not modified in males, approximately as wide as second tooth; first and second teeth distinctly separated from each other; first tooth normal, distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin serrate (sometimes slightly) or not serrate, medially dilated, and produced in large rounded angle. Metatibia strongly curved and notched medially; inner margin not denticulate, produced in small angle apically.
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE42B62FDDCE47D6FB7C02A5" blockId="31.[199,1262,151,248]" pageId="31" pageNumber="132">
|
||
<emphasis id="B91CEAE5AE42B62FDDCE47D6FEB902C0" box="[199,354,153,176]" italics="true" pageId="31" pageNumber="132">Metasternum.</emphasis>
|
||
Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length.
|
||
<emphasis id="B91CEAE5AE42B62FDDCE47F3FEE002A5" box="[199,315,188,213]" italics="true" pageId="31" pageNumber="132">Aedeagus.</emphasis>
|
||
Parameres symmetrical, simple, apex widely rounded and inferiorly angulate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C372657CAE42B610DDCE4790FB1F03A0" lastPageId="32" lastPageNumber="133" pageId="31" pageNumber="132" type="materials_examined">
|
||
<paragraph id="8BD736F7AE42B62FDDCE4790FB350287" blockId="31.[199,1262,151,248]" box="[199,1262,223,248]" pageId="31" pageNumber="132">
|
||
<emphasis id="B91CEAE5AE42B62FDDCE4790FEE00288" bold="true" box="[199,315,223,248]" pageId="31" pageNumber="132">
|
||
<typeStatus id="54D38855AE42B62FDDCE4790FEED0288" box="[199,310,223,248]" pageId="31" pageNumber="132" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Profemur with sinuate anterior margin. Metafemoral posterior margin serrate.
|
||
</paragraph>
|
||
<caption id="DF17667FAE42B62FDD9E4168FAAD042B" httpUri="https://zenodo.org/record/1001951/files/figure.png" pageId="31" pageNumber="132" targetBox="[151,1416,301,1538]" targetPageId="31">
|
||
<paragraph id="8BD736F7AE42B62FDD9E4168FAAD042B" blockId="31.[151,1436,1575,1627]" pageId="31" pageNumber="132">
|
||
<emphasis id="B91CEAE5AE42B62FDD9E4168FEFA044C" bold="true" box="[151,289,1575,1597]" pageId="31" pageNumber="132">FIGURE 17.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C684D74AE42B62FDCC54168FD08044D" authorityName="Tarasov" authorityYear="2017" box="[460,723,1575,1597]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="newthayerae">
|
||
<emphasis id="B91CEAE5AE42B62FDCC54168FD08044D" box="[460,723,1575,1597]" italics="true" pageId="31" pageNumber="132">Parachorius newthayerae</emphasis>
|
||
</taxonomicName>
|
||
. (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD736F7AE42B610DDCE41CCFB1F03A0" blockId="31.[151,1437,1667,2016]" lastBlockId="32.[151,1437,151,464]" lastPageId="32" lastPageNumber="133" pageId="31" pageNumber="132">
|
||
<materialsCitation id="3B003CAAAE42B62FDDCE41CCFD3E04EC" ID-GBIF-Occurrence="1658411487" box="[199,741,1667,1692]" collectorName="Material" country="Laos" location="Laos" pageId="31" pageNumber="132" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="B91CEAE5AE42B62FDDCE41CCFDF404EC" bold="true" box="[199,559,1667,1692]" pageId="31" pageNumber="132">
|
||
<collectorName id="269D5321AE42B62FDDCE41CCFEF404EC" box="[199,303,1667,1692]" pageId="31" pageNumber="132">Material</collectorName>
|
||
examined.
