427 lines
53 KiB
XML
427 lines
53 KiB
XML
<document id="B11FF3C0484E89CC029C1263D05BBA89" ID-CLB-Dataset="20923" ID-DOI="10.11646/zootaxa.5093.4.6" ID-GBIF-Dataset="7b7e1551-458c-4d35-ae65-af38e016ed91" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912813" ID-ZooBank="4691C842-ADF3-4231-B07F-9C9DDD14FEEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354835418" checkinUser="plazi" docAuthor="Nguyen, Tung T., Lam, Dang H. & Nguyen, Anh D." docDate="2022" docId="03C187BCFFA1FFF8FEC0FB0FFE77F8A1" docLanguage="en" docName="zootaxa.5093.4.6.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metaphire giengensis Nguyen & Lam & Nguyen 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="488" masterDocId="FFF8FFC4FFA5FFFDFE57FF85FF94FFA4" masterDocTitle="Two new earthworm species (Annelida, Oligochaeta, Megascolecidae) from the Mekong delta, Vietnam" masterLastPageNumber="492" masterPageNumber="483" pageNumber="487" updateTime="1699304635722" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8B4B7CBD4DEE4BCEBAF7954CCC5ABB89">
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<mods:title id="DF4FC4A310C1BFBC12D461E9C589B233">Two new earthworm species (Annelida, Oligochaeta, Megascolecidae) from the Mekong delta, Vietnam</mods:title>
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<mods:name id="D668F7A8EA1773FE2EC4288C71716962" type="personal">
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<mods:roleTerm id="D07F5DAA5FC31E099158FBCAC36A0FB4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E63CC43B376DCA4E647A70A14D9D471">Nguyen, Tung T.</mods:namePart>
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<mods:affiliation id="42E1716C466BA020CE10800EA0636ED8">Department of Biology, School of Education, Can Tho University, Can Tho City, Vietnam.</mods:affiliation>
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</mods:name>
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<mods:name id="71E391BAFEC6432E4AB767C8FEC195AD" type="personal">
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<mods:roleTerm id="2136D1A95ED45A30C41F9CC0EC41755A">Author</mods:roleTerm>
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<mods:namePart id="9968C2DDB262D6BC74FE9736688686AD">Lam, Dang H.</mods:namePart>
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<mods:nameIdentifier id="BB14E3AD86A66CB5FAFC14AEC90DE0B3" type="ORCID">0000-0002-1236-1087</mods:nameIdentifier>
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<mods:affiliation id="181829FF7006040ED0C93F33A5DAF462">Department of Biology, School of Education, Can Tho University, Can Tho City, Vietnam. & lamhaidangct @ gmail. com; https: // orcid. org / 0000 - 0002 - 1236 - 1087</mods:affiliation>
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<mods:nameIdentifier id="E0E5F2A493B683A0E835061A8809C731" type="email">lamhaidangct@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="6D1F4F0E812734FE300E2A4FECC71093">Author</mods:roleTerm>
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<mods:namePart id="62EA9BD09062AD25E837478AEF876599">Nguyen, Anh D.</mods:namePart>
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<mods:affiliation id="4795AE2923EEE782BF540415B924EF64">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
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<mods:typeOfResource id="517ECB1A8A5E74FA9EEBD3A3D200FAFB">text</mods:typeOfResource>
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<mods:title id="D780CB505B7807EE44B6ED39728A53D0">Zootaxa</mods:title>
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<mods:part id="1A5DB5905DDF2FDDC463869301C0A58E">
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<mods:date id="F6A03E166AE87E790F3F17284C211B4F">2022</mods:date>
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<mods:number id="C9075B710F32C53088F168D495DD00BF">2022-01-28</mods:number>
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<mods:number id="54CC6F9744323F83BC40C075005B332C">5093</mods:number>
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<treatment id="03C187BCFFA1FFF8FEC0FB0FFE77F8A1" ID-DOI="http://doi.org/10.5281/zenodo.5914111" ID-GBIF-Taxon="192701309" ID-Zenodo-Dep="5914111" LSID="urn:lsid:plazi:treatment:03C187BCFFA1FFF8FEC0FB0FFE77F8A1" httpUri="http://treatment.plazi.org/id/03C187BCFFA1FFF8FEC0FB0FFE77F8A1" lastPageId="5" lastPageNumber="488" pageId="4" pageNumber="487">
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<subSubSection id="C3726521FFA1FFF9FEC0FB0FFD94FB00" box="[151,512,1162,1189]" pageId="4" pageNumber="487" type="nomenclature">
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<paragraph id="8BD736AAFFA1FFF9FEC0FB0FFD94FB00" blockId="4.[151,512,1162,1224]" box="[151,512,1162,1189]" pageId="4" pageNumber="487">
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<heading id="D09F81C6FFA1FFF9FEC0FB0FFD94FB00" bold="true" box="[151,512,1162,1189]" fontSize="11" level="1" pageId="4" pageNumber="487" reason="1">
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<emphasis id="B91CEAB8FFA1FFF9FEC0FB0FFD94FB00" bold="true" box="[151,512,1162,1189]" pageId="4" pageNumber="487">
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<taxonomicName id="4C684D29FFA1FFF9FEC0FB0FFE0FFB00" authority="Nguyen & Lam & Nguyen, 2022" authorityName="Nguyen & Lam & Nguyen" authorityYear="2022" box="[151,411,1162,1189]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="487" phylum="Annelida" rank="species" species="giengensis" status="sp. nov.">
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<emphasis id="B91CEAB8FFA1FFF9FEC0FB0FFE0FFB00" bold="true" box="[151,411,1162,1189]" italics="true" pageId="4" pageNumber="487">Metaphire giengensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A22F57C3FFA1FFF9FFF5FB0FFD94FB00" box="[418,512,1162,1188]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3726521FFA1FFF9FEC0FB2BFF7FFB6C" box="[151,235,1198,1224]" pageId="4" pageNumber="487" type="description">
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<paragraph id="8BD736AAFFA1FFF9FEC0FB2BFF7FFB6C" blockId="4.[151,512,1162,1224]" box="[151,235,1198,1224]" pageId="4" pageNumber="487">
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(
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<figureCitation id="13532A2FFFA1FFF9FEF7FB2BFF76FB6C" box="[160,226,1198,1224]" captionStart="FIGURE 3" captionStartId="6.[151,250,1809,1833]" captionTargetBox="[158,1431,181,1784]" captionTargetId="figure-21@6.[151,1436,181,1785]" captionTargetPageId="6" captionText="FIGURE 3. Metaphire giengensis sp. nov., Holotype (CTU-EW.017.h01). A1 & A2. Ventral view of the male region. B. Dorsal view of testis sacs and seminal vesicles. C1 & C2. Right spermatheca. D. Ventral view of the spermathecal region. E1 & E2. Intestinal caecum; F1 & F2. Right prostate gland. G. Male pore region transverse body section (Non-type: CTU- EW.017.03). H. Body wall transverse section. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5912819" httpUri="https://zenodo.org/record/5912819/files/figure.png" pageId="4" pageNumber="487">Fig. 3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3726521FFA1FFF9FEC0FB73FC5EFA61" pageId="4" pageNumber="487" type="materials_examined">
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<paragraph id="8BD736AAFFA1FFF9FEC0FB73FC5EFA61" blockId="4.[151,1437,1270,2017]" pageId="4" pageNumber="487">
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||
<emphasis id="B91CEAB8FFA1FFF9FEC0FB73FE10FAB4" bold="true" box="[151,388,1270,1296]" pageId="4" pageNumber="487">Material examined.</emphasis>
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<materialsCitation id="3B003CF7FFA1FFF9FFD9FB73FEC3FAFC" ID-GBIF-Occurrence="3458738301" collectingDate="2010-02-02" collectionCode="CTU-EW" collectorName="Ho Minh Thuan." country="Vietnam" elevation="2" latitude="10.511027" location="Tan My Village" longLatPrecision="1" longitude="105.50075" municipality="Cho Moi District" pageId="4" pageNumber="487" specimenCount="1" stateProvince="An Giang" typeStatus="holotype">
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<emphasis id="B91CEAB8FFA1FFF9FFD9FB73FE6AFAB4" box="[398,510,1270,1296]" italics="true" pageId="4" pageNumber="487">
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<typeStatus id="54D38808FFA1FFF9FFD9FB73FE6CFAB4" box="[398,504,1270,1296]" pageId="4" pageNumber="487" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
1 mature, (
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||
<collectionCode id="ED79AE6FFFA1FFF9FCDFFB73FD6AFAB5" box="[648,766,1270,1297]" pageId="4" pageNumber="487">CTU-EW</collectionCode>
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||
.017.h01),
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<collectingCountry id="F37F763AFFA1FFF9FD27FB73FC46FAB4" box="[880,978,1270,1296]" name="Vietnam" pageId="4" pageNumber="487">Vietnam</collectingCountry>
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,
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<collectingRegion id="49ACF848FFA1FFF9FD88FB73FB54FAB4" box="[991,1216,1270,1296]" country="Vietnam" name="An Giang" pageId="4" pageNumber="487">An Giang Province</collectingRegion>
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||
,
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||
<collectingMunicipality id="6BB3ACD0FFA1FFF9FA98FB73FA0CFAB4" box="[1231,1432,1270,1296]" pageId="4" pageNumber="487">Cho Moi District</collectingMunicipality>
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,
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||
<location id="8EB76071FFA1FFF9FEC0FA9EFED8FA90" LSID="urn:lsid:plazi:treatment:03C187BCFFA1FFF8FEC0FB0FFE77F8A1:8EB76071FFA1FFF9FEC0FA9EFED8FA90" box="[151,332,1306,1333]" country="Vietnam" latitude="10.511027" longLatPrecision="1" longitude="105.50075" municipality="Cho Moi District" name="Tan My Village" pageId="4" pageNumber="487" stateProvince="An Giang">Tan My Village</location>
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||
(Gieng islet), mango garden (
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<geoCoordinate id="EE5C506DFFA1FFF9FCFDFA9FFCA9FA90" box="[682,829,1306,1332]" degrees="10" direction="north" minutes="30" orientation="latitude" pageId="4" pageNumber="487" precision="1" seconds="39.7" value="10.511027">10°30’39.7N</geoCoordinate>
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-
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<geoCoordinate id="EE5C506DFFA1FFF9FD0DFA9FFC6EFA91" box="[858,1018,1306,1333]" degrees="105" direction="east" minutes="30" orientation="longitude" pageId="4" pageNumber="487" precision="1" seconds="02.7" value="105.50075">105°30’02.7E</geoCoordinate>
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||
),
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||
<elevation id="0045D199FFA1FFF9FA46FA9FFBEAFA90" box="[1041,1150,1306,1333]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="487" unit="m" value="2.0">
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<quantity id="4C909B4FFFA1FFF9FA46FA9FFBABFA91" box="[1041,1087,1306,1333]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="487" unit="m" value="2.0">2 m</quantity>
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||
a.s.l.
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||
</elevation>
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||
,
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||
<date id="FFD6106AFFA1FFF9FADAFA9FFAC9FA90" box="[1165,1373,1306,1332]" pageId="4" pageNumber="487" value="2010-02-02">
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<collectingDate id="EF92E982FFA1FFF9FADAFA9FFAC9FA90" box="[1165,1373,1306,1332]" pageId="4" pageNumber="487" value="2010-02-02">02 February 2010</collectingDate>
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||
</date>
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||
, coll.
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||
<collectorName id="269D537CFFA1FFF9FEC0FABAFEC3FAFC" box="[151,343,1342,1369]" pageId="4" pageNumber="487">Ho Minh Thuan.</collectorName>
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</materialsCitation>
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<materialsCitation id="3B003CF7FFA1FFF9FF08FABAFA83FAD9" ID-GBIF-Occurrence="3458738306" collectingDate="2020-11-02" collectionCode="CTU-EW" collectorName="Lam Hai Dang." country="Vietnam" elevation="2" latitude="10.462" location="Tan My Village" longLatPrecision="1" longitude="105.52986" municipality="Cho Moi District" pageId="4" pageNumber="487" specimenCount="3" stateProvince="An Giang" typeStatus="paratype">
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||
<emphasis id="B91CEAB8FFA1FFF9FF08FABAFE52FAFC" box="[351,454,1343,1368]" italics="true" pageId="4" pageNumber="487">
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||
<typeStatus id="54D38808FFA1FFF9FF08FABAFE52FAFC" box="[351,454,1343,1368]" pageId="4" pageNumber="487" type="paratype">Paratype</typeStatus>
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||
</emphasis>
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||
s: 12 matures, (
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||
<collectionCode id="ED79AE6FFFA1FFF9FC20FABBFD78FAFC" box="[631,748,1342,1368]" pageId="4" pageNumber="487">CTU-EW</collectionCode>
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||
.017.p02), same data as for holotype.
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<emphasis id="B91CEAB8FFA1FFF9FAC2FABAFA93FAFC" box="[1173,1287,1343,1368]" italics="true" pageId="4" pageNumber="487">Non-types</emphasis>
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||
: 31 matures, (
|
||
<collectionCode id="ED79AE6FFFA1FFF9FEC8FAE7FE85FAD8" box="[159,273,1378,1404]" pageId="4" pageNumber="487">CTU-EW</collectionCode>
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||
.017.03), same data as for
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||
<typeStatus id="54D38808FFA1FFF9FC1CFAE7FD24FAD8" box="[587,688,1378,1404]" pageId="4" pageNumber="487" type="holotype">holotype</typeStatus>
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||
, but in mango garden (
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||
<geoCoordinate id="EE5C506DFFA1FFF9FD9DFAE7FBDCFAD9" box="[970,1096,1378,1405]" degrees="10" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="487" precision="15" seconds="45" value="10.479166">10°28’45N</geoCoordinate>
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||
-
|
||
<geoCoordinate id="EE5C506DFFA1FFF9FA3EFAE7FA9EFAD9" box="[1129,1290,1378,1405]" degrees="105" direction="east" minutes="32" orientation="longitude" pageId="4" pageNumber="487" precision="1" seconds="56.6" value="105.54906">105°32’56.6E</geoCoordinate>
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||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B003CF7FFA1FFF9FB74FAE7FC5EFA61" ID-GBIF-Occurrence="3458738303" collectingDate="2020-11-02" collectionCode="CTU-EW" collectorName="Lam Hai Dang." country="Vietnam" elevation="2" latitude="10.462" location="Tan My Village" longLatPrecision="1" longitude="105.52986" municipality="Cho Moi District" pageId="4" pageNumber="487" specimenCount="3" stateProvince="An Giang" typeStatus="holotype">
|
||
5 matures, (
|
||
<collectionCode id="ED79AE6FFFA1FFF9FEC8FA03FE9BFA04" box="[159,271,1414,1440]" pageId="4" pageNumber="487">CTU-EW</collectionCode>
|
||
.017.04) and 6 matures, (
|
||
<collectionCode id="ED79AE6FFFA1FFF9FC73FA03FD01FA04" box="[548,661,1414,1440]" pageId="4" pageNumber="487">CTU-EW</collectionCode>
|
||
.DNA.017.04) same locality as for
|
||
<typeStatus id="54D38808FFA1FFF9FA4BFA03FB15FA04" box="[1052,1153,1414,1440]" pageId="4" pageNumber="487" type="holotype">holotype</typeStatus>
|
||
, but in banana plantation (
|
||
<geoCoordinate id="EE5C506DFFA1FFF9FEC8FA2FFEA6FA60" box="[159,306,1450,1476]" degrees="10" direction="north" minutes="27" orientation="latitude" pageId="4" pageNumber="487" precision="1" seconds="43,2" value="10.462">10°27’43,2N</geoCoordinate>
|
||
-
|
||
<geoCoordinate id="EE5C506DFFA1FFF9FF1DFA2FFE7DFA61" box="[330,489,1450,1477]" degrees="105" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="487" precision="1" seconds="47,5" value="105.52986">105°31’47,5E</geoCoordinate>
|
||
),
|
||
<date id="FFD6106AFFA1FFF9FFA9FA2FFD4DFA60" box="[510,729,1450,1476]" pageId="4" pageNumber="487" value="2020-11-02">
|
||
<collectingDate id="EF92E982FFA1FFF9FFA9FA2FFD4DFA60" box="[510,729,1450,1476]" pageId="4" pageNumber="487" value="2020-11-02">02 November 2020</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="269D537CFFA1FFF9FD4BFA2EFC5EFA61" box="[796,970,1450,1477]" pageId="4" pageNumber="487">Lam Hai Dang.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726521FFA1FFF9FE90FA4BFCC7FA4C" box="[199,851,1486,1513]" pageId="4" pageNumber="487" type="etymology">
|
||
<paragraph id="8BD736AAFFA1FFF9FE90FA4BFCC7FA4C" blockId="4.[151,1437,1270,2017]" box="[199,851,1486,1513]" pageId="4" pageNumber="487">
|
||
<emphasis id="B91CEAB8FFA1FFF9FE90FA4BFED8FA4C" bold="true" box="[199,332,1486,1512]" pageId="4" pageNumber="487">Etymology.</emphasis>
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||
named after its
|
||
<typeStatus id="54D38808FFA1FFF9FFA8FA4AFDBBFA4D" box="[511,559,1487,1513]" pageId="4" pageNumber="487">type</typeStatus>
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||
locality, the “gieng” islet.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726521FFA1FFF9FE90FA77FB80F938" pageId="4" pageNumber="487" type="diagnosis">
|
||
<paragraph id="8BD736AAFFA1FFF9FE90FA77FB80F938" blockId="4.[151,1437,1270,2017]" pageId="4" pageNumber="487">
|
||
<emphasis id="B91CEAB8FFA1FFF9FE90FA77FED6F9A8" bold="true" box="[199,322,1522,1548]" pageId="4" pageNumber="487">Diagnosis.</emphasis>
|
||
Small-sized worm, length
|
||
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|
||
, diameter 1.8–2.0 mm, segments 76–131. Prostomium epilobous. First dorsal pore in 11/12. Two pairs of spermathecal pores in ventral intersegments 5/6/7. Male pores paired in xviii; ventral distance between
|
||
<specimenCount id="9D6EFD23FFA1FFF9FC01F9BEFD2FF9F0" box="[598,699,1594,1621]" pageId="4" pageNumber="487" type="male">two male</specimenCount>
|
||
pores about 0.20–0.22X body circumference. No genital markings in the spermathecal pore region, but two pairs, bean-shaped in xviii, ventrally next to secondary male pore. Holandric. Testis sacs connected. Intestinal caeca simple. Septa 8/9 thickened, 9/10 absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726521FFA1FFF8FE90F923FCFAFCF9" lastPageId="5" lastPageNumber="488" pageId="4" pageNumber="487" type="description">
|
||
<paragraph id="8BD736AAFFA1FFF9FE90F923FF55F8D0" blockId="4.[151,1437,1270,2017]" pageId="4" pageNumber="487">
|
||
<emphasis id="B91CEAB8FFA1FFF9FE90F923FECDF964" bold="true" box="[199,345,1702,1728]" pageId="4" pageNumber="487">Description.</emphasis>
|
||
<emphasis id="B91CEAB8FFA1FFF9FF32F923FDC5F964" box="[357,593,1702,1728]" italics="true" pageId="4" pageNumber="487">External characters.</emphasis>
|
||
Body cylindrical, small size, length
|
||
<quantity id="4C909B4FFFA1FFF9FA5DF923FB1EF965" box="[1034,1162,1702,1729]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="6.7" metricValueMin="3.9" pageId="4" pageNumber="487" unit="mm" value="53.0" valueMax="67.0" valueMin="39.0">39–67 mm</quantity>
|
||
, diameter 1.8–2.0 mm, segments 76–131. Body colouration uniformly light brown; clitellum dark brown. Prostomium 1/3 epilobous. First dorsal pore in 11/12. Setae perichaetine, pre-clitellar setae stouter and denser than post-clitellar ones; setal numbers:
|
||
<quantity id="4C909B4FFFA1FFF9FEC0F897FF6EF888" box="[151,250,1810,1836]" metricMagnitude="0" metricUnit="m" metricValue="2.0193" metricValueMax="2.1082" metricValueMin="1.9304" pageId="4" pageNumber="487" unit="in" value="79.5" valueMax="83.0" valueMin="76.0">76–83 in</quantity>
|
||
viii,
|
||
<quantity id="4C909B4FFFA1FFF9FF63F897FE03F888" box="[308,407,1810,1836]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.5493999999999999" metricValueMin="1.2446000000000002" pageId="4" pageNumber="487" unit="in" value="55.0" valueMax="61.0" valueMin="49.0">49–61 in</quantity>
|
||
xxx, 6–9 between male porophores in xviii; setal distance: aa=ab, zz>zy. Clitellum annular, within xiv–xvi, without dorsal pores; clitellar setae present only in xvi ventrally. Female pore single, in mid-ventral xiv.
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA1FFF8FE90F8FAFC17FF5D" blockId="4.[151,1437,1270,2017]" lastBlockId="5.[151,1437,151,1797]" lastPageId="5" lastPageNumber="488" pageId="4" pageNumber="487">Two pairs of spermathecal pores located ventrally in intersegment 5/6/7; ventral distance between pores about 0.15-0.18X body circumference. No genital markings in the spermathecal region. Male pores located inside copulatory pouches which open ventrally on the setal ring of xviii; ventral distance between two openings of copulatory pouches about 0.20-0.22X body circumference; Each copulatory pouch is a shallow depression of the body wall with a large skin on the lateral side forming the openings of the pouch. Two pairs of genital markings in xviii, located medially next to the openings of copulatory pouches.</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FE86FE88FE2D" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FE86FE38FEB9" box="[199,428,259,285]" italics="true" pageId="5" pageNumber="488">Internal characters.</emphasis>
|
||
Septa 5/6/7/8/9 thick, 9/10 absent, 10/11/12/13 thin. Oesophageal gizzard within viii. Intestinal origin at xvi; caeca simple, originating at xxvii and extending anteriorly to xxv. Last hearts in xiii. Pharyngeal micronephridia meroic, developed in 4/5/6/7. Typhlosole simple, lamelliform. Lymph glands present from 29/30.
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FE16FC8FFE51" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">Spermathecae paired in vi and vii. Ampulla oval-shaped to capsule-shaped; ducts about 1/2 ampulla, robust. Diverticula cylindrical, not coiled, somewhat slightly wavy, ca. twice as long as ampulla; seminal chamber very reduced. No accessory glands in the spermathecal region.</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FE7AFD65FD21" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">Holandric. Testis sacs developed, connected in x–xi. Seminal vesicles well developed in xi–xii enclosing the second pair of testis sacs laterally. Ovaries well developed xiii, on ventral posterior face of septum 12/13. Ovisac invisible. Prostate glands rudimentary lobes, paired within xvii–xx; ducts robust, enlarged basally and directly attached to the body wall. Accessory glands invisible.</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FD0BFC6DFCB1" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FD0BFEEFFD0D" bold="true" box="[199,379,654,681]" pageId="5" pageNumber="488">DNA barcode.</emphasis>
|
||
DNA materials were extracted from CTU-EW.DNA.017.04. Partial sequences of the mitochondrial gene cytochrome oxidase subunit I (COI) were uploaded to GenBank with the accession numbers
|
||
<accessionNumber id="943BAB49FFA0FFF8FEC0FD52FE8DFD55" box="[151,281,727,753]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/OM234761" pageId="5" pageNumber="488" type="GenBank">OM234761</accessionNumber>
|
||
and
|
||
<accessionNumber id="943BAB49FFA0FFF8FF02FD52FE4EFD55" box="[341,474,727,753]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/OM234762" pageId="5" pageNumber="488" type="GenBank">OM234762</accessionNumber>
|
||
. The new species has a close COI identity of 86.56% and 85.89% with
|
||
<taxonomicName id="4C684D29FFA0FFF8FB7EFD52FE87FCB1" baseAuthorityName="Michaelsen" baseAuthorityYear="1928" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="tschiliensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FB7EFD52FE87FCB1" italics="true" pageId="5" pageNumber="488">Metaphire tschiliensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<accessionNumber id="943BAB49FFA0FFF8FF75FD7FFE36FCB1" box="[290,418,762,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ835673" pageId="5" pageNumber="488">DQ835673</accessionNumber>
|
||
) and
|
||
<taxonomicName id="4C684D29FFA0FFF8FFB7FD7EFD5CFCB1" baseAuthorityName="Vaillant" baseAuthorityYear="1868" box="[480,712,763,789]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="posthuma">
|
||
<emphasis id="B91CEAB8FFA0FFF8FFB7FD7EFD5CFCB1" box="[480,712,763,789]" italics="true" pageId="5" pageNumber="488">Metaphire posthuma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<accessionNumber id="943BAB49FFA0FFF8FC81FD7FFCCDFCB1" box="[726,857,762,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU565289" pageId="5" pageNumber="488">KU565289</accessionNumber>
|
||
), respectively.
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FC9BFCFAFCF9" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FC9BFE50FC9D" bold="true" box="[199,452,798,825]" pageId="5" pageNumber="488">Habitat and ecology:</emphasis>
|
||
The species was found in the depth of
|
||
<quantity id="4C909B4FFFA0FFF8FDDCFC9AFC72FC9D" box="[907,998,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="5" pageNumber="488" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10cm</quantity>
|
||
of alluvial soils in banana and mango plantations. Feces were small, pellet-like and lying underground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726521FFA0FFF8FE90FCE3FD0BFC25" box="[199,671,870,897]" pageId="5" pageNumber="488" type="distribution">
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FCE3FD0BFC25" blockId="5.[151,1437,151,1797]" box="[199,671,870,897]" pageId="5" pageNumber="488">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FCE3FEF7FC24" bold="true" box="[199,355,870,896]" pageId="5" pageNumber="488">Distribution:</emphasis>
|
||
Only known from
|
||
<collectingCountry id="F37F763AFFA0FFF8FC6EFCE2FD08FC25" box="[569,668,871,897]" name="Vietnam" pageId="5" pageNumber="488">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726521FFA0FFF8FE90FC0EFE77F8A1" pageId="5" pageNumber="488" type="discussion">
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FC0EFE76FB91" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FC0EFEA6FC01" bold="true" box="[199,306,907,933]" pageId="5" pageNumber="488">Remark:</emphasis>
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FF6EFC0EFDA4FC01" author="Sims, R. W. & Easton, E. G." box="[313,560,907,933]" pageId="5" pageNumber="488" pagination="169 - 268" refId="ref6136" refString="Sims, R. W. & Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4 (3), 169 - 268. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims & Easton (1972)</bibRefCitation>
|
||
arranged the species of the genus
|
||
<taxonomicName id="4C684D29FFA0FFF8FDFBFC0EFBB4FC01" authorityName="Sims & Easton" authorityYear="1972" box="[940,1056,907,933]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDFBFC0EFBB4FC01" box="[940,1056,907,933]" italics="true" pageId="5" pageNumber="488">Metaphire</emphasis>
|
||
</taxonomicName>
|
||
into lots of species-groups at very detailed levels. However,
|
||
<taxonomicName id="4C684D29FFA0FFF8FF95FC2AFDA2FC6D" authorityName="Sims & Easton" authorityYear="1972" box="[450,566,943,969]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF95FC2AFDA2FC6D" box="[450,566,943,969]" italics="true" pageId="5" pageNumber="488">Metaphire</emphasis>
|
||
</taxonomicName>
|
||
’s species with two pairs of spermathecal pores in 5/6/7 and bithecate were not yet included in any groups, and up to now, only three species with that spermathecal configuration have been recorded:
|
||
<taxonomicName id="4C684D29FFA0FFF8FF5FFC72FD9DFBB5" authority="(Gates, 1935)" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[264,521,1014,1041]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exilis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF5FFC72FEFCFBB5" box="[264,360,1015,1041]" italics="true" pageId="5" pageNumber="488">M. exilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FF21FC72FD94FBB5" author="Gates, G. E." box="[374,512,1014,1041]" pageId="5" pageNumber="488" pagination="1 - 19" refId="ref5062" refString="Gates, G. E. (1935) New earthworms from China: with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Institution, 3293 (3), 1 - 19." type="journal article" year="1935">Gates, 1935</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D29FFA0FFF8FC43FC72FCADFBB6" authority="(Chen, 1936)" baseAuthorityName="Chen" baseAuthorityYear="1936" box="[532,825,1015,1042]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exiloides">
|
||
<emphasis id="B91CEAB8FFA0FFF8FC43FC72FD09FBB5" box="[532,669,1015,1041]" italics="true" pageId="5" pageNumber="488">M. exiloides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FCFCFC72FCA6FBB6" author="Chen, Y." box="[683,818,1015,1042]" pageId="5" pageNumber="488" pagination="269 - 306" refId="ref4782" refString="Chen, Y. (1936) On the Terrestrial Oligochaeta from Szechuan, II.: With the Notes on Gates' Types. Contributions from the Biological Laboratory of the Science Society of China. Zoological series, 11 (8), 269 - 306." type="journal article" year="1936">Chen, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C684D29FFA0FFF8FD23FC72FA92FBB5" authority="(Thai & Huynh, 1993)" baseAuthorityName="Thai & Huynh" baseAuthorityYear="1993" box="[884,1286,1015,1041]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="easupana">
|
||
<emphasis id="B91CEAB8FFA0FFF8FD23FC72FB90FBB4" box="[884,1028,1015,1040]" italics="true" pageId="5" pageNumber="488">M. easupana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FA44FC72FB6AFBB5" author="Thai, B. T. & Do, N. V. & Huynh, H. T. K." box="[1043,1278,1015,1041]" pageId="5" pageNumber="488" pagination="12 - 15" refId="ref6225" refString="Thai, B. T., Do, N. V. & Huynh, H. T. K. (1993) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae - Oligochaeta) from Yokdon, Dac Lac Province. Tap chi Sinh hoc, 15 (4), 12 - 15. [in Vietnamese]" type="journal article" year="1993">Thai & Huynh, 1993</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FB43FC72FA03FBB5" author="Gates, G. E." box="[1300,1431,1014,1041]" pageId="5" pageNumber="488" pagination="1 - 19" refId="ref5062" refString="Gates, G. E. (1935) New earthworms from China: with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Institution, 3293 (3), 1 - 19." type="journal article" year="1935">Gates 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FEC0FB9EFE80FB91" author="Chen, Y." box="[151,276,1051,1077]" pageId="5" pageNumber="488" pagination="269 - 306" refId="ref4782" refString="Chen, Y. (1936) On the Terrestrial Oligochaeta from Szechuan, II.: With the Notes on Gates' Types. Contributions from the Biological Laboratory of the Science Society of China. Zoological series, 11 (8), 269 - 306." type="journal article" year="1936">Chen 1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FF77FB9EFE41FB91" author="Thai, B. T. & Do, N. V. & Huynh, H. T. K." box="[288,469,1051,1077]" pageId="5" pageNumber="488" pagination="12 - 15" refId="ref6225" refString="Thai, B. T., Do, N. V. & Huynh, H. T. K. (1993) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae - Oligochaeta) from Yokdon, Dac Lac Province. Tap chi Sinh hoc, 15 (4), 12 - 15. [in Vietnamese]" type="journal article" year="1993">
|
||
Thai
|
||
<emphasis id="B91CEAB8FFA0FFF8FF0DFB9EFE1FFB91" box="[346,395,1051,1077]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
|
||
. 1993
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FBBAFBB5FB61" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<taxonomicName id="4C684D29FFA0FFF8FE90FBBAFE2EFBFC" authority="Nguyen & Lam & Nguyen, 2022" authorityName="Nguyen & Lam & Nguyen" authorityYear="2022" box="[199,442,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="giengensis" status="sp. nov.">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FBBAFE2EFBFC" box="[199,442,1087,1113]" italics="true" pageId="5" pageNumber="488">Metaphire giengensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CEAB8FFA0FFF8FF93FBBAFDB4FBFD" bold="true" box="[452,544,1087,1113]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="A22F57C3FFA0FFF8FF93FBBAFDB4FBFD" box="[452,544,1087,1113]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is fairly similar to
|
||
<taxonomicName id="4C684D29FFA0FFF8FD56FBBAFCF0FBFD" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[769,868,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exilis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FD56FBBAFCF0FBFD" box="[769,868,1087,1113]" italics="true" pageId="5" pageNumber="488">M. exilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C684D29FFA0FFF8FDC8FBBAFBBFFBFD" baseAuthorityName="Chen" baseAuthorityYear="1936" box="[927,1067,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exiloides">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDC8FBBAFBBFFBFD" box="[927,1067,1087,1113]" italics="true" pageId="5" pageNumber="488">M. exiloides</emphasis>
|
||
</taxonomicName>
|
||
in the prostomium
|
||
<typeStatus id="54D38808FFA0FFF8FB58FBBAFAABFBFD" box="[1295,1343,1087,1113]" pageId="5" pageNumber="488">type</typeStatus>
|
||
and the presence of septum 8/9. However, the new species is different from
|
||
<taxonomicName id="4C684D29FFA0FFF8FDF1FBE6FB9CFBD9" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[934,1032,1123,1149]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exilis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDF1FBE6FB9CFBD9" box="[934,1032,1123,1149]" italics="true" pageId="5" pageNumber="488">M. exilis</emphasis>
|
||
</taxonomicName>
|
||
in having genital markings only in xviii, connected testis sacs, oval/capsule-shaped spermathecal ampulla; while
|
||
<taxonomicName id="4C684D29FFA0FFF8FA56FB02FBF5FB05" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[1025,1121,1159,1185]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exilis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FA56FB02FBF5FB05" box="[1025,1121,1159,1185]" italics="true" pageId="5" pageNumber="488">M. exilis</emphasis>
|
||
</taxonomicName>
|
||
has genital markings paired in xvii and xix, separated testis sacs, and strongly reduced ampulla (
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FDC7FB2EFB81FB61" author="Gates, G. E." box="[912,1045,1194,1221]" pageId="5" pageNumber="488" pagination="1 - 19" refId="ref5062" refString="Gates, G. E. (1935) New earthworms from China: with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Institution, 3293 (3), 1 - 19." type="journal article" year="1935">Gates 1935</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FB4AFB0CFA96" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
The new species is distinguished from
|
||
<taxonomicName id="4C684D29FFA0FFF8FC23FB4AFD68FB4D" baseAuthorityName="Chen" baseAuthorityYear="1936" box="[628,764,1231,1257]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exiloides">
|
||
<emphasis id="B91CEAB8FFA0FFF8FC23FB4AFD68FB4D" box="[628,764,1231,1257]" italics="true" pageId="5" pageNumber="488">M. exiloides</emphasis>
|
||
</taxonomicName>
|
||
by having two pairs of genital markings in xviii, the absence of septum 9/10, spermatheal diverticula being longer than ampulla; while
|
||
<taxonomicName id="4C684D29FFA0FFF8FDB9FB76FBEDFAA9" baseAuthorityName="Chen" baseAuthorityYear="1936" box="[1006,1145,1267,1293]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="exiloides">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDB9FB76FBEDFAA9" box="[1006,1145,1267,1293]" italics="true" pageId="5" pageNumber="488">M. exiloides</emphasis>
|
||
</taxonomicName>
|
||
has no genital markings, presence of septum 9/10, diverticula being wavy and shorter than the ampulla (
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FA5CFA92FB1FFA96" author="Chen, Y." box="[1035,1163,1303,1330]" pageId="5" pageNumber="488" pagination="269 - 306" refId="ref4782" refString="Chen, Y. (1936) On the Terrestrial Oligochaeta from Szechuan, II.: With the Notes on Gates' Types. Contributions from the Biological Laboratory of the Science Society of China. Zoological series, 11 (8), 269 - 306." type="journal article" year="1936">Chen 1936</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FABEFD48FA65" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
<taxonomicName id="4C684D29FFA0FFF8FE90FABEFE2FFAF0" authority="Nguyen & Lam & Nguyen, 2022" authorityName="Nguyen & Lam & Nguyen" authorityYear="2022" box="[199,443,1339,1365]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="giengensis" status="sp. nov.">
|
||
<emphasis id="B91CEAB8FFA0FFF8FE90FABEFE2FFAF0" box="[199,443,1339,1365]" italics="true" pageId="5" pageNumber="488">Metaphire giengensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CEAB8FFA0FFF8FF92FABEFDB5FAF1" bold="true" box="[453,545,1339,1365]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="A22F57C3FFA0FFF8FF92FABEFDB5FAF1" box="[453,545,1339,1365]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C684D29FFA0FFF8FCE9FABEFCC5FAF0" baseAuthorityName="Thai & Huynh" baseAuthorityYear="1993" box="[702,849,1339,1364]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="easupana">
|
||
<emphasis id="B91CEAB8FFA0FFF8FCE9FABEFCC5FAF0" box="[702,849,1339,1364]" italics="true" pageId="5" pageNumber="488">M. easupana</emphasis>
|
||
</taxonomicName>
|
||
in having the spermathecal pores placed ventrally, first dorsal pore in 11/12, two pairs of genital markings in xviii, intestinal origin at xv with simple caeca; while
|
||
<taxonomicName id="4C684D29FFA0FFF8FB29FADAFE96FA38" baseAuthorityName="Thai & Huynh" baseAuthorityYear="1993" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="easupana">
|
||
<emphasis id="B91CEAB8FFA0FFF8FB29FADAFE96FA38" italics="true" pageId="5" pageNumber="488">M. easupana</emphasis>
|
||
</taxonomicName>
|
||
has spermathecal pores located more dorsally, first dorsal pore in 12/13, no genital markings, and intestine origin at xiv with manicate caeca (
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FC4EFA22FD44FA65" author="Thai, B. T. & Do, N. V. & Huynh, H. T. K." box="[537,720,1447,1473]" pageId="5" pageNumber="488" pagination="12 - 15" refId="ref6225" refString="Thai, B. T., Do, N. V. & Huynh, H. T. K. (1993) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae - Oligochaeta) from Yokdon, Dac Lac Province. Tap chi Sinh hoc, 15 (4), 12 - 15. [in Vietnamese]" type="journal article" year="1993">
|
||
Thai
|
||
<emphasis id="B91CEAB8FFA0FFF8FC02FA22FD12FA65" box="[597,646,1447,1473]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
|
||
. 1993
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736AAFFA0FFF8FE90FA4EFE77F8A1" blockId="5.[151,1437,151,1797]" pageId="5" pageNumber="488">
|
||
The presence of septum 8/9 was also recorded in
|
||
<taxonomicName id="4C684D29FFA0FFF8FCA7FA4EFBACFA42" authority="(Vaillant, 1868)" baseAuthorityName="Vaillant" baseAuthorityYear="1868" box="[752,1080,1483,1510]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="posthuma">
|
||
<emphasis id="B91CEAB8FFA0FFF8FCA7FA4EFC16FA41" box="[752,898,1483,1509]" italics="true" pageId="5" pageNumber="488">M. posthuma</emphasis>
|
||
(Vaillant, 1868)
|
||
</taxonomicName>
|
||
re-described by
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FAA3FA4EFE93F9AD" author="Bantaowong, U. & Chanabun, R. & Tongkerd, P. & Sutcharit, C. & James, S. & Panha, S." pageId="5" pageNumber="488" pagination="55 - 69" refId="ref4607" refString="Bantaowong, U., Chanabun, R., Tongkerd, P., Sutcharit, C., James, S. & Panha, S. (2011) A new species of the terrestrial earthworm of the genus Metaphire Sims and Easton, 1972 from Thailand with redescription of some species. Tropical Natural History, 11 (1), 55 - 69. [https: // li 01. tci-thaijo. org / index. php / tnh / article / view / 103012]" type="journal article" year="2011">
|
||
Bantaowong
|
||
<emphasis id="B91CEAB8FFA0FFF8FBDFFA4EFF39F9AD" italics="true" pageId="5" pageNumber="488">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C684D29FFA0FFF8FF44FA6AFDE2F9AD" authority="(Thai, 1982)" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[275,630,1519,1545]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="perelae">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF44FA6AFE75F9AD" box="[275,481,1519,1545]" italics="true" pageId="5" pageNumber="488">Metaphire perelae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FFA7FA6AFDFAF9AD" author="Thai, T. B." box="[496,622,1519,1545]" pageId="5" pageNumber="488" pagination="817 - 830" refId="ref6290" refString="Thai, T. B. (1982) The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Jurnal, 61 (6), 817 - 830. [in Russian]" type="journal article" year="1982">Thai, 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D29FFA0FFF8FCD5FA6AFC72F9AD" authority="(Thai, 1984)" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[642,998,1519,1545]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="truongsonensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FCD5FA6AFCC5F9AC" box="[642,849,1519,1544]" italics="true" pageId="5" pageNumber="488">M. truongsonensis</emphasis>
|
||
(Thai, 1984)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C684D29FFA0FFF8FA75FA6AFE1CF989" authority="Nguyen, Nguyen, Lam & Nguyen, 2020" authorityName="Nguyen, Nguyen, Lam & Nguyen" authorityYear="2020" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="songbeensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FA75FA6AFB5AF9AD" box="[1058,1230,1519,1545]" italics="true" pageId="5" pageNumber="488">M. songbeensis</emphasis>
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FA81FA6AFE1CF989" author="Nguyen, T. T. & Nguyen, N. Q. & Lam, D. H. & Nguyen, A. D." pageId="5" pageNumber="488" pagination="220 - 236" refId="ref5811" refString="Nguyen, T. T., Nguyen, N. Q., Lam, D. H. & Nguyen, A. D. (2020 b) Six new species of the genus Metaphire Sims & Easton, 1972 (Annelida: Oligochaeta: Megascolecidae) from southeastern Vietnam. Raffles Bulletin of Zoology, 68, 220 - 236. https: // doi. org / 10.26107 / RBZ- 2020 - 0019" type="journal article" year="2020">Nguyen, Nguyen, Lam & Nguyen, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FFC2F996FD9FF989" author="Thai, T. B." box="[405,523,1555,1581]" pageId="5" pageNumber="488" pagination="817 - 830" refId="ref6290" refString="Thai, T. B. (1982) The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Jurnal, 61 (6), 817 - 830. [in Russian]" type="journal article" year="1982">Thai 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FC43F996FD0AF989" author="Thai, T. B." box="[532,670,1555,1581]" pageId="5" pageNumber="488" pagination="284 - 288" refId="ref6379" refString="Thai, T. B. (1984 b) New data on taxonomy of the genus Pheretima (Oligochaeta, Megascolecidae) of the fauna of Vietnam. Zoologichesky Zhurnal, 63 (2), 284 - 288. [in Russian]" type="journal article" year="1984">Thai, 1984b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF94B5BFFA0FFF8FCF0F996FC18F98A" author="Nguyen, T. T. & Nguyen, N. Q. & Lam, D. H. & Nguyen, A. D." box="[679,908,1555,1582]" pageId="5" pageNumber="488" pagination="220 - 236" refId="ref5811" refString="Nguyen, T. T., Nguyen, N. Q., Lam, D. H. & Nguyen, A. D. (2020 b) Six new species of the genus Metaphire Sims & Easton, 1972 (Annelida: Oligochaeta: Megascolecidae) from southeastern Vietnam. Raffles Bulletin of Zoology, 68, 220 - 236. https: // doi. org / 10.26107 / RBZ- 2020 - 0019" type="journal article" year="2020">
|
||
Nguyen
|
||
<emphasis id="B91CEAB8FFA0FFF8FD52F996FCA1F989" box="[773,821,1555,1581]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
|
||
. 2020b
|
||
</bibRefCitation>
|
||
). The new species is clearly different from these species in having two pairs of spermathecal pores in ventrally 5/6/7 (vs. laterally 6/7/
|
||
<quantity id="4C909B4FFFA0FFF8FA11F9B2FBFBF9F5" box="[1094,1135,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="488" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName id="4C684D29FFA0FFF8FA22F9B2FB78F9F5" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[1141,1260,1591,1617]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="perelae">
|
||
<emphasis id="B91CEAB8FFA0FFF8FA22F9B2FB78F9F5" box="[1141,1260,1591,1617]" italics="true" pageId="5" pageNumber="488">M. perelae</emphasis>
|
||
</taxonomicName>
|
||
, ventrolaterally 5/6/7/8/
|
||
<quantity id="4C909B4FFFA0FFF8FEB8F9DFFE8FF9D1" box="[239,283,1626,1653]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="5" pageNumber="488" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="4C684D29FFA0FFF8FF73F9DEFE2CF9D1" baseAuthorityName="Vaillant" baseAuthorityYear="1868" box="[292,440,1627,1653]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="posthuma">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF73F9DEFE2CF9D1" box="[292,440,1627,1653]" italics="true" pageId="5" pageNumber="488">M. posthuma</emphasis>
|
||
</taxonomicName>
|
||
, and dorsally 5/6/7/8/
|
||
<quantity id="4C909B4FFFA0FFF8FCE2F9DFFD75F9D1" box="[693,737,1626,1653]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="5" pageNumber="488" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="4C684D29FFA0FFF8FCBDF9DEFC2FF9D0" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[746,955,1627,1652]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="truongsonensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FCBDF9DEFC2FF9D0" box="[746,955,1627,1652]" italics="true" pageId="5" pageNumber="488">M. truongsonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C684D29FFA0FFF8FDA2F9DEFB30F9D1" authorityName="Nguyen, Nguyen, Lam & Nguyen" authorityYear="2020" box="[1013,1188,1627,1653]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="songbeensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDA2F9DEFB30F9D1" box="[1013,1188,1627,1653]" italics="true" pageId="5" pageNumber="488">M. songbeensis</emphasis>
|
||
</taxonomicName>
|
||
), two pairs of genital markings in xviii (vs. paired but not in xviii (
|
||
<taxonomicName id="4C684D29FFA0FFF8FCFCF9FAFCD4F93D" baseAuthorityName="Vaillant" baseAuthorityYear="1868" box="[683,832,1663,1689]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="posthuma">
|
||
<emphasis id="B91CEAB8FFA0FFF8FCFCF9FAFCD4F93D" box="[683,832,1663,1689]" italics="true" pageId="5" pageNumber="488">M. posthuma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C684D29FFA0FFF8FD2BF9FAFBBFF93D" authorityName="Nguyen, Nguyen, Lam & Nguyen" authorityYear="2020" box="[892,1067,1663,1689]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="songbeensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FD2BF9FAFBBFF93D" box="[892,1067,1663,1689]" italics="true" pageId="5" pageNumber="488">M. songbeensis</emphasis>
|
||
</taxonomicName>
|
||
), absent in the male region (
|
||
<taxonomicName id="4C684D29FFA0FFF8FB29F9FAFF7EF919" baseAuthorityName="Thai" baseAuthorityYear="1982" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="perelae">
|
||
<emphasis id="B91CEAB8FFA0FFF8FB29F9FAFF7EF919" italics="true" pageId="5" pageNumber="488">M. perelae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C684D29FFA0FFF8FF74F926FE60F918" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[291,500,1699,1724]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="truongsonensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF74F926FE60F918" box="[291,500,1699,1724]" italics="true" pageId="5" pageNumber="488">M. truongsonensis</emphasis>
|
||
</taxonomicName>
|
||
)). In addition, the new species also differs from
|
||
<taxonomicName id="4C684D29FFA0FFF8FA75F926FB22F919" baseAuthorityName="Vaillant" baseAuthorityYear="1868" box="[1058,1206,1699,1725]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="posthuma">
|
||
<emphasis id="B91CEAB8FFA0FFF8FA75F926FB22F919" box="[1058,1206,1699,1725]" italics="true" pageId="5" pageNumber="488">M. posthuma</emphasis>
|
||
</taxonomicName>
|
||
in prostomium
|
||
<typeStatus id="54D38808FFA0FFF8FB3BF926FA08F919" box="[1388,1436,1699,1725]" pageId="5" pageNumber="488">type</typeStatus>
|
||
(epilobous vs. prolobous),
|
||
<taxonomicName id="4C684D29FFA0FFF8FF9CF942FDD3F945" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[459,583,1735,1761]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="perelae">
|
||
<emphasis id="B91CEAB8FFA0FFF8FF9CF942FDD3F945" box="[459,583,1735,1761]" italics="true" pageId="5" pageNumber="488">M. perelae</emphasis>
|
||
</taxonomicName>
|
||
in caeca (simple vs serrated),
|
||
<taxonomicName id="4C684D29FFA0FFF8FDFBF942FBCFF945" authorityName="Nguyen, Nguyen, Lam & Nguyen" authorityYear="2020" box="[940,1115,1735,1761]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="488" phylum="Annelida" rank="species" species="songbeensis">
|
||
<emphasis id="B91CEAB8FFA0FFF8FDFBF942FBCFF945" box="[940,1115,1735,1761]" italics="true" pageId="5" pageNumber="488">M. songbeensis</emphasis>
|
||
</taxonomicName>
|
||
in the state of septum 9/10 (absent vs. present) (
|
||
<tableCitation id="C6EA0311FFA0FFF8FF28F96EFE42F8A1" box="[383,470,1771,1797]" captionStart="TABLE 2" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 2. Character comparison between M. giengensis sp. nov., M. perelae, M. posthuma, M. truongsonensis, and M. songbeensis" httpUri="http://table.plazi.org/id/DF176622FFA2FFFAFEC0FF12FE8CFF77" pageId="5" pageNumber="488" tableUuid="DF176622FFA2FFFAFEC0FF12FE8CFF77">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |