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<document id="83EC585809762E2EE4FD1BE78C7067B1" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE40B62EDD9E42F4FB380288" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius maruyamai Masumoto, Ochi" docType="treatment" docVersion="12" lastPageNumber="130" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="130" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="DE4E045FDC63A9AC2E717D9BAB8E65F0">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="8A71284FCCE005EF8D91C1A3F4D5F311">Tarasov, Sergei</mods:namePart>
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<mods:date id="3618DF29B6AC1F6AC21BA1444D290B7B">2017</mods:date>
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<mods:identifier id="63070E2F8C5C8F39BCF4964722E094F4" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<treatment id="03C187E1AE40B62EDD9E42F4FB380288" ID-DOI="http://doi.org/10.5281/zenodo.6010832" ID-GBIF-Taxon="135106140" ID-Zenodo-Dep="6010832" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288" httpUri="http://treatment.plazi.org/id/03C187E1AE40B62EDD9E42F4FB380288" lastPageId="30" lastPageNumber="130" pageId="29" pageNumber="130">
<subSubSection id="C372657CAE40B62DDD9E42F4FEE40786" pageId="29" pageNumber="130" type="nomenclature">
<paragraph id="8BD736F7AE40B62DDD9E42F4FC6107A5" blockId="29.[151,954,1467,1526]" box="[151,954,1467,1493]" pageId="29" pageNumber="130">
<heading id="D09F819BAE40B62DDD9E42F4FC6107A5" bold="true" box="[151,954,1467,1493]" fontSize="11" level="1" pageId="29" pageNumber="130" reason="1">
<emphasis id="B91CEAE5AE40B62DDD9E42F4FC6107A5" bold="true" box="[151,954,1467,1493]" pageId="29" pageNumber="130">
<taxonomicName id="4C684D74AE40B62DDD9E42F4FD4307A5" ID-CoL="75NLV" authority="Masumoto, Ochi" authorityName="Masumoto, Ochi" box="[151,664,1467,1493]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="maruyamai">
<emphasis id="B91CEAE5AE40B62DDD9E42F4FE6407A5" bold="true" box="[151,447,1467,1493]" italics="true" pageId="29" pageNumber="130">Parachorius maruyamai</emphasis>
Masumoto, Ochi
</taxonomicName>
, &amp; Sakchoowong, 2012
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE40B62DDD9E4292FEE40786" blockId="29.[151,954,1467,1526]" box="[151,319,1501,1526]" pageId="29" pageNumber="130">
(
<figureCitation id="13532A72AE40B62DDD964292FF270786" box="[159,252,1501,1526]" captionStart="FIGURE 16" captionStartId="30.[151,250,803,825]" captionTargetBox="[346,1241,307,778]" captionTargetId="figure@30.[340,1247,303,781]" captionTargetPageId="30" captionText="FIGURE 16. Morphology of Parachorius maruyamai according to Masumoto et al. (2012). (A) habitus (male, holotype). (B) head and pronotum (dorsal view, scale 1 mm). (C) left metafemur (male). (D) aedeagus." httpUri="https://zenodo.org/record/1001949/files/figure.png" pageId="29" pageNumber="130">Figs. 16</figureCitation>
,
<figureCitation id="13532A72AE40B62DDC014292FEEC0786" box="[264,311,1501,1526]" captionStart="FIGURE 26" captionStartId="43.[151,250,1905,1927]" captionTargetBox="[151,1436,265,1825]" captionTargetId="figure@43.[151,1436,265,1825]" captionTargetPageId="43" captionText="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001969/files/figure.png" pageId="29" pageNumber="130">26E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE40B62DDD9E416AFCB3044B" box="[151,872,1573,1596]" pageId="29" pageNumber="130" type="description">
<paragraph id="8BD736F7AE40B62DDD9E416AFCB3044B" blockId="29.[151,872,1573,1596]" box="[151,872,1573,1596]" pageId="29" pageNumber="130">
<taxonomicName id="4C684D74AE40B62DDD9E416AFD99044B" authority="Masumoto, Ochi" authorityName="Masumoto, Ochi" box="[151,578,1573,1596]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="maruyamai">
<emphasis id="B91CEAE5AE40B62DDD9E416AFE4B044C" box="[151,400,1573,1596]" italics="true" pageId="29" pageNumber="130">Parachorius maruyamai</emphasis>
Masumoto, Ochi
</taxonomicName>
, &amp; Sakchoowong, 2012: 104
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE40B62DDD9E4127FC6804F1" box="[151,947,1640,1665]" pageId="29" pageNumber="130" type="materials_examined">
<paragraph id="8BD736F7AE40B62DDD9E4127FC6804F1" blockId="29.[151,1437,1640,2025]" box="[151,947,1640,1665]" pageId="29" pageNumber="130">
<materialsCitation id="3B003CAAAE40B62DDD9E4127FC6804F1" ID-GBIF-Occurrence="1658411485" box="[151,947,1640,1665]" country="Thailand" location="Khao Yai National Park" municipality="Kong Kaew" pageId="29" pageNumber="130" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEAE5AE40B62DDD9E4127FEF604F1" bold="true" box="[151,301,1640,1665]" pageId="29" pageNumber="130">
<typeStatus id="54D38855AE40B62DDD9E4127FF0904F1" box="[151,210,1640,1665]" pageId="29" pageNumber="130">Type</typeStatus>
locality
</emphasis>
.
<location id="8EB7602CAE40B62DDC354127FD9204F1" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE40B62DDC354127FD9204F1" box="[316,585,1640,1665]" country="Thailand" municipality="Kong Kaew" name="Khao Yai National Park" pageId="29" pageNumber="130">Khao Yai National Park</location>
,
<collectingMunicipality id="6BB3AC8DAE40B62DDF5C4126FD0004F0" box="[597,731,1641,1665]" pageId="29" pageNumber="130">Kong Kaew</collectingMunicipality>
[central
<collectingCountry id="F37F7667AE40B62DDE364127FC7204F1" box="[831,937,1640,1665]" name="Thailand" pageId="29" pageNumber="130">Thailand</collectingCountry>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE40B62DDDCE41C3FF220561" pageId="29" pageNumber="130" type="distribution">
<paragraph id="8BD736F7AE40B62DDDCE41C3FBF904D5" blockId="29.[151,1437,1640,2025]" box="[199,1058,1676,1701]" pageId="29" pageNumber="130">
<emphasis id="B91CEAE5AE40B62DDDCE41C3FEBB04D5" bold="true" box="[199,352,1676,1701]" pageId="29" pageNumber="130">Distribution.</emphasis>
Known only from Khao Tai National Park in central
<collectingCountry id="F37F7667AE40B62DDEBE41C3FBC504D5" box="[951,1054,1676,1701]" name="Thailand" pageId="29" pageNumber="130">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8BD736F7AE40B62DDDCE41FFFF220561" blockId="29.[151,1437,1640,2025]" pageId="29" pageNumber="130">
<emphasis id="B91CEAE5AE40B62DDDCE41FFFE4204B9" bold="true" box="[199,409,1712,1737]" pageId="29" pageNumber="130">Taxonomic notes.</emphasis>
Unfortunately, I was not able to study the
<typeStatus id="54D38855AE40B62DDE9D41FEFC1F04B9" box="[916,964,1713,1737]" pageId="29" pageNumber="130">type</typeStatus>
specimens of this species. However, the original description provides sufficient information to formulate its diagnosis and elucidate its phylogenetic position.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE40B62DDDCE4053FB6905B5" pageId="29" pageNumber="130" type="diagnosis">
<paragraph id="8BD736F7AE40B62DDDCE4053FE5005D1" blockId="29.[151,1437,1640,2025]" pageId="29" pageNumber="130">
<emphasis id="B91CEAE5AE40B62DDDCE4053FE9A0545" bold="true" box="[199,321,1820,1845]" pageId="29" pageNumber="130">Diagnosis.</emphasis>
Such characters as sharp frontal angles of pronotum and strongly curved metatibia suggest that this species is most closely related to
<taxonomicName id="4C684D74AE40B62DDF21400EFD0C0529" authorityName="Tarasov" authorityYear="2017" box="[552,727,1856,1881]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="newthayerae">
<emphasis id="B91CEAE5AE40B62DDF21400EFD0C0529" box="[552,727,1856,1881]" italics="true" pageId="29" pageNumber="130">P. newthayerae</emphasis>
</taxonomicName>
.
<emphasis id="B91CEAE5AE40B62DDFE3400FFD260529" bold="true" box="[746,765,1856,1881]" pageId="29" pageNumber="130">T</emphasis>
he aedeagus shape is similar in these two species, but
<taxonomicName id="4C684D74AE40B62DD88E400EFECD050C" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="maruyamai">
<emphasis id="B91CEAE5AE40B62DD88E400EFECD050C" italics="true" pageId="29" pageNumber="130">P. maruyamai</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C684D74AE40B62DDCA6402AFD80050D" authorityName="Tarasov" authorityYear="2017" box="[431,603,1892,1917]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="newthayerae">
<emphasis id="B91CEAE5AE40B62DDCA6402AFD80050D" box="[431,603,1892,1917]" italics="true" pageId="29" pageNumber="130">P. newthayerae</emphasis>
</taxonomicName>
by strongly sharped frontal angles of pronotum and not dilated posterior margin of metafemur.
</paragraph>
<paragraph id="8BD736F7AE40B62DDDCE40E3FB6905B5" blockId="29.[151,1437,1640,2025]" box="[199,1202,1964,1989]" pageId="29" pageNumber="130">
The identification using females is ambiguous, see
<taxonomicName id="4C684D74AE40B62DDE0D40E2FC7405B5" authorityName="Tarasov" authorityYear="2017" box="[772,943,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="130" phylum="Arthropoda" rank="species" species="newthayerae">
<emphasis id="B91CEAE5AE40B62DDE0D40E2FC7405B5" box="[772,943,1964,1989]" italics="true" pageId="29" pageNumber="130">P. newthayerae</emphasis>
</taxonomicName>
description for details.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE40B62EDDCE409FFB380288" lastPageId="30" lastPageNumber="131" pageId="29" pageNumber="130" type="materials_examined">
<paragraph id="8BD736F7AE40B62EDDCE409FFB380288" blockId="29.[151,1437,1640,2025]" lastBlockId="30.[151,1437,151,248]" lastPageId="30" lastPageNumber="131" pageId="29" pageNumber="130">
<materialsCitation id="3B003CAAAE40B62EDDCE409FFE4802A4" ID-GBIF-Occurrence="1658411491" collectingDate="2007-09-22" collectingDateMax="2007-09-28" collectingDateMin="2007-09-22" collectionCode="NSMT" collectorName="Maruyama &amp; Katayama Y." country="Thailand" county="Kong Kaew" elevation="700" lastPageId="30" lastPageNumber="131" latitude="14.438889" location="Material" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" pageId="29" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B91CEAE5AE40B62DDDCE409FFCBA0599" bold="true" box="[199,865,2000,2025]" pageId="29" pageNumber="130">
<location id="8EB7602CAE40B62DDDCE409FFEF40599" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE40B62DDDCE409FFEF40599" box="[199,303,2000,2025]" country="Thailand" county="Kong Kaew" latitude="14.438889" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" name="Material" pageId="29" pageNumber="130">Material</location>
examined in
<bibRefCitation id="EFF94B06AE40B62DDCDB409FFD3C0599" box="[466,743,2000,2025]" pageId="29" pageNumber="130" refString="Masumoto, K., Ochi, T. &amp; Sakehoowong, W. (2012) Scarabaeid beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Khao Yai in central Thailand collected by Dr. Munetoshi Maruyama. Kogane, 13, 103 - 124." type="journal article">
Masumoto
<emphasis id="B91CEAE5AE40B62DDF53409EFD490599" bold="true" box="[602,658,2000,2025]" italics="true" pageId="29" pageNumber="130">et al.</emphasis>
(2012)
</bibRefCitation>
.
<typeStatus id="54D38855AE40B62DDFFD409FFCBA0599" box="[756,865,2000,2025]" pageId="29" pageNumber="130" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F37F7667AE40B62DDE78409FFC050599" box="[881,990,2000,2025]" name="Thailand" pageId="29" pageNumber="130">
<emphasis id="B91CEAE5AE40B62DDE78409FFC050599" bold="true" box="[881,990,2000,2025]" pageId="29" pageNumber="130">Thailand</emphasis>
</collectingCountry>
:
<specimenCount id="9D6EFD7EAE40B62DDEE7409FFBE60599" box="[1006,1085,2000,2025]" pageId="29" pageNumber="130" type="male">
<emphasis id="B91CEAE5AE40B62DDEE7409FFBE60599" bold="true" box="[1006,1085,2000,2025]" pageId="29" pageNumber="130">1 male</emphasis>
</specimenCount>
, &quot;
<location id="8EB7602CAE40B62DD95F409FFB4B0599" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE40B62DD95F409FFB4B0599" box="[1110,1168,2000,2025]" country="Thailand" county="Kong Kaew" latitude="14.438889" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" name="Thai" pageId="29" pageNumber="130">Thai</location>
:
<location id="8EB7602CAE40B62DD995409FFA4B0599" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE40B62DD995409FFA4B0599" box="[1180,1424,2000,2025]" country="Thailand" county="Kong Kaew" latitude="14.438889" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" name="Nakhon Ratchashima" pageId="29" pageNumber="130">Nakhon Ratchashima</location>
,/
<collectingMunicipality id="6BB3AC8DAE43B62EDD9E47D8FE7002C0" box="[151,427,151,176]" pageId="30" pageNumber="131">Khao Yai National Park</collectingMunicipality>
,/
<collectingCounty id="62B64E7BAE43B62EDCB147D8FD9B02DF" box="[440,576,151,176]" pageId="30" pageNumber="131">Kong Kaew</collectingCounty>
(
<elevation id="0045D1C4AE43B62EDF5847D7FD4F02C0" box="[593,660,152,176]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="131" unit="m" value="700.0">
<quantity id="4C909B12AE43B62EDF5847D7FD4F02C0" box="[593,660,152,176]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="131" unit="m" value="700.0">700m</quantity>
</elevation>
)/
<collectingDate id="EF92E9DFAE43B62EDFAC47D7FC8B02C0" box="[677,848,152,176]" pageId="30" pageNumber="131" value="2007-09-22" valueMax="2007-09-28" valueMin="2007-09-22">
<date id="FFD61037AE43B62EDFAC47D7FC8B02C0" box="[677,848,152,176]" pageId="30" pageNumber="131" value="2007-09-22" valueMax="2007-09-28" valueMin="2007-09-22">22-28 IX 2007</date>
</collectingDate>
,
<collectorName id="269D5321AE43B62EDE5447D7FC0202C0" box="[861,985,152,176]" pageId="30" pageNumber="131">Maruyama</collectorName>
/M. &amp;
<collectorName id="269D5321AE43B62ED92E47D7FB6002C0" box="[1063,1211,152,176]" pageId="30" pageNumber="131">Katayama Y.</collectorName>
(FIT)&quot;,
<geoCoordinate id="EE5C5030AE43B62ED81847D8FA4302C0" box="[1297,1432,151,176]" direction="north" orientation="latitude" pageId="30" pageNumber="131" precision="15" value="14.438889">14°26'20&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE43B62EDD9E47F3FEFE02A4" box="[151,293,188,212]" direction="east" orientation="longitude" pageId="30" pageNumber="131" precision="15" value="101.37195">101°22'19&quot;E</geoCoordinate>
(
<collectionCode id="ED79AE32AE43B62EDC3E47F3FE5002A4" box="[311,395,188,212]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="30" pageNumber="131">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE43B62EDCA947F3FB040288" ID-GBIF-Occurrence="1658411470" collectingDate="2007-09-22" collectingDateMax="2007-09-28" collectingDateMin="2007-09-22" collectionCode="NSMT" collectorName="Maruyama &amp; Katayama Y." country="Thailand" county="Kong Kaew" elevation="700" latitude="14.438889" location="Thai" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" pageId="30" pageNumber="131" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<typeStatus id="54D38855AE43B62EDCA947F3FDD602A5" box="[416,525,188,213]" pageId="30" pageNumber="131" type="paratype">
<emphasis id="B91CEAE5AE43B62EDCA947F3FDD602A5" bold="true" box="[416,525,188,213]" pageId="30" pageNumber="131">Paratype</emphasis>
</typeStatus>
:
<specimenCount id="9D6EFD7EAE43B62EDF1747F3FD5402A5" box="[542,655,188,213]" pageId="30" pageNumber="131" type="female">
<emphasis id="B91CEAE5AE43B62EDF1747F3FD5402A5" bold="true" box="[542,655,188,213]" pageId="30" pageNumber="131">3 females</emphasis>
</specimenCount>
, &quot;
<location id="8EB7602CAE43B62EDFA047F3FD3802A5" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE43B62EDFA047F3FD3802A5" box="[681,739,188,213]" country="Thailand" county="Kong Kaew" latitude="14.438889" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" name="Thai" pageId="30" pageNumber="131">Thai</location>
:
<location id="8EB7602CAE43B62EDFF947F3FC3E02A5" LSID="urn:lsid:plazi:treatment:03C187E1AE40B62EDD9E42F4FB380288:8EB7602CAE43B62EDFF947F3FC3E02A5" box="[752,997,188,213]" country="Thailand" county="Kong Kaew" latitude="14.438889" longLatPrecision="21" longitude="101.37195" municipality="Khao Yai National Park" name="Nakhon Ratchashima" pageId="30" pageNumber="131">Nakhon Ratchashima</location>
,/
<collectingMunicipality id="6BB3AC8DAE43B62EDEFB47F3FADC02A5" box="[1010,1287,188,213]" pageId="30" pageNumber="131">Khao Yai National Park</collectingMunicipality>
,/
<collectingCounty id="62B64E7BAE43B62ED81A47F3FA4702A4" box="[1299,1436,188,213]" pageId="30" pageNumber="131">Kong Kaew</collectingCounty>
(
<elevation id="0045D1C4AE43B62EDD9747AFFF3A0288" box="[158,225,224,248]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="131" unit="m" value="700.0">
<quantity id="4C909B12AE43B62EDD9747AFFF3A0288" box="[158,225,224,248]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="131" unit="m" value="700.0">700m</quantity>
</elevation>
)/
<collectingDate id="EF92E9DFAE43B62EDDF847AFFE450288" box="[241,414,224,248]" pageId="30" pageNumber="131" value="2007-09-22" valueMax="2007-09-28" valueMin="2007-09-22">
<date id="FFD61037AE43B62EDDF847AFFE450288" box="[241,414,224,248]" pageId="30" pageNumber="131" value="2007-09-22" valueMax="2007-09-28" valueMin="2007-09-22">22–28 IX 2007</date>
</collectingDate>
,
<collectorName id="269D5321AE43B62EDCA047AFFDFE0288" box="[425,549,224,248]" pageId="30" pageNumber="131">Maruyama</collectorName>
/M. &amp;
<collectorName id="269D5321AE43B62EDF6447AFFCDB0288" box="[621,768,224,248]" pageId="30" pageNumber="131">Katayama Y.</collectorName>
(FIT)&quot;,
<geoCoordinate id="EE5C5030AE43B62EDE5D4790FC010288" box="[852,986,223,248]" direction="north" orientation="latitude" pageId="30" pageNumber="131" precision="15" value="14.438889">14°26'20&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE43B62EDEEC47AFFBAF0288" box="[997,1140,224,248]" direction="east" orientation="longitude" pageId="30" pageNumber="131" precision="15" value="101.37195">101°22'19&quot;E</geoCoordinate>
(
<collectionCode id="ED79AE32AE43B62ED98A47AFFB0C0288" box="[1155,1239,224,248]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="30" pageNumber="131">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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