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<document id="63E5A910C759516CB78CBC5619F3FC1E" ID-CLB-Dataset="302486" ID-DOI="10.11646/phytotaxa.394.4.4" ID-GBIF-Dataset="36d8d2bb-d1b1-4550-b379-c6b16b430595" ID-ISSN="1179-3163" ID-Zenodo-Dep="13718420" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725654170130" checkinUser="felipe" docAuthor="Salmaki, Yasaman &amp; Müller, Jochen" docDate="2019" docId="03BD0133FFB0FFC0FF49FA83FB64A8F2" docLanguage="en" docName="phytotaxa.394.4.4.pdf" docOrigin="Phytotaxa 394 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.394.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Scutellaria xylorrhiza Bornmuller 1911" docType="treatment" docVersion="2" lastPageNumber="274" masterDocId="FF84794BFFB5FFC7FFC1FFF7FFB1A963" masterDocTitle="Rediscovery of the enigmatic Scutellaria xylorrhiza (Scutellarioideae; Lamiaceae) - a rare endemic species from Iran" masterLastPageNumber="275" masterPageNumber="267" pageNumber="272" updateTime="1725664772960" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="643555A5D602B36459F31614FEE5298E">Rediscovery of the enigmatic Scutellaria xylorrhiza (Scutellarioideae; Lamiaceae) - a rare endemic species from Iran</mods:title>
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<mods:namePart id="7D2AA7156B9F615E550674FB4BCE2AA0">Müller, Jochen</mods:namePart>
<mods:affiliation id="286B75581B686DD9A677C24B826438E0">Herbarium Haussknecht, Institut für Spezielle Botanik, Friedrich-Schiller-Universität, Fürstengraben 1, 07737 Jena, Germany. * Author for correspondence</mods:affiliation>
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<paragraph id="8BABB025FFB0FFC2FF49FA83FC8DACEC" blockId="5.[136,828,1396,1423]" box="[136,828,1396,1423]" pageId="5" pageNumber="272">
<heading id="D0E30749FFB0FFC2FF49FA83FC8DACEC" box="[136,828,1396,1423]" fontSize="11" level="2" pageId="5" pageNumber="272" reason="3">
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<treatmentCitation id="0AB59634FFB0FFC2FF49FA83FD37ACEC" author="Bornmuller, J." box="[136,646,1396,1423]" page=")" pageId="5" pageNumber="272" year="1911">
<taxonomicName id="4C14CBA6FFB0FFC2FF49FA83FD37ACEC" ID-CoL="6XYFG" authority="Bornmuller (1911 a: 7)" authorityName="Bornmuller" authorityPageNumber="7" authorityYear="1911" box="[136,646,1396,1423]" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="272" phylum="Tracheophyta" rank="species" species="xylorrhiza">
<emphasis id="B9606C37FFB0FFC2FF49FA83FE32ACEC" bold="true" box="[136,387,1396,1423]" italics="true" pageId="5" pageNumber="272">Scutellaria xylorrhiza</emphasis>
<bibRefCitation id="EF85CDD4FFB0FFC2FE4BFA81FD37ACEC" author="Bornmuller, J." box="[394,646,1398,1423]" pageId="5" pageNumber="272" pagination="7 - 8" refId="ref3469" refString="Bornmuller, J. (1911 a) Scutellaria xylorrhiza. Russkii Botanicheskii Zhurnal 1: 7 - 8." type="journal article" year="1911">Bornmüller (1911a: 7)</bibRefCitation>
</taxonomicName>
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(
<figureCitation id="132FACA0FFB0FFC2FD54FA81FD53ACEC" box="[661,738,1398,1423]" captionStart="FIGURE 1" captionStartId="3.[136,229,2007,2028]" captionTargetBox="[151,1429,193,1979]" captionTargetId="figure-14@3.[151,1436,187,1979]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Scutellaria xylorrhiza (JE00001284)." figureDoi="http://doi.org/10.5281/zenodo.13718422" httpUri="https://zenodo.org/record/13718422/files/figure.png" pageId="5" pageNumber="272">Figs. 1</figureCitation>
,
<figureCitation id="132FACA0FFB0FFC2FD31FA81FC82ACEC" box="[752,819,1398,1423]" captionStart="FIGURE 2" captionStartId="4.[136,229,1613,1634]" captionTargetBox="[208,1379,191,1585]" captionTargetId="figure-15@4.[208,1379,190,1588]" captionTargetPageId="4" captionText="FIGURE 2. (A, B) Scutellaria xylorrhiza in its natural habitat, (C) distribution map of S. xylorrhiza in Iran." figureDoi="http://doi.org/10.5281/zenodo.13718424" httpUri="https://zenodo.org/record/13718424/files/figure.png" pageId="5" pageNumber="272">2AB</figureCitation>
)
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<subSubSection id="C30EE3AEFFB0FFC2FF49FA6DFA12A174" pageId="5" pageNumber="272" type="materials_examined">
<paragraph id="8BABB025FFB0FFC2FF49FA6DFE82ACB4" blockId="5.[136,1452,1434,2071]" pageId="5" pageNumber="272">
Type (
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, designated here):—[
<collectingCountry id="F303F0B5FFB0FFC2FD87FA6DFDC6ACD0" box="[582,631,1434,1459]" name="Iran" pageId="5" pageNumber="272">Iran</collectingCountry>
] Kohrud,
<date id="FFAA96E5FFB0FFC2FD35FA6DFC28ACD0" box="[756,921,1434,1459]" pageId="5" pageNumber="272" value="1904-06-20">20 June 1904</date>
,
<emphasis id="B9606C37FFB0FFC2FC6AFA6DFB9EACD0" box="[939,1071,1434,1459]" italics="true" pageId="5" pageNumber="272">Th. Strauss</emphasis>
s.n. (
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00001284!,
<typeStatus id="54AF0E87FFB0FFC2FAE5FA6DFA1AACD0" box="[1316,1451,1434,1459]" pageId="5" pageNumber="272" type="isolectotype">isolectotype</typeStatus>
<collectionCode id="ED0528E0FFB0FFC2FF49FA49FF2BACB4" box="[136,154,1470,1495]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="5" pageNumber="272" type="Herbarium">B</collectionCode>
100241792!).
</paragraph>
<paragraph id="8BABB025FFB0FFC2FF7CFA15FEF1AEAC" blockId="5.[136,1452,1434,2071]" pageId="5" pageNumber="272">
Plants perennial, saxicolous, suffruticose herbs with woody base. Stems thin and fragile, numerous, erect to decumbent,
<quantity id="4CEC1DC0FFB0FFC2FED3F9F1FEC2AF7C" box="[274,371,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="5" pageNumber="272" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">410 cm</quantity>
tall, covered by papillate short simple hairs. Basal leaves long-petiolate; blade ovate to broadly ovate, 1.22.2 ×
<quantity id="4CEC1DC0FFB0FFC2FE8AF9DDFE08AF20" box="[331,441,1578,1603]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="5" pageNumber="272" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">12.5 cm</quantity>
, subcrenate to entire, ± obtuse at apex, cuneate to rarely rounded at base, upper surface sparsely covered by short curved papillate simple hairs (
<figureCitation id="132FACA0FFB0FFC2FCD3F9B9FCDFAF04" box="[786,878,1614,1639]" captionStart="FIGURE 3" captionStartId="5.[136,229,1218,1239]" captionTargetBox="[152,1436,191,1193]" captionTargetId="figure-14@5.[151,1436,190,1194]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electron micrographs of leaf and calyx of Scutellaria xylorrhiza. A: leaf; BD: calyx." figureDoi="http://doi.org/10.5281/zenodo.13718430" httpUri="https://zenodo.org/record/13718430/files/figure.png" pageId="5" pageNumber="272">Fig. 3A</figureCitation>
), lower surface rarely with peltate glandular hairs (
<figureCitation id="132FACA0FFB0FFC2FF51F985FF5DAFE8" box="[144,236,1650,1675]" captionStart="FIGURE 3" captionStartId="5.[136,229,1218,1239]" captionTargetBox="[152,1436,191,1193]" captionTargetId="figure-14@5.[151,1436,190,1194]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electron micrographs of leaf and calyx of Scutellaria xylorrhiza. A: leaf; BD: calyx." figureDoi="http://doi.org/10.5281/zenodo.13718430" httpUri="https://zenodo.org/record/13718430/files/figure.png" pageId="5" pageNumber="272">Fig. 3D</figureCitation>
); petiole length equal or subequal to that of the blade. Cauline leaves shortly petiolate; blade broadly ovate to ovate-rounded, 12 ×
<quantity id="4CEC1DC0FFB0FFC2FE5DF961FDADAFCC" box="[412,540,1686,1711]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.1" metricValueMin="0.9" pageId="5" pageNumber="272" unit="cm" value="1.5" valueMax="2.1" valueMin="0.9">0.92.1 cm</quantity>
, leaf margins and indumentum similar to the basal leaves (leaf margin sometimes dentate); petiole
<quantity id="4CEC1DC0FFB0FFC2FE89F94DFE03AFB0" box="[328,434,1722,1747]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="272" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">11.2 cm</quantity>
long. Inflorescence secund, compressed and terminal, dense,
<quantity id="4CEC1DC0FFB0FFC2FBB4F94DFB7BAFB0" box="[1141,1226,1722,1747]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="5" pageNumber="272" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">47 cm</quantity>
long, verticillasters 2-flowered. Bracts subsessile or with a short stalk (up to
<quantity id="4CEC1DC0FFB0FFC2FCE5F929FCD9AF94" box="[804,872,1758,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="272" unit="mm" value="1.0">1 mm</quantity>
long), ovate to lanceolate,
<quantity id="4CEC1DC0FFB0FFC2FB66F929FAB7AF94" box="[1191,1286,1758,1783]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="272" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long, covered by stalked glandular hairs and curved simple hairs. Pedicel
<quantity id="4CEC1DC0FFB0FFC2FCEFF8F5FC3DAE78" box="[814,908,1794,1819]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="272" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long. Calyx
<quantity id="4CEC1DC0FFB0FFC2FBE1F8F5FBCFAE78" box="[1056,1150,1794,1819]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="272" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, growing to
<quantity id="4CEC1DC0FFB0FFC2FA8AF8F5FA3FAE78" box="[1355,1422,1794,1819]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="272" unit="mm" value="5.0">5 mm</quantity>
in fruit, densely covered by exerted, long-stalked glandular hairs, sessile to subsessile peltate glandular hairs and curved papillate simple hairs (
<figureCitation id="132FACA0FFB0FFC2FE48F8BDFDB4AE00" box="[393,517,1866,1891]" captionStart="FIGURE 3" captionStartId="5.[136,229,1218,1239]" captionTargetBox="[152,1436,191,1193]" captionTargetId="figure-14@5.[151,1436,190,1194]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electron micrographs of leaf and calyx of Scutellaria xylorrhiza. A: leaf; BD: calyx." figureDoi="http://doi.org/10.5281/zenodo.13718430" httpUri="https://zenodo.org/record/13718430/files/figure.png" pageId="5" pageNumber="272">Fig. 3BD</figureCitation>
). Corolla
<quantity id="4CEC1DC0FFB0FFC2FDB4F8BDFD5EAE00" box="[629,751,1866,1891]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="5" pageNumber="272" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">1620 mm</quantity>
long, tubular, apically bilabiate, with hooded upper lip, violet with a large white spot on lower lip, covered by stalked and sessile glandular hairs and simple short hairs outside, glabrous inside; corolla tube curved at base. Mericarps ovate to elliptic in outline, covered with appressed grey hairs, c. 1.3 ×
<quantity id="4CEC1DC0FFB0FFC2FF22F841FE8DAEAC" box="[227,316,1974,1999]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="272" unit="mm" value="0.8">0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BABB025FFB0FFC2FF7CF82DFA12A174" blockId="5.[136,1452,1434,2071]" pageId="5" pageNumber="272">
<emphasis id="B9606C37FFB0FFC2FF7CF82DFE2FAE90" bold="true" box="[189,414,2010,2035]" pageId="5" pageNumber="272">Species examined:</emphasis>
<collectingCountry id="F303F0B5FFB0FFC2FE7DF82DFE5AAE90" box="[444,491,2010,2035]" name="Iran" pageId="5" pageNumber="272">Iran</collectingCountry>
. Prov.
<collectingRegion id="49D07EC7FFB0FFC2FD80F82DFD29AE90" box="[577,664,2010,2035]" country="Iran" name="Esfahan" pageId="5" pageNumber="272">Isfahan</collectingRegion>
: south of
<collectingRegion id="49D07EC7FFB0FFC2FCD7F82DFCDAAE90" box="[790,875,2010,2035]" country="Iran" name="Esfahan" pageId="5" pageNumber="272">Isfahan</collectingRegion>
, N-facing slopes, Sofeh Mountains,
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, 51°6465″E, ca.
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,
<date id="FFAA96E5FFB0FFC2FE63F809FDF9A174" box="[418,584,2046,2071]" pageId="5" pageNumber="272" value="2017-08-01">1 August 2017</date>
,
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<collectionCode id="ED0528E0FFB0FFC2FD92F809FDD5A174" box="[595,612,2046,2071]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="5" pageNumber="272" type="Herbarium">Y</collectionCode>
. Salmaki &amp;
<collectionCode id="ED0528E0FFB0FFC2FD2DF809FD4CA174" box="[748,765,2046,2071]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="5" pageNumber="272" type="Herbarium">S</collectionCode>
. Zarre
</emphasis>
45417 (
<collectionCode id="ED0528E0FFB0FFC2FC62F809FC57A174" box="[931,998,2046,2071]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13098" lsid="urn:lsid:biocol.org:col:13098" name="Tehran University" pageId="5" pageNumber="272" type="Herbarium">TUH</collectionCode>
!),
<emphasis id="B9606C37FFB0FFC2FC3FF809FB43A174" box="[1022,1266,2046,2071]" italics="true" pageId="5" pageNumber="272">
<collectionCode id="ED0528E0FFB0FFC2FC3FF809FBBEA174" box="[1022,1039,2046,2071]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="5" pageNumber="272" type="Herbarium">Y</collectionCode>
. Salmaki &amp;
<collectionCode id="ED0528E0FFB0FFC2FB56F809FB19A174" box="[1175,1192,2046,2071]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="5" pageNumber="272" type="Herbarium">S</collectionCode>
. Zarre
</emphasis>
45418 (
<collectionCode id="ED0528E0FFB0FFC2FA8EF809FA22A174" box="[1359,1427,2046,2071]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13098" lsid="urn:lsid:biocol.org:col:13098" name="Tehran University" pageId="5" pageNumber="272" type="Herbarium">TUH</collectionCode>
!).
</paragraph>
</subSubSection>
<caption id="DF6BE0ADFFB3FFC1FF49FA5BFB5DAC86" ID-DOI="http://doi.org/10.5281/zenodo.13718432" ID-Zenodo-Dep="13718432" httpUri="https://zenodo.org/record/13718432/files/figure.png" pageId="6" pageNumber="273" startId="6.[136,229,1452,1473]" targetBox="[206,1381,312,1364]" targetPageId="6" targetType="figure">
<paragraph id="8BABB025FFB3FFC1FF49FA5BFB5DAC86" blockId="6.[136,1452,1452,1509]" pageId="6" pageNumber="273">
<emphasis id="B9606C37FFB3FFC1FF49FA5BFF4FACA2" bold="true" box="[136,254,1452,1473]" pageId="6" pageNumber="273">FIGURE 4.</emphasis>
Majority rule consensus tree inferred from Bayesian analysis of nrDNA ITS sequences showing the phylogenetic placement of
<taxonomicName id="4C14CBA6FFB3FFC1FF63FA27FEC2AC86" authorityName="Bornmuller" authorityYear="1911" box="[162,371,1488,1509]" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="273" phylum="Tracheophyta" rank="species" species="xylorrhiza">
<emphasis id="B9606C37FFB3FFC1FF63FA27FEC2AC86" box="[162,371,1488,1509]" italics="true" pageId="6" pageNumber="273">Scutellaria xylorrhiza</emphasis>
</taxonomicName>
. Posterior probability and bootstrap support (PP/BS) values are reported above at the nodes.
</paragraph>
</caption>
<subSubSection id="C30EE3AEFFB3FFC0FF7CF9E2FB64A8F2" lastPageId="7" lastPageNumber="274" pageId="6" pageNumber="273" type="distribution">
<paragraph id="8BABB025FFB3FFC1FF7CF9E2FCB8AF15" blockId="6.[136,1452,1557,2086]" pageId="6" pageNumber="273">
<emphasis id="B9606C37FFB3FFC1FF7CF9E2FE49AF4D" bold="true" box="[189,504,1557,1582]" pageId="6" pageNumber="273">Distribution and ecology:</emphasis>
<taxonomicName id="4C14CBA6FFB3FFC1FDD5F9E2FCBFAF4D" authorityName="Bornmuller" authorityYear="1911" box="[532,782,1557,1582]" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="273" phylum="Tracheophyta" rank="species" species="xylorrhiza">
<emphasis id="B9606C37FFB3FFC1FDD5F9E2FCBFAF4D" box="[532,782,1557,1582]" italics="true" pageId="6" pageNumber="273">Scutellaria xylorrhiza</emphasis>
</taxonomicName>
is found on bare rocks at N-facing slopes of the Sofeh Mountains south of
<collectingRegion id="49D07EC7FFB3FFC1FEAFF9CEFE72AF31" box="[366,451,1593,1618]" country="Iran" name="Esfahan" pageId="6" pageNumber="273">Isfahan</collectingRegion>
, a province capital in the central part of
<collectingCountry id="F303F0B5FFB3FFC1FC59F9CEFC74AF31" box="[920,965,1593,1618]" name="Iran" pageId="6" pageNumber="273">Iran</collectingCountry>
(
<figureCitation id="132FACA0FFB3FFC1FC17F9CEFB80AF31" box="[982,1073,1593,1618]" captionStart="FIGURE 2" captionStartId="4.[136,229,1613,1634]" captionTargetBox="[208,1379,191,1585]" captionTargetId="figure-15@4.[208,1379,190,1588]" captionTargetPageId="4" captionText="FIGURE 2. (A, B) Scutellaria xylorrhiza in its natural habitat, (C) distribution map of S. xylorrhiza in Iran." figureDoi="http://doi.org/10.5281/zenodo.13718424" httpUri="https://zenodo.org/record/13718424/files/figure.png" pageId="6" pageNumber="273">Fig. 2C</figureCitation>
). The small and cushion-forming subalpine species grows at an elevation of
<quantity id="4CEC1DC0FFB3FFC1FDA7F9AAFCB5AF15" box="[614,772,1629,1654]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="273" unit="m" value="2000.0" valueMax="2200.0" valueMin="1800.0">18002200 m</quantity>
.
</paragraph>
<paragraph id="8BABB025FFB3FFC1FF7CF976FAD7AFF9" blockId="6.[136,1452,1557,2086]" box="[189,1382,1665,1690]" pageId="6" pageNumber="273">
<emphasis id="B9606C37FFB3FFC1FF7CF976FEF2AFF9" bold="true" box="[189,323,1665,1690]" pageId="6" pageNumber="273">Phenology:</emphasis>
—Flowering between late June and early August and fruiting between middle and late August.
</paragraph>
<paragraph id="8BABB025FFB3FFC1FF7CF952FE92AEF5" blockId="6.[136,1452,1557,2086]" pageId="6" pageNumber="273">
<emphasis id="B9606C37FFB3FFC1FF7CF952FE65AFDD" bold="true" box="[189,468,1701,1726]" pageId="6" pageNumber="273">Conservation remarks:</emphasis>
<taxonomicName id="4C14CBA6FFB3FFC1FE31F952FD56AFDD" authorityName="Bornmuller" authorityYear="1911" box="[496,743,1701,1726]" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="273" phylum="Tracheophyta" rank="species" species="xylorrhiza">
<emphasis id="B9606C37FFB3FFC1FE31F952FD56AFDD" box="[496,743,1701,1726]" italics="true" pageId="6" pageNumber="273">Scutellaria xylorrhiza</emphasis>
</taxonomicName>
can be considered as Critically Endangered (CR) subcategory A1 according to the IUCN Red List (
<bibRefCitation id="EF85CDD4FFB3FFC1FDE7F93EFD1CAF81" author="IUCN" box="[550,685,1737,1762]" pageId="6" pageNumber="273" refId="ref3748" refString="IUCN. (2010) The IUCN red list of threatened species, version 2010.4. IUCN Red List Unit, Cambridge U. K. Available from: http: // www. iucnredlist. org / (accessed 19 May 2011)" type="url" year="2010">IUCN 2010</bibRefCitation>
) categories. It is known only from an area which is not very close to the
<typeStatus id="54AF0E87FFB3FFC1FF73F91AFF53AE65" box="[178,226,1773,1798]" pageId="6" pageNumber="273">type</typeStatus>
locality (Barzuk Mountain near Kohrud) in
<collectingRegion id="49D07EC7FFB3FFC1FD19F91AFC28AE65" box="[728,921,1773,1798]" country="Iran" name="Esfahan" pageId="6" pageNumber="273">Isfahan Province</collectingRegion>
. We found about 70 individuals of this plant in the single locality, but we presume there should be also populations in the
<typeStatus id="54AF0E87FFB3FFC1FC13F8E6FBB3AE49" box="[978,1026,1809,1834]" pageId="6" pageNumber="273">type</typeStatus>
locality which are not easy to access. As there are no additional specimens except those that were collected by Strauss and Stapf in 1904,
<bibRefCitation id="EF85CDD4FFB3FFC1FAC5F8C2FA1DAE2D" author="Jamzad, Z." box="[1284,1452,1845,1870]" pageId="6" pageNumber="273" pagination="253 - 337" refId="ref3792" refString="Jamzad, Z. (2012) Eremostachys, and Phlomis. In: Assadi, M., Maassoumi, A. A. &amp; Mozaffarian, V. (Eds.) Flora of Iran 76: 253 - 337." type="book chapter" year="2012">Jamzad (2012)</bibRefCitation>
considered this species as presumably extinct because the distribution area of this species is heavily disturbed by human usage.
</paragraph>
<paragraph id="8BABB025FFB3FFC1FF7CF856FBFAA145" blockId="6.[136,1452,1557,2086]" pageId="6" pageNumber="273">
<emphasis id="B9606C37FFB3FFC1FF7CF856FD30AED9" bold="true" box="[189,641,1951,1978]" pageId="6" pageNumber="273">
Typification of
<taxonomicName id="4C14CBA6FFB3FFC1FEB8F868FDC6AED9" authorityName="Bornmuller" authorityYear="1911" box="[377,631,1951,1978]" class="Magnoliopsida" family="Lamiaceae" genus="Scutellaria" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="273" phylum="Tracheophyta" rank="species" species="xylorrhiza">
<emphasis id="B9606C37FFB3FFC1FEB8F868FDC6AED9" bold="true" box="[377,631,1951,1978]" italics="true" pageId="6" pageNumber="273">Scutellaria xylorrhiza</emphasis>
</taxonomicName>
:
</emphasis>
—The name was published in the journal named Russkij Botaniceskij Zurnal (
<bibRefCitation id="EF85CDD4FFB3FFC1FF22F832FE04AEBD" author="Bornmuller, J." box="[227,437,1989,2014]" pageId="6" pageNumber="273" pagination="7 - 8" refId="ref3469" refString="Bornmuller, J. (1911 a) Scutellaria xylorrhiza. Russkii Botanicheskii Zhurnal 1: 7 - 8." type="journal article" year="1911">Bornmüller 1911a</bibRefCitation>
), and in the same year, re-published in Beihefte zum Botanischen Centralblatt, Abteilung (
<bibRefCitation id="EF85CDD4FFB3FFC1FF4EF81EFED5A161" author="Bornmuller, J." box="[143,356,2025,2050]" pageId="6" pageNumber="273" pagination="458 - 535" refId="ref3489" refString="Bornmuller, J. (1911 b) Collectiones Straussianae novae. Beihefte zum Botanischen Centralblatt Abt. 2. 28: 458 - 535." type="journal article" year="1911">Bornmüller 1911b</bibRefCitation>
). Since the latter article addresses to the publication in Russkij Botaniceskij Zurnal, this may be considered as the protologue, although an earlier publication of the first is not proven.
</paragraph>
<paragraph id="8BABB025FFB2FFC0FF7CFF57FB64A8F2" blockId="7.[136,1452,160,401]" pageId="7" pageNumber="274">
The protologue (
<bibRefCitation id="EF85CDD4FFB2FFC0FEB2FF57FDF1A9DA" author="Bornmuller, J." box="[371,576,160,185]" pageId="7" pageNumber="274" pagination="7 - 8" refId="ref3469" refString="Bornmuller, J. (1911 a) Scutellaria xylorrhiza. Russkii Botanicheskii Zhurnal 1: 7 - 8." type="journal article" year="1911">Bornmüller 1911a</bibRefCitation>
) refers to a specimen from Kuh-i-Barsuk, collected on
<date id="FFAA96E5FFB2FFC0FB56FF57FA99A9DA" box="[1175,1320,160,185]" pageId="7" pageNumber="274" value="1904-06-22">22 June 1904</date>
by Theodor Strauss, but without designation of that specimen as the
<typeStatus id="54AF0E87FFB2FFC0FD3CFF33FCD2A9BE" box="[765,867,196,221]" pageId="7" pageNumber="274" type="holotype">holotype</typeStatus>
. No specimen with that collection date was found in the collections of B, JE and W, although W houses a specimen from Kuh Barsuk collected on
<date id="FFAA96E5FFB2FFC0FB5DFF1FFA85A862" box="[1180,1332,232,257]" pageId="7" pageNumber="274" value="1904-06-23">23 June 1904</date>
. Since this specimen has no original label, there might have been a transcription error. Another gathering at B and JE, labelled Kohrud,
<date id="FFAA96E5FFB2FFC0FF2DFEC7FE35A82A" box="[236,388,304,329]" pageId="7" pageNumber="274" value="1904-06-20">20 June 1904</date>
, and cited by
<bibRefCitation id="EF85CDD4FFB2FFC0FDE1FEC7FCB1A82A" author="Bornmuller, J." box="[544,768,304,329]" pageId="7" pageNumber="274" pagination="458 - 535" refId="ref3489" refString="Bornmuller, J. (1911 b) Collectiones Straussianae novae. Beihefte zum Botanischen Centralblatt Abt. 2. 28: 458 - 535." type="journal article" year="1911">Bornmüller (1911b)</bibRefCitation>
should be regarded as an additional
<typeStatus id="54AF0E87FFB2FFC0FB5AFEC7FB47A82A" box="[1179,1270,304,329]" pageId="7" pageNumber="274" type="syntype">syntype</typeStatus>
. As Barsuk is in the district of Kohrud, Strauss may discarded to write the name on all sheets. We here select the sample (JE00001284) at JE as the
<typeStatus id="54AF0E87FFB2FFC0FECFFE8FFEC4A8F2" box="[270,373,376,401]" pageId="7" pageNumber="274" type="lectotype">lectotype</typeStatus>
and, the voucher specimen (B100241792) deposited at B as the
<typeStatus id="54AF0E87FFB2FFC0FB87FE8FFB60A8F2" box="[1094,1233,376,401]" pageId="7" pageNumber="274" type="isolectotype">isolectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>