|
||
<typeStatus id="54D38855AE42B62FDCCB41CCFDF404EC" box="[450,559,1667,1692]" pageId="31" pageNumber="132" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="F37F7667AE42B62FDF4A41CCFDA604EC" box="[579,637,1667,1692]" name="Laos" pageId="31" pageNumber="132">
|
||
<emphasis id="B91CEAE5AE42B62FDF4A41CCFDA604EC" bold="true" box="[579,637,1667,1692]" pageId="31" pageNumber="132">Laos</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<specimenCount id="9D6EFD7EAE42B62FDF9941CCFD3E04EC" box="[656,741,1667,1692]" pageId="31" pageNumber="132" type="male">
|
||
<emphasis id="B91CEAE5AE42B62FDF9941CCFD3E04EC" bold="true" box="[656,741,1667,1692]" pageId="31" pageNumber="132">1 male</emphasis>
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B003CAAAE42B62FDFF141CCFC110494" ID-GBIF-Occurrence="1658411480" collectingDate="2008-06-08" collectingDateMax="2008-06-16" collectingDateMin="2008-06-08" collectionCode="FMHD" collectorName="A. Newton & M. Thayer" country="Laos" elevation="1095" latitude="15.238133" location="Xekatam" longLatPrecision="1" longitude="106.53152" pageId="31" pageNumber="132" specimenCode="FMHD#2008-037" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
|
||
"
|
||
<collectingCountry id="F37F7667AE42B62FDE0841CCFC8A04EB" box="[769,849,1667,1691]" name="Laos" pageId="31" pageNumber="132">LAOS</collectingCountry>
|
||
:
|
||
<collectingRegion id="49ACF815AE42B62FDE6941CCFC3804EC" box="[864,995,1667,1692]" country="Laos" name="Champasak" pageId="31" pageNumber="132">Champasak</collectingRegion>
|
||
Pr.: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE42B62FDDA441E8FECC04B0" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE42B62FDDA441E8FECC04B0" box="[173,279,1703,1728]" country="Laos" latitude="15.238133" longLatPrecision="1" longitude="106.53152" name="Xekatam" pageId="31" pageNumber="132" stateProvince="Champasak">Xékatam</location>
|
||
) vic., old logging road N of village
|
||
<elevation id="0045D1C4AE42B62FDFA841E8FD2B04B0" box="[673,752,1703,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="31" pageNumber="132" unit="m" value="1095.0">
|
||
<quantity id="4C909B12AE42B62FDFA841E8FD2B04B0" box="[673,752,1703,1728]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="31" pageNumber="132" unit="m" value="1095.0">1095m</quantity>
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE5C5030AE42B62FDFF341E8FC5304B0" box="[762,904,1703,1728]" direction="north" orientation="latitude" pageId="31" pageNumber="132" precision="1" value="15.238133">15°14.288'N</geoCoordinate>
|
||
<geoCoordinate id="EE5C5030AE42B62FDE8741E8FBF604B0" box="[910,1069,1703,1728]" direction="east" orientation="longitude" pageId="31" pageNumber="132" precision="1" value="106.53152">106°31.891'E</geoCoordinate>
|
||
,
|
||
<collectingDate id="EF92E9DFAE42B62FD93841E8FB6604B0" box="[1073,1213,1703,1728]" pageId="31" pageNumber="132" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">
|
||
<date id="FFD61037AE42B62FD93841E8FB6604B0" box="[1073,1213,1703,1728]" pageId="31" pageNumber="132" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8-16.vi.2008</date>
|
||
</collectingDate>
|
||
/ selectively logged forest;
|
||
<specimenCode id="DBCE9E8CAE42B62FDDEE4184FE6F0493" box="[231,436,1739,1763]" collectionCode="FMHD" pageId="31" pageNumber="132">FMHD#2008-037</specimenCode>
|
||
,
|
||
<collectingMethod id="52294EE0AE42B62FDCC84184FD450493" box="[449,670,1739,1764]" pageId="31" pageNumber="132">flight intercept trap</collectingMethod>
|
||
,
|
||
<collectorName id="269D5321AE42B62FDFA24184FCF30493" box="[683,808,1739,1763]" pageId="31" pageNumber="132">A. Newton</collectorName>
|
||
&
|
||
<collectorName id="269D5321AE42B62FDE464183FC110494" box="[847,970,1739,1764]" pageId="31" pageNumber="132">M. Thayer</collectorName>
|
||
</materialsCitation>
|
||
; ANMT site 1231 ex 95% FIELD MUS. NAT. HIST. / HOLOTYPUS
|
||
<taxonomicName id="4C684D74AE42B62FDCF741A0FCC40578" authorityName="Tarasov" authorityYear="2017" box="[510,799,1775,1800]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="newthayerae">Parachorius newthayerae</taxonomicName>
|
||
/ S. Tarasov det. 2011", 15°14.288'N, 106°31.899'E (FMNH).
|
||
<emphasis id="B91CEAE5AE42B62FDC19405CFD4E055C" bold="true" box="[272,661,1811,1836]" pageId="31" pageNumber="132">Paratypes (3 males, 3 females)</emphasis>
|
||
:
|
||
<materialsCitation id="3B003CAAAE42B62FDFA5405CFBEE053F" ID-GBIF-Occurrence="1658411481" collectionCode="FMHD" country="Laos" location="Xekatam" pageId="31" pageNumber="132" specimenCode="FMHD#2008-034" specimenCount="1" specimenCount-male="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<emphasis id="B91CEAE5AE42B62FDFA5405CFCA7055C" bold="true" box="[684,892,1811,1836]" pageId="31" pageNumber="132">
|
||
<specimenCount id="9D6EFD7EAE42B62FDFA5405CFCDA055C" box="[684,769,1811,1836]" pageId="31" pageNumber="132" type="male">1 male</specimenCount>
|
||
<typeStatus id="54D38855AE42B62FDE19405CFCA7055C" box="[784,892,1811,1836]" pageId="31" pageNumber="132" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE42B62FDE98405CFB51055C" box="[913,1162,1811,1836]" country="Laos" name="Champasak" pageId="31" pageNumber="132">Champasak Province</collectingRegion>
|
||
: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE42B62FDC214078FE4A0520" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE42B62FDC214078FE4A0520" box="[296,401,1847,1872]" country="Laos" name="Xekatam" pageId="31" pageNumber="132" stateProvince="Champasak Province">Xékatam</location>
|
||
) vicinity, old logging road N of village,
|
||
<specimenCode id="DBCE9E8CAE42B62FDE6E4078FBEE053F" box="[871,1077,1847,1871]" collectionCode="FMHD" pageId="31" pageNumber="132">FMHD#2008-034</specimenCode>
|
||
</materialsCitation>
|
||
, ANMT site 1230, 15°13.96'N, 106°31.731'E,
|
||
<quantity id="4C909B12AE42B62FDC4B4014FE450503" box="[322,414,1883,1908]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="31" pageNumber="132" unit="m" value="1035.0">1035 m</quantity>
|
||
, selectively logged forest, on
|
||
<date id="FFD61037AE42B62FDE0D4014FC410504" box="[772,922,1883,1908]" pageId="31" pageNumber="132" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
|
||
(A. Newton, M. Thayer) (FMNH);
|
||
<materialsCitation id="3B003CAAAE42B62FD8414014FE6505CB" ID-GBIF-Occurrence="1658411469" collectionCode="FMHD" country="Laos" location="Xekatam" pageId="31" pageNumber="132" specimenCode="FMHD#2008-037" specimenCount="1" specimenCount-male="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<emphasis id="B91CEAE5AE42B62FD8414014FED905E8" bold="true" pageId="31" pageNumber="132">
|
||
<specimenCount id="9D6EFD7EAE42B62FD8414014FA470504" box="[1352,1436,1883,1908]" pageId="31" pageNumber="132" type="male">1 male</specimenCount>
|
||
<typeStatus id="54D38855AE42B62FDD9E4030FED905E8" box="[151,258,1919,1944]" pageId="31" pageNumber="132" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE42B62FDC1D4030FDD005E8" box="[276,523,1919,1944]" country="Laos" name="Champasak" pageId="31" pageNumber="132">Champasak Province</collectingRegion>
|
||
: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE42B62FDEBD4030FBC505E8" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE42B62FDEBD4030FBC505E8" box="[948,1054,1919,1944]" country="Laos" name="Xekatam" pageId="31" pageNumber="132" stateProvince="Champasak Province">Xékatam</location>
|
||
) vicinity, old logging road N of village,
|
||
<specimenCode id="DBCE9E8CAE42B62FDDF840ECFE6505CB" box="[241,446,1955,1979]" collectionCode="FMHD" pageId="31" pageNumber="132">FMHD#2008-037</specimenCode>
|
||
</materialsCitation>
|
||
, ANMT site 1231, 15°14.288'N, 106°31.899'E,
|
||
<quantity id="4C909B12AE42B62FDED040ECFBEA05CB" box="[985,1073,1955,1980]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="31" pageNumber="132" unit="m" value="1095.0">1095 m</quantity>
|
||
, selectively logged forest, on
|
||
<date id="FFD61037AE42B62FD88940ECFECB0590" pageId="31" pageNumber="132" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8– 16/vi/2008</date>
|
||
(A. Newton, M. Thayer) (FMNH);
|
||
<materialsCitation id="3B003CAAAE42B610DFBD4088FBF002C0" ID-GBIF-Occurrence="1658411478" collectionCode="FMHD" country="Laos" lastPageId="32" lastPageNumber="133" location="Xekatam" pageId="31" pageNumber="132" specimenCode="FMHD#2008-038" specimenCount="1" specimenCount-female="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<emphasis id="B91CEAE5AE42B62FDFBD4088FC4A0590" bold="true" box="[692,913,1991,2016]" pageId="31" pageNumber="132">
|
||
<specimenCount id="9D6EFD7EAE42B62FDFBD4088FCC00590" box="[692,795,1991,2016]" pageId="31" pageNumber="132" type="female">1 female</specimenCount>
|
||
<typeStatus id="54D38855AE42B62FDE2F4088FC4A0590" box="[806,913,1991,2016]" pageId="31" pageNumber="132" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE42B62FDEAB4088FB4C0590" box="[930,1175,1991,2016]" country="Laos" name="Champasak" pageId="31" pageNumber="132">Champasak Province</collectingRegion>
|
||
: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE7DB610DC2E47D8FE4A02C0" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE7DB610DC2E47D8FE4A02C0" box="[295,401,151,176]" country="Laos" name="Xekatam" pageId="32" pageNumber="133" stateProvince="Champasak Province">Xékatam</location>
|
||
) vicinity, old logging road N of village,
|
||
<specimenCode id="DBCE9E8CAE7DB610DE5747D7FBF002C0" box="[862,1067,152,176]" collectionCode="FMHD" pageId="32" pageNumber="133">FMHD#2008-038</specimenCode>
|
||
</materialsCitation>
|
||
, ANMT site 1231, 15°14.288'N, 106°31.891'E,
|
||
<quantity id="4C909B12AE7DB610DC4B47F3FE4502A4" box="[322,414,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.095" pageId="32" pageNumber="133" unit="m" value="1095.0">1095 m</quantity>
|
||
, selectively logged forest, on
|
||
<date id="FFD61037AE7DB610DE0F47F3FC4002A5" box="[774,923,188,213]" pageId="32" pageNumber="133" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
|
||
(A. Solodovnikov, M. Thayer, A. Newton) (FMNH);
|
||
<materialsCitation id="3B003CAAAE7DB610DC054790FD43036C" ID-GBIF-Occurrence="1658411475" collectionCode="FMHD" country="Laos" location="Xekatam" pageId="32" pageNumber="133" specimenCode="FMHD#2008-040" specimenCount="2" specimenCount-female="2" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<emphasis id="B91CEAE5AE7DB610DC054790FE280288" bold="true" box="[268,499,223,248]" pageId="32" pageNumber="133">
|
||
<specimenCount id="9D6EFD7EAE7DB610DC054790FEA80288" box="[268,371,223,248]" pageId="32" pageNumber="133" type="female">2 female</specimenCount>
|
||
<typeStatus id="54D38855AE7DB610DC744790FE280288" box="[381,499,223,248]" pageId="32" pageNumber="133" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE7DB610DF0D4790FD230288" box="[516,760,223,248]" country="Laos" name="Champasak" pageId="32" pageNumber="133">Champasak Province</collectingRegion>
|
||
: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE7DB610D9904790FAD90288" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE7DB610D9904790FAD90288" box="[1177,1282,223,248]" country="Laos" name="Xekatam" pageId="32" pageNumber="133" stateProvince="Champasak Province">Xékatam</location>
|
||
) vicinity, old logging road N of village,
|
||
<specimenCode id="DBCE9E8CAE7DB610DCC3464BFD43036C" box="[458,664,260,284]" collectionCode="FMHD" pageId="32" pageNumber="133">FMHD#2008-040</specimenCode>
|
||
</materialsCitation>
|
||
, ANMT site 1232, 15°14.494'N, 106°31.807'E,
|
||
<materialsCitation id="3B003CAAAE7DB610D9B7464BFCBA0315" ID-GBIF-Occurrence="1658411495" collectingDate="2008-06-08" collectingDateMax="2008-06-16" collectingDateMin="2008-06-08" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Laos" elevation="1170" latitude="15.245684" location="Ban Thongvay" longLatPrecision="1" longitude="106.53194" municipality="Bolaven Plateau" pageId="32" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<elevation id="0045D1C4AE7DB610D9B7464BFACC036C" box="[1214,1303,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="32" pageNumber="133" unit="m" value="1170.0">
|
||
<quantity id="4C909B12AE7DB610D9B7464BFACC036C" box="[1214,1303,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="32" pageNumber="133" unit="m" value="1170.0">1170 m</quantity>
|
||
</elevation>
|
||
, selectively logged forest, on
|
||
<collectingDate id="EF92E9DFAE7DB610DC664668FDDE0330" box="[367,517,295,320]" pageId="32" pageNumber="133" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">
|
||
<date id="FFD61037AE7DB610DC664668FDDE0330" box="[367,517,295,320]" pageId="32" pageNumber="133" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
|
||
</collectingDate>
|
||
(
|
||
<collectorName id="269D5321AE7DB610DF124667FD44034F" box="[539,671,296,320]" pageId="32" pageNumber="133">A. Newton</collectorName>
|
||
,
|
||
<collectorName id="269D5321AE7DB610DFB84667FCEB0330" box="[689,816,295,320]" pageId="32" pageNumber="133">M. Thayer</collectorName>
|
||
) (
|
||
<collectionCode id="ED79AE32AE7DB610DE444667FC7E0330" box="[845,933,296,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="32" pageNumber="133">FMNH</collectionCode>
|
||
);
|
||
<emphasis id="B91CEAE5AE7DB610DEB74668FB570330" bold="true" box="[958,1164,295,320]" pageId="32" pageNumber="133">
|
||
<specimenCount id="9D6EFD7EAE7DB610DEB74668FBCF0330" box="[958,1044,295,320]" pageId="32" pageNumber="133" type="male">1 male</specimenCount>
|
||
<typeStatus id="54D38855AE7DB610D9284668FB570330" box="[1057,1164,295,320]" pageId="32" pageNumber="133" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE7DB610D9A84668FA430330" box="[1185,1432,295,320]" country="Laos" name="Champasak" pageId="32" pageNumber="133">Champasak Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BB3AC8DAE7DB610DD9E4603FE8D0315" box="[151,342,332,357]" pageId="32" pageNumber="133">Bolaven Plateau</collectingMunicipality>
|
||
,
|
||
<location id="8EB7602CAE7DB610DC6E4602FDCF0315" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE7DB610DC6E4602FDCF0315" box="[359,532,332,357]" country="Laos" latitude="15.245684" longLatPrecision="1" longitude="106.53194" municipality="Bolaven Plateau" name="Ban Thongvay" pageId="32" pageNumber="133" stateProvince="Champasak Province">Ban Thongvay</location>
|
||
,
|
||
<geoCoordinate id="EE5C5030AE7DB610DF2C4603FD6E0315" box="[549,693,332,357]" direction="north" orientation="latitude" pageId="32" pageNumber="133" precision="1" value="15.245684">15°14.741'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5C5030AE7DB610DFCF4603FCBA0315" box="[710,865,332,357]" direction="east" orientation="longitude" pageId="32" pageNumber="133" precision="1" value="106.53194">106°31.916'E</geoCoordinate>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B003CAAAE7DB610DE7B4603FD2D03DD" ID-GBIF-Occurrence="1658411503" collectionCode="FMHD" collectorName="S. Tarasov" country="Laos" elevation="1150" location="Xekatam" pageId="32" pageNumber="133" specimenCount="1" specimenCount-female="1" stateProvince="Champasak Province" typeStatus="paratype">
|
||
<elevation id="0045D1C4AE7DB610DE7B4603FC150314" box="[882,974,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="32" pageNumber="133" unit="m" value="1150.0">
|
||
<quantity id="4C909B12AE7DB610DE7B4603FC150314" box="[882,974,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="32" pageNumber="133" unit="m" value="1150.0">1150 m</quantity>
|
||
</elevation>
|
||
, disturbed montane forest (
|
||
<collectorName id="269D5321AE7DB610D8114603FA4F0314" box="[1304,1428,332,356]" pageId="32" pageNumber="133">S. Tarasov</collectorName>
|
||
) (CST).
|
||
<emphasis id="B91CEAE5AE7DB610DDF84620FDB603F8" bold="true" box="[241,621,367,392]" pageId="32" pageNumber="133">
|
||
Additional specimen:
|
||
<specimenCount id="9D6EFD7EAE7DB610DF0A4620FDB603F8" box="[515,621,367,392]" pageId="32" pageNumber="133" type="female">1 female</specimenCount>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="49ACF815AE7DB610DF8B4620FCA103F8" box="[642,890,367,392]" country="Laos" name="Champasak" pageId="32" pageNumber="133">Champasak Province</collectingRegion>
|
||
: Bolaven Plateau, Ban Thôngvay (=
|
||
<location id="8EB7602CAE7DB610D8224620FA4F03F8" LSID="urn:lsid:plazi:treatment:03C187E1AE43B610DD9E44E7FB1F03A0:8EB7602CAE7DB610D8224620FA4F03F8" box="[1323,1428,367,392]" country="Laos" name="Xekatam" pageId="32" pageNumber="133" stateProvince="Champasak Province">Xékatam</location>
|
||
) vicinity, old logging road N of village,
|
||
<collectionCode id="ED79AE32AE7DB610DF5346DBFD7503DD" box="[602,686,404,429]" pageId="32" pageNumber="133">FMHD</collectionCode>
|
||
#2008
|
||
</materialsCitation>
|
||
-035, ANMT site 1230, 15°13.96'N, 106°31.731'E,
|
||
<quantity id="4C909B12AE7DB610D83746DBFA4C03DC" box="[1342,1431,404,429]" metricMagnitude="3" metricUnit="m" metricValue="1.035" pageId="32" pageNumber="133" unit="m" value="1035.0">1035 m</quantity>
|
||
, selectively logged forest, on
|
||
<date id="FFD61037AE7DB610DCD346F8FDB503A0" box="[474,622,439,464]" pageId="32" pageNumber="133" value="2008-06-08" valueMax="2008-06-16" valueMin="2008-06-08">8–16/vi/2008</date>
|
||
(A. Solodovnikov, M. Thayer, A. Newton) (FMNH).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |