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<document id="2FB081519FAE779A1AD685E0BA740154" ID-DOI="10.11646/zootaxa.3478.1.32" ID-GBIF-Dataset="f3e4ca1e-a40b-465a-a900-7a898e472cde" ID-ISSN="1175-5326" ID-Zenodo-Dep="282268" ID-ZooBank="FC92E481-4FF7-4DD8-B7C9-9F192F373D2E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524755954" checkinUser="plazi" docAuthor="Aarvik, Leif, Karisch, Timm &amp; Marthinsen, Gunnhild" docDate="2012" docId="03C13418FFEEFF911AE7FC50388AFA67" docLanguage="en" docName="zt03478p372.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dracontogena tonitrualis Meyrick 1934" docType="treatment" docVersion="13" lastPageNumber="356" masterDocId="FFF84C60FFE5FF9A1A70FFA73C2AFFB5" masterDocTitle="Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species" masterLastPageNumber="372" masterPageNumber="345" pageNumber="356" updateTime="1698315014682" updateUser="plazi">
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<mods:title id="CA83A4E78606F007D54E022B68541FE0">Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species</mods:title>
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<paragraph id="8BD7850EFFEEFF911AE7FC503EBCFBA4" blockId="11.[151,662,1015,1074]" box="[151,662,1015,1041]" pageId="11" pageNumber="356">
<heading id="D09F3262FFEEFF911AE7FC503EBCFBA4" bold="true" box="[151,662,1015,1041]" fontSize="11" level="1" pageId="11" pageNumber="356" reason="1">
<taxonomicName id="4C68FE8DFFEEFF911AE7FC503EBCFBA4" ID-CoL="94S3P" authority="Meyrick, 1934" authorityName="Meyrick" authorityYear="1934" box="[151,662,1015,1041]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitrualis">
<emphasis id="B91C591CFFEEFF911AE7FC503EBCFBA4" bold="true" box="[151,662,1015,1041]" pageId="11" pageNumber="356">
<emphasis id="B91C591CFFEEFF911AE7FC503DE3FBA4" bold="true" box="[151,457,1015,1041]" italics="true" pageId="11" pageNumber="356">Dracontogena tonitrualis</emphasis>
(
<bibRefCitation id="EFF9F8FFFFEEFF911BA7FC503EA4FBA4" author="Meyrick, E." box="[471,654,1015,1041]" firstAuthor="Meyrick" journalOrPublisher="The Annals and Magazine of Natural History, Series 10" pageId="11" pageNumber="356" pagination="401 - 411" part="14" refId="ref14786" refString="Meyrick, E. (1934) The Percy Sladen and Godman Trusts Expedition to the Islands in the Gulf of Guinea, October 1932 - March, 1933. - III. Micro-Lepidoptera. The Annals and Magazine of Natural History, Series 10, 14, 401 - 411." title="The Percy Sladen and Godman Trusts Expedition to the Islands in the Gulf of Guinea, October 1932 - March, 1933. - III. Micro-Lepidoptera" type="journal article" year="1934">Meyrick, 1934</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C372D685FFEEFF911AE7FBBD3CD3FB87" box="[151,249,1050,1074]" pageId="11" pageNumber="356" type="description">
<paragraph id="8BD7850EFFEEFF911AE7FBBD3CD3FB87" blockId="11.[151,662,1015,1074]" box="[151,249,1050,1074]" pageId="11" pageNumber="356">
(
<figureCitation id="1353998BFFEEFF911AEFFBBD3CDBFB87" box="[159,241,1050,1074]" captionStart="FIGURES 41 46" captionStartId="15.[151,264,1745,1768]" captionTargetBox="[168,1435,469,1734]" captionTargetId="figure@15.[151,1436,469,1757]" captionTargetPageId="15" captionText="FIGURES 41 46. Male genitalia in Dracontogena. 41. D. bernardi Karisch. (slide L. Aarvik 2808). 42. D. bernardi Karisch, phallus (slide L. Aarvik 2808). 43. D. solii n. sp. (slide NHMO 2010). 44. D. solii n. sp. phallus (slide NHMO 2010). 45. D. tonitrualis (Meyrick). 46. D. metamorphica (Meyrick). Scale figs. 41, 43: 1 mm., figs. 42, 44: 0.5 mm. Figs. 45 46 from Karisch (2005)." httpUri="https://zenodo.org/record/282275/files/figure.png" pageId="11" pageNumber="356">Fig. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEEFF911AE7FBC43818FB00" pageId="11" pageNumber="356" type="reference_group">
<paragraph id="8BD7850EFFEEFF911AE7FBC43E13FBCD" blockId="11.[151,1074,1122,1206]" box="[151,569,1122,1144]" pageId="11" pageNumber="356">
<treatmentCitationGroup id="AB78A220FFEEFF911AE7FBC43E13FBCD" box="[151,569,1122,1144]" pageId="11" pageNumber="356">
<treatmentCitation id="0AC9A31FFFEEFF911AE7FBC43E1FFBCD" author="Meyrick" box="[151,565,1122,1144]" page="405" pageId="11" pageNumber="356" year="1934">
<taxonomicName id="4C68FE8DFFEEFF911AE7FBC43E1FFBCD" ID-CoL="6GV9X" authority="Meyrick, 1934: 405" authorityName="Meyrick" authorityPageNumber="405" authorityYear="1934" box="[151,565,1122,1144]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitrualis">
<emphasis id="B91C591CFFEEFF911AE7FBC43D49FBCD" box="[151,355,1122,1144]" italics="true" pageId="11" pageNumber="356">Eucosma tonitrualis</emphasis>
<bibRefCitation id="EFF9F8FFFFEEFF911B19FBC53E1FFBCD" author="Meyrick, E." box="[361,565,1122,1144]" firstAuthor="Meyrick" journalOrPublisher="The Annals and Magazine of Natural History, Series 10" page="405" pageId="11" pageNumber="356" pagination="401 - 411" part="14" refId="ref14786" refString="Meyrick, E. (1934) The Percy Sladen and Godman Trusts Expedition to the Islands in the Gulf of Guinea, October 1932 - March, 1933. - III. Micro-Lepidoptera. The Annals and Magazine of Natural History, Series 10, 14, 401 - 411." title="The Percy Sladen and Godman Trusts Expedition to the Islands in the Gulf of Guinea, October 1932 - March, 1933. - III. Micro-Lepidoptera" type="journal article" year="1934">Meyrick, 1934: 405</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BD7850EFFEEFF911AE7FB253FB6FB22" blockId="11.[151,1074,1122,1206]" box="[151,924,1153,1175]" pageId="11" pageNumber="356">
<treatmentCitationGroup id="AB78A220FFEEFF911AE7FB253FB6FB22" box="[151,924,1153,1175]" pageId="11" pageNumber="356">
<taxonomicName id="4C68FE8DFFEEFF911AE7FB253D4EFB22" box="[151,356,1153,1175]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitraulis">
<emphasis id="B91C591CFFEEFF911AE7FB253D4EFB22" box="[151,356,1153,1175]" italics="true" pageId="11" pageNumber="356">Eucosma tonitraulis</emphasis>
</taxonomicName>
(misspelling),
<treatmentCitation id="0AC9A31FFFEEFF911B8CFB263E84FB22" author="Clarke" box="[508,686,1153,1175]" page="392" pageId="11" pageNumber="356" year="1958">
<bibRefCitation id="EFF9F8FFFFEEFF911B8CFB263E84FB22" author="Clarke, J. F. G." box="[508,686,1153,1175]" firstAuthor="Clarke" journalOrPublisher="Tortricidae, Olethreutidae, Noctuidae" page="392" pageId="11" pageNumber="356" pagination="1 - 600" refId="ref14308" refString="Clarke, J. F. G. (1958) Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. III. Tortricidae, Olethreutidae, Noctuidae, 1 - 600." title="Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. III" type="book chapter" year="1958">Clarke 1958: 392</bibRefCitation>
</treatmentCitation>
, pl. 195, figs. 2, 2a, 2b.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD7850EFFEEFF911AE7FB063818FB00" blockId="11.[151,1074,1122,1206]" box="[151,1074,1184,1206]" pageId="11" pageNumber="356">
<treatmentCitationGroup id="AB78A220FFEEFF911AE7FB063818FB00" box="[151,1074,1184,1206]" pageId="11" pageNumber="356">
<taxonomicName id="4C68FE8DFFEEFF911AE7FB063E57FB00" ID-CoL="94S3P" authority="Diakonoff 1970: 124" authorityName="Diakonoff" authorityPageNumber="124" authorityYear="1970" box="[151,637,1184,1206]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitrualis">
<emphasis id="B91C591CFFEEFF911AE7FB063DB5FB03" box="[151,415,1184,1206]" italics="true" pageId="11" pageNumber="356">Dracontogena tonitrualis,</emphasis>
<treatmentCitation id="0AC9A31FFFEEFF911BD5FB073E57FB00" author="Diakonoff" box="[421,637,1184,1206]" page="124" pageId="11" pageNumber="356" year="1970">
<bibRefCitation id="EFF9F8FFFFEEFF911BD5FB073E57FB00" author="Diakonoff, A." box="[421,637,1184,1206]" firstAuthor="Diakonoff" journalOrPublisher="Memoires O. R. S. T. O. M." page="124" pageId="11" pageNumber="356" pagination="103 - 150" part="37" refId="ref14353" refString="Diakonoff, A. (1970) Lepidoptera Tortricoidea from Tsaratanana range. Memoires O. R. S. T. O. M., 37, 103 - 150." title="Lepidoptera Tortricoidea from Tsaratanana range" type="journal article" year="1970">Diakonoff 1970: 124</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AC9A31FFFEEFF9118F7FB073F6EFB03" author="Karisch" box="[647,836,1184,1206]" page="461" pageId="11" pageNumber="356" year="2005">
<bibRefCitation id="EFF9F8FFFFEEFF9118F7FB073F6EFB03" author="Karisch, T." box="[647,836,1184,1206]" firstAuthor="Karisch" journalOrPublisher="Linzer biologische Beitrage" page="461" pageId="11" pageNumber="356" pagination="457 - 476" part="37" refId="ref14694" refString="Karisch, T. (2005) Ubersicht uber die Gattung Dracontogena Diakonoff 1970 (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini). Linzer biologische Beitrage, 37, 457 - 476." title="Ubersicht uber die Gattung Dracontogena Diakonoff 1970 (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini)" type="journal article" year="2005">Karisch 2005: 461</bibRefCitation>
</treatmentCitation>
, text fig. 9, pl. 1, fig. 9.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEEFF911AE7FB453DB5FAAB" pageId="11" pageNumber="356" type="materials_examined">
<paragraph id="8BD7850EFFEEFF911AE7FB453DB5FAAB" blockId="11.[151,1436,1250,1490]" pageId="11" pageNumber="356">
Material examined.
<materialsCitation id="3B008F53FFEEFF911BF4FB453DB5FAAB" ID-GBIF-Occurrence="3712346307" collectingDate="1932-11-05" collectionCode="BMNH" collectorName="W. H. T. Tams" country="Sao Tome and Principe" location="Sao Tome" pageId="11" pageNumber="356" specimenCode="BMNH 7012" specimenCount="1">
Holotype of
<taxonomicName id="4C68FE8DFFEEFF91186DFB433F2CFB4E" box="[541,774,1251,1275]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitrualis">
<emphasis id="B91C591CFFEEFF91186DFB433F2CFB4E" box="[541,774,1251,1275]" italics="true" pageId="11" pageNumber="356">
<originalDetermination id="A8763DF2FFEEFF91186DFB433F2CFB4E" box="[541,774,1251,1275]" pageId="11" pageNumber="356">Eucosma tonitrualis</originalDetermination>
</emphasis>
</taxonomicName>
, 3,
<collectingCountry id="F37FC59EFFEEFF911933FB453F97FB4F" box="[835,957,1250,1274]" name="Sao Tome and Principe" pageId="11" pageNumber="356">Sao Tomé</collectingCountry>
,
<date id="FFD6A3CEFFEEFF9119BEFB453810FB4F" box="[974,1082,1250,1274]" pageId="11" pageNumber="356" value="1932-11-05">
<collectingDate id="EF925A26FFEEFF9119BEFB453810FB4F" box="[974,1082,1250,1274]" pageId="11" pageNumber="356" value="1932-11-05">5.xi.1932</collectingDate>
</date>
,
<collectorName id="269DE0D8FFEEFF911E3CFB4438C2FB4F" box="[1100,1256,1251,1274]" pageId="11" pageNumber="356">W.H.T. Tams</collectorName>
, genitalia slide
<specimenCode id="DBCE2D75FFEEFF911AE7FAA03D00FAAB" box="[151,298,1286,1310]" pageId="11" pageNumber="356">BMNH 7012</specimenCode>
(
<collectionCode id="ED791DCBFFEEFF911B48FAA13DB9FAAB" box="[312,403,1286,1310]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="11" pageNumber="356">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
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<subSubSection id="C372D685FFEEFF911AB7FA8D3881FA1B" pageId="11" pageNumber="356" type="diagnosis">
<paragraph id="8BD7850EFFEEFF911AB7FA8D3881FA1B" blockId="11.[151,1436,1250,1490]" pageId="11" pageNumber="356">
Diagnosis. Wingspan 17.0 mm. Together with
<taxonomicName id="4C68FE8DFFEEFF9118A4FA8B3F5CFAF6" box="[724,886,1323,1347]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="niphadonta">
<emphasis id="B91C591CFFEEFF9118A4FA8B3F5CFAF6" box="[724,886,1323,1347]" italics="true" pageId="11" pageNumber="356">D. niphadonta</emphasis>
</taxonomicName>
characterised externally by the combination of a strongly notched hindwing and a patch of black sex scales. The other African species with a patch of black sex scales on the hindwing upperside,
<taxonomicName id="4C68FE8DFFEEFF911866FAD33E42FA3E" box="[534,616,1395,1419]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="solii" status="sp. nov.">
<emphasis id="B91C591CFFEEFF911866FAD33E42FA3E" box="[534,616,1395,1419]" italics="true" pageId="11" pageNumber="356">D. solii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FE467FFEEFF91181EFAD43E86FA3F" box="[622,684,1395,1418]" pageId="11" pageNumber="356" rank="species">n. sp.</taxonomicNameLabel>
, has the hindwing termen only slightly concave. The male genitalia of
<taxonomicName id="4C68FE8DFFEEFF911AC5FA3F3D61FA1B" box="[181,331,1430,1455]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="tonitrualis">
<emphasis id="B91C591CFFEEFF911AC5FA3F3CE3FA1A" box="[181,201,1432,1455]" italics="true" pageId="11" pageNumber="356">D</emphasis>
.
<emphasis id="B91C591CFFEEFF911AA7FA313D61FA1B" box="[215,331,1430,1454]" italics="true" pageId="11" pageNumber="356">tonitrualis</emphasis>
</taxonomicName>
differ from all other
<taxonomicName id="4C68FE8DFFEEFF911849FA3F3EF1FA1A" box="[569,731,1432,1455]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="B91C591CFFEEFF911849FA3F3EF1FA1A" box="[569,731,1432,1455]" italics="true" pageId="11" pageNumber="356">Dracontogena</emphasis>
</taxonomicName>
species by the slender cucullus (
<figureCitation id="1353998BFFEEFF911E3AFA3138B4FA1B" box="[1098,1182,1430,1454]" captionStart="FIGURES 41 46" captionStartId="15.[151,264,1745,1768]" captionTargetBox="[168,1435,469,1734]" captionTargetId="figure@15.[151,1436,469,1757]" captionTargetPageId="15" captionText="FIGURES 41 46. Male genitalia in Dracontogena. 41. D. bernardi Karisch. (slide L. Aarvik 2808). 42. D. bernardi Karisch, phallus (slide L. Aarvik 2808). 43. D. solii n. sp. (slide NHMO 2010). 44. D. solii n. sp. phallus (slide NHMO 2010). 45. D. tonitrualis (Meyrick). 46. D. metamorphica (Meyrick). Scale figs. 41, 43: 1 mm., figs. 42, 44: 0.5 mm. Figs. 45 46 from Karisch (2005)." httpUri="https://zenodo.org/record/282275/files/figure.png" pageId="11" pageNumber="356">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8BD7850EFFEEFF911AB7FA1D388AFA67" blockId="11.[151,1436,1250,1490]" box="[199,1184,1466,1490]" pageId="11" pageNumber="356">
Distribution. Known only from
<collectingCountry id="F37FC59EFFEEFF91185CFA1D3EB5FA67" box="[556,671,1466,1490]" name="Sao Tome and Principe" pageId="11" pageNumber="356">Sao Tomé</collectingCountry>
, an island in the Gulf of
<collectingCountry id="F37FC59EFFEEFF9119C4FA1D3822FA67" box="[948,1032,1466,1490]" name="Guinea-Bissau" pageId="11" pageNumber="356">Guinea</collectingCountry>
, West Africa.
</paragraph>
</subSubSection>
</treatment>
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