treatments-xml/data/03/BB/0D/03BB0D181F28FFE4718BF934FD09FF46.xml

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<mods:title id="D75E520ECDE69D0C0D36C626FE0B12F4">Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)</mods:title>
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<heading id="D0E50B621F28FFE1718BF934FC2DF971" bold="true" box="[151,902,1736,1762]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="4C12C78D1F28FFE1718BF934FEF3F971" ID-CoL="WJXT" box="[151,344,1736,1762]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F28FFE1718BF934FEF3F971" bold="true" box="[151,344,1736,1762]" italics="true" pageId="16" pageNumber="17">Cobitis fahireae</emphasis>
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Erk'akan, 
<bibRefCitation id="EF83C1FF1F28FFE170F9F934FC2DF971" author="Erk'akan, F. &amp; Atalay-Ekmekci, F. G. &amp; Nalbant, T. T." box="[485,902,1736,1762]" pageId="16" pageNumber="17" pagination="9 - 15" refId="ref46673" refString="Erk'akan, F., Atalay-Ekmekci, F. G. &amp; Nalbant, T. T. (1998) Four new species and one new subspecies of the genus Cobitis (Pisces: Ostariophysi: Cobitidae) from Turkey. Turkish Journal of Zoology, 22, 9 - 15." type="journal article" year="1998">Atalay-Ekmekçi &amp; Nalbant, 1998</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BADBC0E1F28FFE1718BF917FE8AF897" blockId="16.[151,902,1736,1796]" box="[151,289,1771,1796]" pageId="16" pageNumber="17">
(
<figureCitation id="1329A08B1F28FFE17183F917FEB2F897" box="[159,281,1771,1796]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="19.[151,250,1974,1996]" captionStartId-1="20.[151,250,460,482]" captionStartId-2="20.[151,250,1285,1307]" captionTargetBox-0="[189,1404,193,1957]" captionTargetBox-1="[202,1430,194,414]" captionTargetBox-2="[150,1436,532,1264]" captionTargetId-0="figure@19.[189,1404,193,1957]" captionTargetId-1="figure@20.[151,1435,193,439]" captionTargetId-2="figure@20.[150,1436,532,1264]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="20" captionText-0="FIGURE 11. Cobitis fahireae, FSJF 1975, male, 46 mm SL; female, 62 mm SL; Turkey: Bakır River; FSJF 3703, female, 57 mm SL; Turkey: stream Develi; FSJF 3692, female, 51 mm SL, male 40 mm SL; Turkey: Büyük Menderes River." captionText-1="FIGURE 12. Cobitis fahireae, HUIC uncat., holotype of C. kurui, 44.5 mm SL; Turkey: Küçük Menderes River." captionText-2="FIGURE 13. Cobitis fahireae, from the top: HUIC, uncat., holotype of C. damlae, female, 64 mm SL; Turkey: stream Bayındır. Photo: Füsun Erk'akan; FSJF 3692, female, 55 mm SL; Turkey: stream Bayındır." httpUri-0="https://zenodo.org/record/2615797/files/figure.png" httpUri-1="https://zenodo.org/record/2615799/files/figure.png" httpUri-2="https://zenodo.org/record/2615801/files/figure.png" pageId="16" pageNumber="17">Fig. 11–13</figureCitation>
)
</paragraph>
<paragraph id="8BADBC0E1F28FFE1718BF8CFFEBFF8FB" blockId="16.[151,1436,1843,1988]" pageId="16" pageNumber="17">
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<taxonomicName id="4C12C78D1F28FFE1718BF8CFFE94F8DA" ID-CoL="WJXT" box="[151,319,1843,1865]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F28FFE1718BF8CFFE94F8DA" box="[151,319,1843,1865]" italics="true" pageId="16" pageNumber="17">Cobitis fahireae</emphasis>
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Erk'akan, 
<treatmentCitation id="0AB39A1F1F28FFE170AFF8CFFCE9F8DA" author="Erk'akan, F. &amp; Atalay-Ekmekci, F. G. &amp; Nalbant, T. T." box="[435,834,1843,1865]" page="10" pageId="16" pageNumber="17" year="1998">
<bibRefCitation id="EF83C1FF1F28FFE170AFF8CFFCB3F8DA" author="Erk'akan, F. &amp; Atalay-Ekmekci, F. G. &amp; Nalbant, T. T." box="[435,792,1843,1865]" pageId="16" pageNumber="17" pagination="9 - 15" refId="ref46673" refString="Erk'akan, F., Atalay-Ekmekci, F. G. &amp; Nalbant, T. T. (1998) Four new species and one new subspecies of the genus Cobitis (Pisces: Ostariophysi: Cobitidae) from Turkey. Turkish Journal of Zoology, 22, 9 - 15." type="journal article" year="1998">Atalay-Ekmekçi &amp; Nalbant, 1998</bibRefCitation>
: 10
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, fig. 2 (
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locality: 
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: Küçük Menderes, Selçuk- 
<collectingRegion id="49D672EC1F28FFE171DBF8AEFEA2F8FB" box="[199,265,1874,1896]" country="Turkey" name="Aydin" pageId="16" pageNumber="17">Aydin</collectingRegion>
).
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</paragraph>
<paragraph id="8BADBC0E1F28FFE1718BF88CFE52F836" blockId="16.[151,1436,1843,1988]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB029B201F28FFE1718BF88CFE52F836" pageId="16" pageNumber="17">
<taxonomicName id="4C12C78D1F28FFE1718BF88CFE0BF815" ID-CoL="WJZ2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[151,416,1904,1926]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="vardarensis" subSpecies="kurui">
<emphasis id="B966601C1F28FFE1718BF88CFE0BF815" box="[151,416,1904,1926]" italics="true" pageId="16" pageNumber="17">Cobitis vardarensis kurui</emphasis>
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Erk'akan, 
<treatmentCitation id="0AB39A1F1F28FFE17309F88CFC0EF815" author="Erk'akan, F. &amp; Atalay-Ekmekci, F. G. &amp; Nalbant, T. T." box="[533,933,1904,1926]" page="13" pageId="16" pageNumber="17" year="1998">
<bibRefCitation id="EF83C1FF1F28FFE17309F88CFCD7F815" author="Erk'akan, F. &amp; Atalay-Ekmekci, F. G. &amp; Nalbant, T. T." box="[533,892,1904,1926]" pageId="16" pageNumber="17" pagination="9 - 15" refId="ref46673" refString="Erk'akan, F., Atalay-Ekmekci, F. G. &amp; Nalbant, T. T. (1998) Four new species and one new subspecies of the genus Cobitis (Pisces: Ostariophysi: Cobitidae) from Turkey. Turkish Journal of Zoology, 22, 9 - 15." type="journal article" year="1998">Atalay-Ekmekçi &amp; Nalbant, 1998</bibRefCitation>
: 13
</treatmentCitation>
, fig. 6 (
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locality: 
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: Menderes River, Selçuk-Aydin, Saplik Bridge).
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</paragraph>
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<paragraph id="8BADBC0E1F28FFE0718BF852FC50FE43" blockId="16.[151,1436,1843,1988]" lastBlockId="17.[151,1437,151,2014]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB029B201F28FFE0718BF852FC50FE43" infered="true" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<taxonomicName id="4C12C78D1F28FFE1718BF852FDD3F857" ID-CoL="WJXF" authority="Erkakan &amp; Ozmedir, 2014: 276" authorityName="Erkakan &amp; Ozmedir" authorityPageNumber="276" authorityYear="2014" box="[151,632,1966,1988]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F28FFE1718BF852FE9BF857" box="[151,304,1966,1988]" italics="true" pageId="16" pageNumber="17">Cobitis damlae</emphasis>
<treatmentCitation id="0AB39A1F1F28FFE1702BF852FDD3F857" author="Erk'akan, F. &amp; Ozdemir, F." box="[311,632,1966,1988]" page="276" pageId="16" pageNumber="17" year="2014">
<bibRefCitation id="EF83C1FF1F28FFE1702BF852FDECF857" author="Erk'akan, F. &amp; Ozdemir, F." box="[311,583,1966,1988]" pageId="16" pageNumber="17" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Erkakan &amp; Özmedir, 2014</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
, fig. 1 (type locality: 
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: Gölhisar prov.: Dalaman stream, 37.148 29.661). 
<emphasis id="B966601C1F29FFE0718BFF6BFE2FFF23" bold="true" box="[151,388,151,176]" pageId="17" pageNumber="18">Material examined.</emphasis>
FSJF 1975, 2, 
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SL; 
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: 
<collectingRegion id="49D672EC1F29FFE07242FF6BFC31FF23" box="[862,922,151,176]" country="Turkey" name="Izmir" pageId="17" pageNumber="18">İzmir</collectingRegion>
prov.: Bakır River at Karadere about 
<quantity id="4CEA11EB1F29FFE07443FF64FA37FF23" box="[1375,1436,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="2.0">2 km</quantity>
northeast of Kınık, 39.100 27.400.—FSJF 2294, 16, 
<quantity id="4CEA11EB1F29FFE073FEFF40FCF0FF47" box="[738,859,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" metricValueMax="7.6" metricValueMin="4.2" pageId="17" pageNumber="18" unit="mm" value="59.0" valueMax="76.0" valueMin="42.0">42–76 mm</quantity>
SL; 
<collectingCountry id="F305FC9E1F29FFE0728CFF40FC4FFF46" box="[912,996,188,213]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE072F2FF40FB80FF46" box="[1006,1067,188,213]" country="Turkey" name="Izmir" pageId="17" pageNumber="18">İzmir</collectingRegion>
prov.: stream Çiçekli, a tributary to stream Nif at Çiçekli, 38.505 27.272.—FSJF 2397, 11, 
<quantity id="4CEA11EB1F29FFE0722EFF1CFC07FF64" box="[818,940,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" metricValueMax="6.7" metricValueMin="4.7" pageId="17" pageNumber="18" unit="mm" value="57.0" valueMax="67.0" valueMin="47.0">47–67 mm</quantity>
SL; 
<collectingCountry id="F305FC9E1F29FFE072FBFF23FB90FF6B" box="[999,1083,223,248]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE0755BFF23FB07FF6B" box="[1095,1196,223,248]" country="Turkey" name="Balikesir" pageId="17" pageNumber="18">Balıkesir</collectingRegion>
prov.: stream Madra about 
<quantity id="4CEA11EB1F29FFE071C2FEF8FEB1FE8E" box="[222,282,260,285]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
south of Altınova, 39.218 26.819.—HUIC uncat., 
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, 2, 44– 
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SL; 
<collectingCountry id="F305FC9E1F29FFE075A9FEF8FAA2FE8E" box="[1205,1289,260,285]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
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prov.: stream Nif.—FSJF 3692, 4, 
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SL; 
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: 
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prov.: stream Bayındır 
<quantity id="4CEA11EB1F29FFE0754FFED4FB25FED3" box="[1107,1166,295,320]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
west of Çavdır, 37.149 29.661.—FSJF 3703, 3, 
<quantity id="4CEA11EB1F29FFE070B1FEB0FD8CFEF7" box="[429,551,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.7" metricValueMin="3.3" pageId="17" pageNumber="18" unit="mm" value="45.0" valueMax="57.0" valueMin="33.0">33–57 mm</quantity>
SL; 
<collectingRegion id="49D672EC1F29FFE07343FEB0FD30FEF6" box="[607,667,332,357]" country="Turkey" name="Izmir" pageId="17" pageNumber="18">İzmir</collectingRegion>
prov.: stream Develi 
<quantity id="4CEA11EB1F29FFE0728EFEB0FC66FEF6" box="[914,973,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="km" value="1.0">1 km</quantity>
south of Develi, 38.201 27.171.—HUIC uncat., 
<typeStatus id="54A902AC1F29FFE071F5FE93FEE0FE1B" box="[233,331,367,392]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE0706DFE8CFE65FE1B" ID-CoL="WJZ2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[369,462,368,392]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE0706DFE8CFE65FE1B" box="[369,462,368,392]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
, 
<quantity id="4CEA11EB1F29FFE070C0FE93FDEAFE14" box="[476,577,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="17" pageNumber="18" unit="mm" value="44.5">44.5 mm</quantity>
SL; 
<typeStatus id="54A902AC1F29FFE0736BFE8CFD7CFE1B" box="[631,727,368,392]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE073E2FE8CFCF0FE1B" ID-CoL="WJZ2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[766,859,368,392]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE073E2FE8CFCF0FE1B" box="[766,859,368,392]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
, 
<quantity id="4CEA11EB1F29FFE07275FE8CFC12FE14" box="[873,953,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="17" pageNumber="18" unit="mm" value="41.0">41 mm</quantity>
SL; 
<collectingCountry id="F305FC9E1F29FFE072ECFE93FBEFFE1B" box="[1008,1092,367,392]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE07553FE93FB38FE1B" box="[1103,1171,367,392]" country="Turkey" name="Aydin" pageId="17" pageNumber="18">Aydın</collectingRegion>
prov.: Menderes River, Saplık bridge.—FSJF 3709, 1, 
<quantity id="4CEA11EB1F29FFE0731FFE68FDFCFE3F" box="[515,599,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="18" unit="mm" value="62.0">62 mm</quantity>
SL; 
<collectingCountry id="F305FC9E1F29FFE07389FE68FD42FE3E" box="[661,745,404,429]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE073E4FE68FC90FE3E" box="[760,827,404,429]" country="Turkey" name="Aydin" pageId="17" pageNumber="18">Aydın</collectingRegion>
prov.: Akçay River 
<quantity id="4CEA11EB1F29FFE0752CFE68FBD0FE3E" box="[1072,1147,404,429]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="km" value="10.0">10 km</quantity>
south of Nazilli, 37.810 28.315.—ZMH 4752, 6, 
<quantity id="4CEA11EB1F29FFE070B2FE4BFD8CFE5C" box="[430,551,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.9" metricValueMin="4.2" pageId="17" pageNumber="18" unit="mm" value="50.5" valueMax="59.0" valueMin="42.0">42–59 mm</quantity>
SL; 
<collectingCountry id="F305FC9E1F29FFE07341FE4BFD1AFE43" box="[605,689,439,464]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: Golusar Gölü, SW 
<collectingCountry id="F305FC9E1F29FFE0728EFE4BFC5DFE43" box="[914,1014,439,464]" name="Turkey" pageId="17" pageNumber="18">Anatolia</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F29FFE071DBFE20FA36FD3B" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BADBC0E1F29FFE071DBFE20FA36FD3B" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
<emphasis id="B966601C1F29FFE071DBFE20FD14FE66" bold="true" box="[199,703,476,501]" pageId="17" pageNumber="18">Material used in molecular genetic analysis.</emphasis>
FSJF DNA-308; 
<collectingCountry id="F305FC9E1F29FFE07263FE20FC7BFE66" box="[895,976,476,501]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE072C6FE20FB90FE66" box="[986,1083,476,501]" country="Turkey" name="Balikesir" pageId="17" pageNumber="18">Balıkesir</collectingRegion>
prov.: stream Madra about 
<quantity id="4CEA11EB1F29FFE07479FE20FA37FE66" box="[1381,1436,476,501]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
south of Altınova, 39.218 26.819. (GenBank accession numbers: 
<accessionNumber id="944121ED1F29FFE07278FE03FC7CFD8B" box="[868,983,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553185" pageId="17" pageNumber="18">KJ553185</accessionNumber>
, 
<accessionNumber id="944121ED1F29FFE072FFFE03FBFCFD8B" box="[995,1111,511,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553276" pageId="17" pageNumber="18">KJ553276</accessionNumber>
). 
<emphasis id="B966601C1F29FFE0757EFE03FBD5FD8B" bold="true" box="[1122,1150,511,536]" pageId="17" pageNumber="18">—</emphasis>
FSJF DNA-2815; 
<collectingCountry id="F305FC9E1F29FFE0745BFE03FA32FD8B" box="[1351,1433,511,536]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE0718BFDD8FF4FFDAE" box="[151,228,548,573]" country="Turkey" name="Burdur" pageId="17" pageNumber="18">Burdur</collectingRegion>
prov.: stream Bayındır 
<quantity id="4CEA11EB1F29FFE070FCFDD8FDBCFDAE" box="[480,535,548,573]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
west of Çavdır, 37.149 29.661. (BLOLD accession numbers: BOLD EUFWF2721- 18 to EUFWF2723-18).—FSJF DNA-2855; 
<collectingCountry id="F305FC9E1F29FFE0739BFDBBFD72FDF3" box="[647,729,583,608]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE073F8FDBBFCB4FDF3" box="[740,799,583,608]" country="Turkey" name="Izmir" pageId="17" pageNumber="18">İzmir</collectingRegion>
prov.: stream Develi about 
<quantity id="4CEA11EB1F29FFE07549FDB4FB25FDF3" box="[1109,1166,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="km" value="1.0">1 km</quantity>
south of Develi, 38.201 27.171. (BOLD accession numbers: BOLD EUFWF2736-18 to EUFWF2738-18).—FSJF DNA-266 
<collectingCountry id="F305FC9E1F29FFE0741EFD90FAFFFD16" box="[1282,1364,620,645]" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
: 
<collectingRegion id="49D672EC1F29FFE0747DFD90FA37FD16" box="[1377,1436,620,645]" country="Turkey" name="Izmir" pageId="17" pageNumber="18">İzmir</collectingRegion>
prov.: stream Çiçekli, a tributary to stream Nif at Çiçekli, 38.505 27.272. (BOLD accession number: EUFWF4326-18).
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F29FFE071DBFD48FCF2FCCF" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BADBC0E1F29FFE071DBFD48FCF2FCCF" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
<emphasis id="B966601C1F29FFE071DBFD48FEEAFD5E" bold="true" box="[199,321,692,717]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
<taxonomicName id="4C12C78D1F29FFE07057FD48FDA9FD5F" box="[331,514,692,716]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07057FD48FDA9FD5F" box="[331,514,692,716]" italics="true" pageId="17" pageNumber="18">Cobitis fahireae</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C12C78D1F29FFE07215FD48FC17FD5F" box="[777,956,692,716]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="elongatoides">
<emphasis id="B966601C1F29FFE07215FD48FC17FD5F" box="[777,956,692,716]" italics="true" pageId="17" pageNumber="18">C. elongatoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F29FFE072D1FD48FBEEFD5F" authorityName="Vasil'eva &amp; Vasil'ev" authorityYear="2006" box="[973,1093,692,716]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pontica">
<emphasis id="B966601C1F29FFE072D1FD48FBEEFD5F" box="[973,1093,692,716]" italics="true" pageId="17" pageNumber="18">C. pontica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F29FFE0759EFD48FAA2FD5F" box="[1154,1289,692,716]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tanaitica">
<emphasis id="B966601C1F29FFE0759EFD48FAA2FD5F" box="[1154,1289,692,716]" italics="true" pageId="17" pageNumber="18">C. tanaitica</emphasis>
</taxonomicName>
by having a small or no black spot at the caudal-fin base (vs. roundish or ovoid, large, about size of pupil or eye) or if larger, then having one black spot at the upper and one black spot at the lower caudal-fin base (vs. always one spot only). It is distinguished from other 
<taxonomicName id="4C12C78D1F29FFE070FDFCDCFD99FCAB" box="[481,562,800,824]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F29FFE070FDFCDCFD99FCAB" box="[481,562,800,824]" italics="true" pageId="17" pageNumber="18">Cobitis</emphasis>
</taxonomicName>
species in the eastern 
<collectingRegion id="49D672EC1F29FFE07237FCDCFC2BFCAB" box="[811,896,800,824]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="18">Aegean</collectingRegion>
Sea basin by having one lamina circularis in the male (vs. two in 
<taxonomicName id="4C12C78D1F29FFE0704EFCB8FE5AFCCF" authorityName="Erkakan, Ozdemir &amp; Ozeren" authorityYear="2017" box="[338,497,836,860]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="dorademiri">
<emphasis id="B966601C1F29FFE0704EFCB8FE5AFCCF" box="[338,497,836,860]" italics="true" pageId="17" pageNumber="18">C. dorademiri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F29FFE070E3FCB8FD28FCCF" authorityName="Battalgazi" authorityYear="1944" box="[511,643,836,860]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F29FFE070E3FCB8FD28FCCF" box="[511,643,836,860]" italics="true" pageId="17" pageNumber="18">C. phrygica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F29FFE073A5FCB8FCE3FCCF" authorityName="Karaman" authorityYear="1955" box="[697,840,836,860]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="strumicae">
<emphasis id="B966601C1F29FFE073A5FCB8FCE3FCCF" box="[697,840,836,860]" italics="true" pageId="17" pageNumber="18">C. strumicae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F29FFE071DBFC9BFB78FC36" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BADBC0E1F29FFE071DBFC9BFB78FC36" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
<emphasis id="B966601C1F29FFE071DBFC9BFECBFC13" bold="true" box="[199,352,871,896]" pageId="17" pageNumber="18">Distribution.</emphasis>
<taxonomicName id="4C12C78D1F29FFE07070FC94FD8DFC13" box="[364,550,872,896]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07070FC94FD8DFC13" box="[364,550,872,896]" italics="true" pageId="17" pageNumber="18">Cobitis fahireae</emphasis>
</taxonomicName>
occurs in the eastern 
<collectingRegion id="49D672EC1F29FFE07228FC94FC22FC13" box="[820,905,872,896]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="18">Aegean</collectingRegion>
Sea basin where it is found from the upper Dalaman River drainage (around Gölhisar) in the south to the Madra River drainage in the north.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F29FFE371DBFC53FD62FF64" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BADBC0E1F29FFE071DBFC53FA88FB31" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
<emphasis id="B966601C1F29FFE071DBFC53FE92FC5B" bold="true" box="[199,313,943,968]" pageId="17" pageNumber="18">Remarks.</emphasis>
Erk'akan 
<emphasis id="B966601C1F29FFE070A8FC4DFE41FC5B" box="[436,490,944,968]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) described 
<taxonomicName id="4C12C78D1F29FFE073D7FC4CFCE7FC5B" box="[715,844,944,968]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE073D7FC4CFCE7FC5B" box="[715,844,944,968]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
from &quot;Küçük Menderes, Selçuk-Aydin&quot; (= Küçük Menderes River at Selçuk in the 
<collectingRegion id="49D672EC1F29FFE0731FFC28FD1AFC7E" box="[515,689,980,1006]" country="Turkey" name="Aydin" pageId="17" pageNumber="18">Aydın province</collectingRegion>
) and they described another species, 
<taxonomicName id="4C12C78D1F29FFE07552FC29FA84FC7E" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1102,1327,981,1005]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="subSpecies" species="vardarensis" subSpecies="kurui">
<emphasis id="B966601C1F29FFE07552FC29FA84FC7E" box="[1102,1327,981,1005]" italics="true" pageId="17" pageNumber="18">C. vardarensis kurui</emphasis>
</taxonomicName>
, from the (Küçük) &quot;Menderes River, Selçuk-Aydin, Saplik Bridge&quot;. Both species have been collected at the same day and probably from the same spot or close by. Despite several visits, we failed to find 
<taxonomicName id="4C12C78D1F29FFE0750EFBE1FBCAFBA6" box="[1042,1121,1053,1077]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F29FFE0750EFBE1FBCAFBA6" box="[1042,1121,1053,1077]" italics="true" pageId="17" pageNumber="18">Cobitis</emphasis>
</taxonomicName>
in the remains of the Küçük Menderes River drainage. This might be related to the massive environmental changes in the area in the past 34 years. We examined one 
<typeStatus id="54A902AC1F29FFE0707FFB9AFE6AFBED" box="[355,449,1126,1150]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE070FAFB9AFDCAFBEE" box="[486,609,1125,1149]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE070FAFB9AFDCAFBEE" box="[486,609,1125,1149]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
(from the Gediz River drainage) and the 
<typeStatus id="54A902AC1F29FFE07536FB99FB21FBED" box="[1066,1162,1125,1150]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
and four 
<typeStatus id="54A902AC1F29FFE075E9FB9AFAF5FBED" box="[1269,1374,1126,1150]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE0749FFB9AFF43FB32" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE0749FFB9AFF43FB32" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
from the type locality. Three 
<typeStatus id="54A902AC1F29FFE07331FB76FD3DFB31" box="[557,662,1162,1186]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE073A6FB76FC80FB32" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[698,811,1161,1185]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE073A6FB76FC80FB32" box="[698,811,1161,1185]" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
are identified as 
<taxonomicName id="4C12C78D1F29FFE072FAFB76FBFBFB32" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[998,1104,1161,1185]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="afifeae">
<emphasis id="B966601C1F29FFE072FAFB76FBFBFB32" box="[998,1104,1161,1185]" italics="true" pageId="17" pageNumber="18">C. afifeae</emphasis>
</taxonomicName>
(described below).
</paragraph>
<paragraph id="8BADBC0E1F29FFE071DBFB51FD17FA0D" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
Erk'akan 
<emphasis id="B966601C1F29FFE07033FB52FEF4FB56" box="[303,351,1197,1221]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998, 1999) report 
<taxonomicName id="4C12C78D1F29FFE07351FB52FD62FB56" box="[589,713,1197,1221]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07351FB52FD62FB56" box="[589,713,1197,1221]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
also from the lower Simav River (Marmara Sea basin; identified as 
<taxonomicName id="4C12C78D1F29FFE071ABFB2EFEB4FB7A" authorityName="Linnaeus" authorityYear="1758" box="[183,287,1234,1257]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="taenia">
<emphasis id="B966601C1F29FFE071ABFB2EFEB4FB7A" box="[183,287,1234,1257]" italics="true" pageId="17" pageNumber="18">C. taenia</emphasis>
</taxonomicName>
, see below) and from the Bakır and Gediz River drainages (
<collectingRegion id="49D672EC1F29FFE072C8FB2EFB87FB79" box="[980,1068,1234,1258]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="18">Aegean</collectingRegion>
Sea basin) and they report 
<taxonomicName id="4C12C78D1F29FFE07474FB2EFF79FA9E" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE07474FB2EFF79FA9E" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
from the Gediz and the Tahtalı (=Sasal) Rivers (all 
<collectingRegion id="49D672EC1F29FFE0723AFB0AFCD0FA9D" box="[806,891,1270,1294]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="18">Aegean</collectingRegion>
Sea basin). This suggests that both, 
<taxonomicName id="4C12C78D1F29FFE07401FB0AFA37FA9E" box="[1309,1436,1269,1293]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07401FB0AFA37FA9E" box="[1309,1436,1269,1293]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F29FFE071D4FAE6FE94FAA2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[200,319,1305,1329]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE071D4FAE6FE94FAA2" box="[200,319,1305,1329]" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
should occur in sympatry in the Küçük Menderes and in the Gediz River drainages. We were not able to find more than one 
<taxonomicName id="4C12C78D1F29FFE070C2FAC1FD84FAC6" box="[478,559,1341,1365]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F29FFE070C2FAC1FD84FAC6" box="[478,559,1341,1365]" italics="true" pageId="17" pageNumber="18">Cobitis</emphasis>
</taxonomicName>
species in the Gediz River drainage but 
<bibRefCitation id="EF83C1FF1F29FFE0750BFAC1FA82FAC5" author="Guclu, S. S. &amp; Kucuk, F." box="[1047,1321,1341,1366]" pageId="17" pageNumber="18" pagination="202 - 214" refId="ref47461" refString="Guclu, S. S. &amp; Kucuk, F. (2015) The Ichthyofauna of Gediz River (Turkey): Taxonomic and zoogeographic features. Annual Research &amp; Review in Biology, 6 (3), 202 - 214. https: // doi. org / 10.9734 / ARRB / 2015 / 14889" type="journal article" year="2015">Güçlü &amp; Küçük (2015)</bibRefCitation>
report 
<taxonomicName id="4C12C78D1F29FFE0749EFAC2FF5FFAEA" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE0749EFAC2FF5FFAEA" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F29FFE07036FA9EFE2CFAEA" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[298,391,1377,1401]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE07036FA9EFE2CFAEA" box="[298,391,1377,1401]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
from the Gediz and distinguish both species by the colour of the spot at the upper caudal-fin base (black in 
<taxonomicName id="4C12C78D1F29FFE07025FA7AFE1CFA0E" box="[313,439,1413,1437]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07025FA7AFE1CFA0E" box="[313,439,1413,1437]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
vs. brown in 
<taxonomicName id="4C12C78D1F29FFE07353FA7AFD00FA0E" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[591,683,1413,1437]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE07353FA7AFD00FA0E" box="[591,683,1413,1437]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BADBC0E1F29FFE071DBFA55FAD4F9E5" blockId="17.[151,1437,151,2014]" pageId="17" pageNumber="18">
Erk'akan 
<emphasis id="B966601C1F29FFE07028FA56FEC2FA52" box="[308,361,1449,1473]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) described 
<taxonomicName id="4C12C78D1F29FFE0735BFA55FD50FA52" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[583,763,1449,1473]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE0735BFA55FD50FA52" box="[583,763,1449,1473]" italics="true" pageId="17" pageNumber="18">Cobitis v. kurui</emphasis>
</taxonomicName>
as a subspecies of 
<taxonomicName id="4C12C78D1F29FFE072F8FA56FB20FA52" authorityName="Karaman" authorityYear="1928" box="[996,1163,1449,1473]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="vardarensis">
<emphasis id="B966601C1F29FFE072F8FA56FB20FA52" box="[996,1163,1449,1473]" italics="true" pageId="17" pageNumber="18">C. vardarensis</emphasis>
</taxonomicName>
and distinguished both subspecies by the brown, very small or absent spot at the upper caudal-fin base in 
<taxonomicName id="4C12C78D1F29FFE07537FA32FB34FA76" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1067,1183,1485,1509]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE07537FA32FB34FA76" box="[1067,1183,1485,1509]" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
(vs. large and black in 
<taxonomicName id="4C12C78D1F29FFE0718BFA0EFEF1F99A" box="[151,346,1521,1545]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" variety="vardarensis">
<emphasis id="B966601C1F29FFE0718BFA0EFEF1F99A" box="[151,346,1521,1545]" italics="true" pageId="17" pageNumber="18">C. v. vardarensis</emphasis>
</taxonomicName>
) and having more blotches along Z4 (no details given). Indeed, based on the description by Erk'akan 
<emphasis id="B966601C1F29FFE0701CF9EAFE9BF9BE" box="[256,304,1557,1581]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) and the 
<typeStatus id="54A902AC1F29FFE070F4F9EAFD88F9BD" box="[488,547,1558,1582]" pageId="17" pageNumber="18">types</typeStatus>
examined, both subspecies can be immediately distinguished by the very small or absent upper caudal-fin base spot in 
<taxonomicName id="4C12C78D1F29FFE07356F9C6FD16F9C2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[586,701,1593,1617]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE07356F9C6FD16F9C2" box="[586,701,1593,1617]" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
(vs. large in 
<taxonomicName id="4C12C78D1F29FFE07251F9C6FBACF9C2" box="[845,1031,1593,1617]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" variety="vardarensis">
<emphasis id="B966601C1F29FFE07251F9C6FBACF9C2" box="[845,1031,1593,1617]" italics="true" pageId="17" pageNumber="18">C. v. vardarensis</emphasis>
</taxonomicName>
) and the pigmentation in Z3 formed by one line or a narrow band of brown spots (vs. many small, brown spots the in Z3 giving it a &quot;sandy&quot; pattern).
</paragraph>
<paragraph id="8BADBC0E1F29FFE371DBF97DFD62FF64" blockId="17.[151,1437,151,2014]" lastBlockId="18.[151,1436,151,2037]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Despite 
<taxonomicName id="4C12C78D1F29FFE07039F97DFE78F90A" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[293,467,1665,1689]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE07039F97DFE78F90A" box="[293,467,1665,1689]" italics="true" pageId="17" pageNumber="18">Cobitis v. kurui</emphasis>
</taxonomicName>
having been described from the same place as 
<taxonomicName id="4C12C78D1F29FFE072EEF97EFBDBF90A" box="[1010,1136,1665,1689]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE072EEF97EFBDBF90A" box="[1010,1136,1665,1689]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
, Erk'akan 
<emphasis id="B966601C1F29FFE075F7F97EFAB6F90A" box="[1259,1309,1665,1689]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) do not compare 
<taxonomicName id="4C12C78D1F29FFE07034F95AFE28F92E" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[296,387,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE07034F95AFE28F92E" box="[296,387,1701,1725]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C12C78D1F29FFE070DEF95AFD95F92E" box="[450,574,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE070DEF95AFD95F92E" box="[450,574,1701,1725]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
directly, but state that 
<taxonomicName id="4C12C78D1F29FFE07227F95AFC3DF92E" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[827,918,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE07227F95AFC3DF92E" box="[827,918,1701,1725]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
is almost identical to 
<emphasis id="B966601C1F29FFE07596F95AFA9FF92E" box="[1162,1332,1701,1725]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C12C78D1F29FFE07596F95AFA9BF92E" authorityName="Karaman" authorityYear="1928" box="[1162,1328,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="vardarensis">C. vardarensis</taxonomicName>
.
</emphasis>
Erk'akan 
<emphasis id="B966601C1F29FFE0718BF936FF62F972" box="[151,201,1737,1761]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) distinguish 
<taxonomicName id="4C12C78D1F29FFE070B1F936FD80F972" box="[429,555,1737,1761]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE070B1F936FD80F972" box="[429,555,1737,1761]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
from the &quot; 
<taxonomicName id="4C12C78D1F29FFE073BBF935FC29F972" authorityName="Karaman" authorityYear="1928" box="[679,898,1737,1761]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="vardarensis">
<emphasis id="B966601C1F29FFE073BBF935FC29F972" box="[679,898,1737,1761]" italics="true" pageId="17" pageNumber="18">Cobitis vardarensis</emphasis>
</taxonomicName>
complex&quot; by having a reduced pigmentation in Z3, the pigmentation in Z2 dotted and a rounded, small caudal spot. Indeed, in 
<taxonomicName id="4C12C78D1F29FFE07535F912FB43F896" box="[1065,1256,1773,1797]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" variety="vardarensis">
<emphasis id="B966601C1F29FFE07535F912FB43F896" box="[1065,1256,1773,1797]" italics="true" pageId="17" pageNumber="18">C. v. vardarensis</emphasis>
</taxonomicName>
there are many small, brown spots the in Z3 giving it a &quot;sandy&quot; pattern while in the types of 
<taxonomicName id="4C12C78D1F29FFE07558F8EEFB6DF8BA" box="[1092,1222,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE07558F8EEFB6DF8BA" box="[1092,1222,1809,1833]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F29FFE0741AF8EEFACCF8BA" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1286,1383,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F29FFE0741AF8EEFACCF8BA" box="[1286,1383,1809,1833]" italics="true" pageId="17" pageNumber="18">C. kurui</emphasis>
</taxonomicName>
, the pigmentation in Z3 is &quot;reduced&quot; as there is just one line or a narrow band of brown spots. Erk'akan 
<emphasis id="B966601C1F29FFE07413F8CAFAEAF8DE" box="[1295,1345,1845,1869]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (1998) also state, that the pigmentation in Z2 is dotted in 
<taxonomicName id="4C12C78D1F29FFE073DAF8A6FCE8F8E2" box="[710,835,1881,1905]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE073DAF8A6FCE8F8E2" box="[710,835,1881,1905]" italics="true" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
(vs. short stripes in 
<taxonomicName id="4C12C78D1F29FFE07538F8A6FB33F8E2" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1060,1176,1881,1905]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F29FFE07538F8A6FB33F8E2" box="[1060,1176,1881,1905]" italics="true" pageId="17" pageNumber="18">C. v. kurui</emphasis>
</taxonomicName>
). However, in our own materials (e.g. 
<figureCitation id="1329A08B1F29FFE07023F881FE29F805" box="[319,386,1917,1942]" captionStart="FIGURE 9" captionStartId="15.[151,250,1753,1775]" captionTargetBox="[151,1436,965,1746]" captionTargetId="figure@15.[151,1436,965,1746]" captionTargetPageId="15" captionText="FIGURE 9. Cobitis elazigensis, FSJF 2559, male, 75 mm SL; Turkey: stream Eğri; FSJF 2495, female, 122 mm SL; Turkey: canal below Cipköy reservoir." httpUri="https://zenodo.org/record/2615793/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
) there is a great variation in this character and all intermediate character states are observed. In addition, the spot at the upper caudal-fin base was found to be small and black or brown or absent in individuals of one population and no difference in the size and the shape of the spot could be found between the 
<typeStatus id="54A902AC1F29FFE075EFF83AFAF5F84D" box="[1267,1374,1990,2014]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C12C78D1F29FFE3749EF83AFF5FFF23" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="18" lastPageNumber="19" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F29FFE3749EF83AFF5FFF23" italics="true" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F2AFFE37035FF64FE37FF23" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[297,412,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F2AFFE37035FF64FE37FF23" box="[297,412,152,176]" italics="true" pageId="18" pageNumber="19">C. v. kurui</emphasis>
</taxonomicName>
, while the 
<typeStatus id="54A902AC1F2AFFE37304FF6BFDD1FF23" box="[536,634,151,176]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C12C78D1F2AFFE37382FF64FD51FF23" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[670,762,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F2AFFE37382FF64FD51FF23" box="[670,762,152,176]" italics="true" pageId="18" pageNumber="19">C. kurui</emphasis>
</taxonomicName>
has no black spot. We found all characters distinguishing 
<taxonomicName id="4C12C78D1F2AFFE3749EFF64FF5FFF47" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3749EFF64FF5FFF47" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F2AFFE37036FF40FE35FF47" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[298,414,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F2AFFE37036FF40FE35FF47" box="[298,414,188,212]" italics="true" pageId="18" pageNumber="19">C. v. kurui</emphasis>
</taxonomicName>
to be highly variable even within populations. As we fail to distinguish 
<taxonomicName id="4C12C78D1F2AFFE375D5FF40FA95FF47" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1225,1342,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F2AFFE375D5FF40FA95FF47" box="[1225,1342,188,212]" italics="true" pageId="18" pageNumber="19">C. v. kurui</emphasis>
</taxonomicName>
from 
<emphasis id="B966601C1F2AFFE3749EFF40FF51FF6B" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C12C78D1F2AFFE3749EFF40FF5DFF6B" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">C. fahireae</taxonomicName>
,
</emphasis>
we have to treat both species as just one.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F2AFFE371DBFEF8FC95FE14" pageId="18" pageNumber="19" type="description">
<paragraph id="8BADBC0E1F2AFFE371DBFEF8FC95FE14" blockId="18.[151,1436,151,2037]" pageId="18" pageNumber="19">
As 
<taxonomicName id="4C12C78D1F2AFFE371F3FEF8FEC4FE8F" box="[239,367,260,284]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE371F3FEF8FEC4FE8F" box="[239,367,260,284]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F2AFFE370B6FEF8FD89FE8F" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[426,546,260,284]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F2AFFE370B6FEF8FD89FE8F" box="[426,546,260,284]" italics="true" pageId="18" pageNumber="19">C. v. kurui</emphasis>
</taxonomicName>
were described in the same study, a First Reviser action is needed. The First Reviser action is the principle that in cases of conflicts between simultaneously published names, the first subsequent author can decide which name has precedence (
<bibRefCitation id="EF83C1FF1F2AFFE37255FEB0FBC4FEF7" author="ICZN &amp; International Code of Zoological Nomenclature" box="[841,1135,332,357]" pageId="18" pageNumber="19" refId="ref47544" refString="ICZN (International Code of Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomeclature, London, 306 pp." type="book" year="1999">ICZN 1999: Article 24.2</bibRefCitation>
). Here we act as the First Revisers and give precedence to 
<taxonomicName id="4C12C78D1F2AFFE3731AFE8CFD2FFE1B" box="[518,644,368,392]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3731AFE8CFD2FFE1B" box="[518,644,368,392]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
over 
<taxonomicName id="4C12C78D1F2AFFE373DFFE8CFC9CFE1B" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[707,823,368,392]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="variety" species="vardarensis" variety="kurui">
<emphasis id="B966601C1F2AFFE373DFFE8CFC9CFE1B" box="[707,823,368,392]" italics="true" pageId="18" pageNumber="19">C. v. kurui</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F2AFFE371DBFE68FC1DFBC7" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BADBC0E1F2AFFE371DBFE68FBF9FD7D" blockId="18.[151,1436,151,2037]" pageId="18" pageNumber="19">
<taxonomicName id="4C12C78D1F2AFFE371DBFE68FED9FE3F" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[199,370,404,428]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE371DBFE68FED9FE3F" box="[199,370,404,428]" italics="true" pageId="18" pageNumber="19">Cobitis damlae</emphasis>
</taxonomicName>
was described by Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE373D4FE68FCD7FE3F" author="Erk'akan, F. &amp; Ozdemir, F." box="[712,892,404,429]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
based on one female individual from the stream Bayındır in the upper Dalaman River drainage. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE37228FE4BFC43FE43" author="Erk'akan, F. &amp; Ozdemir, F." box="[820,1000,439,464]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
claim that 
<taxonomicName id="4C12C78D1F2AFFE37579FE44FB72FE5C" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[1125,1241,439,463]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE37579FE44FB72FE5C" box="[1125,1241,439,463]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
is a subterranean species but it has been collected from surface waters and not from an underground habitat. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE37426FE27FF4AFD84" author="Erk'akan, F. &amp; Ozdemir, F." pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
speculate that the sole individual was washed out from underground waters nearby. However, they provide no evidence for this speculation. Baran Yoğurtçuoğlu (
<collectingRegion id="49D672EC1F2AFFE373E0FDDEFCF9FDA8" box="[764,850,546,571]" country="Turkey" name="Ankara" pageId="18" pageNumber="19">Ankara</collectingRegion>
) &amp; JF visited the 
<typeStatus id="54A902AC1F2AFFE37507FDDFFBE0FDA8" box="[1051,1099,547,571]" pageId="18" pageNumber="19">type</typeStatus>
locality of 
<taxonomicName id="4C12C78D1F2AFFE375D0FDDFFAEBFDA8" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[1228,1344,547,571]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE375D0FDDFFAEBFDA8" box="[1228,1344,547,571]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
in 2017 and found no spring or any subterranean waters adjacent to the 
<typeStatus id="54A902AC1F2AFFE37245FDBBFC22FDCC" box="[857,905,583,607]" pageId="18" pageNumber="19">type</typeStatus>
locality. The stream Bayındır flows down from a reservoir and we found no tributaries, which might originate from an underground source. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE37426FD96FF4AFD34" author="Erk'akan, F. &amp; Ozdemir, F." pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
mentioned the caves Keloğlan and Aslanini in the upper Dalaman River drainage about 
<quantity id="4CEA11EB1F2AFFE375A6FD73FAABFD34" box="[1210,1280,654,679]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="km" value="30.0">30 km</quantity>
from the 
<typeStatus id="54A902AC1F2AFFE37470FD73FA37FD34" box="[1388,1436,655,679]" pageId="18" pageNumber="19">type</typeStatus>
locality of 
<taxonomicName id="4C12C78D1F2AFFE3700DFD4EFE2FFD59" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[273,388,690,714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE3700DFD4EFE2FFD59" box="[273,388,690,714]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
, but no troglomorphic fish have ever been reported from these caves. It remains unclear how these caves or other subterranean habitats might be connected to the stream Bayındır.
</paragraph>
<paragraph id="8BADBC0E1F2AFFE371DBFD05FC1DFBC7" blockId="18.[151,1436,151,2037]" pageId="18" pageNumber="19">
The 
<typeStatus id="54A902AC1F2AFFE371E7FD05FEF6FC81" box="[251,349,761,786]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
and single known individual of 
<taxonomicName id="4C12C78D1F2AFFE373D2FD05FCD2FC82" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[718,889,761,785]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE373D2FD05FCD2FC82" box="[718,889,761,785]" italics="true" pageId="18" pageNumber="19">Cobitis damlae</emphasis>
</taxonomicName>
(
<figureCitation id="1329A08B1F2AFFE37296FD05FC76FC81" box="[906,989,761,786]" captionStart="FIGURE 13" captionStartId="20.[151,250,1285,1307]" captionTargetBox="[150,1436,532,1264]" captionTargetId="figure@20.[150,1436,532,1264]" captionTargetPageId="20" captionText="FIGURE 13. Cobitis fahireae, from the top: HUIC, uncat., holotype of C. damlae, female, 64 mm SL; Turkey: stream Bayındır. Photo: Füsun Erk'akan; FSJF 3692, female, 55 mm SL; Turkey: stream Bayındır." httpUri="https://zenodo.org/record/2615801/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
) has no pigmentation and its eye is not reduced in size. Surprisingly, Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE3739CFCE0FC93FCA6" author="Erk'akan, F. &amp; Ozdemir, F." box="[640,824,796,821]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
distinguished 
<taxonomicName id="4C12C78D1F2AFFE372FFFCE1FBF1FCA6" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[995,1114,797,821]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE372FFFCE1FBF1FCA6" box="[995,1114,797,821]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
by having vestigial eyes, a character state not seen from the picture. In subterranean fishes, the eye is usually reduced in size already in the very early phase of adaptation to an underground environment, while the pigmentation pattern is reduced later on, resulting in many subterranean fish with very small eyes but still having a colour patter (
<bibRefCitation id="EF83C1FF1F2AFFE375CEFC74FA3AFC33" author="Proudlove, G." box="[1234,1425,904,929]" pageId="18" pageNumber="19" refId="ref48804" refString="Proudlove, G. (2003) Subterranean fishes of the world: An account of the subterranean (hypogean) fishes described up to 2003 with a bibliography 1541 - 2004. International Society for Subterranean Biology, Moulis, 300 pp." type="book" year="2003">Proudlove 2003</bibRefCitation>
). Furthermore, many subterranean fish with eyes become grey or brown, if exposed to light for some days (own observation). The situation is the opposite in 
<taxonomicName id="4C12C78D1F2AFFE373B1FC2CFC8FFC7B" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[685,804,976,1000]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE373B1FC2CFC8FFC7B" box="[685,804,976,1000]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
. Here the pigmentation is absent, but the eye is fully developed. On the picture (
<figureCitation id="1329A08B1F2AFFE370DBFC0FFDB2FB9F" box="[455,537,1011,1036]" captionStart="FIGURE 13" captionStartId="20.[151,250,1285,1307]" captionTargetBox="[150,1436,532,1264]" captionTargetId="figure@20.[150,1436,532,1264]" captionTargetPageId="20" captionText="FIGURE 13. Cobitis fahireae, from the top: HUIC, uncat., holotype of C. damlae, female, 64 mm SL; Turkey: stream Bayındır. Photo: Füsun Erk'akan; FSJF 3692, female, 55 mm SL; Turkey: stream Bayındır." httpUri="https://zenodo.org/record/2615801/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
), the eye-background is orange, suggesting, that the animal is albinotic. While it is impossible to exclude that the 
<typeStatus id="54A902AC1F2AFFE3730EFBEBFDDFFBA3" box="[530,628,1047,1072]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C12C78D1F2AFFE37381FBE4FCB8FBBC" ID-CoL="WJXF" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[669,787,1047,1071]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE37381FBE4FCB8FBBC" box="[669,787,1047,1071]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
originates from an underground habitat, there are strong indications that it is just an albinotic individual of a surface population.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F2AFFE471DBFBA2FD09FF46" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BADBC0E1F2AFFE371DBFBA2FBDCF867" blockId="18.[151,1436,151,2037]" pageId="18" pageNumber="19">
Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE3704CFBA2FDAEFBE4" author="Erk'akan, F. &amp; Ozdemir, F." box="[336,517,1118,1143]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
identified the pigmented 
<taxonomicName id="4C12C78D1F2AFFE37237FBA2FCD7FBE5" box="[811,892,1118,1142]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE37237FBA2FCD7FBE5" box="[811,892,1118,1142]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
from the stream Bayındır as 
<taxonomicName id="4C12C78D1F2AFFE375D7FBA3FACDFBE5" authorityName="Bacescu" authorityYear="1962" box="[1227,1382,1118,1142]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F2AFFE375D7FBA3FACDFBE5" box="[1227,1382,1118,1142]" italics="true" pageId="18" pageNumber="19">C. battalgilae</emphasis>
</taxonomicName>
(see discussion above, identifying this population as 
<taxonomicName id="4C12C78D1F2AFFE373ADFB7FFC9DFB09" authorityName="Battalgazi" authorityYear="1944" box="[689,822,1154,1178]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE373ADFB7FFC9DFB09" box="[689,822,1154,1178]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
) but we found 
<taxonomicName id="4C12C78D1F2AFFE372FCFB7FFBF5FB09" box="[992,1118,1154,1178]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE372FCFB7FFBF5FB09" box="[992,1118,1154,1178]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
to be common in the stream Bayındır. Indeed, the absence of pigmentation and the fact that the 
<typeStatus id="54A902AC1F2AFFE372B9FB5AFBACFB2C" box="[933,1031,1190,1215]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C12C78D1F2AFFE3752EFB5BFB03FB2D" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[1074,1192,1190,1214]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE3752EFB5BFB03FB2D" box="[1074,1192,1190,1214]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
is a female makes it difficult to identify it as one of the two 
<taxonomicName id="4C12C78D1F2AFFE37366FB36FD60FB71" box="[634,715,1226,1250]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE37366FB36FD60FB71" box="[634,715,1226,1250]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
species present in the area (
<taxonomicName id="4C12C78D1F2AFFE3753FFB36FB0EFB71" box="[1059,1189,1226,1250]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3753FFB36FB0EFB71" box="[1059,1189,1226,1250]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2AFFE375ABFB36FAEBFB71" authorityName="Battalgazi" authorityYear="1944" box="[1207,1344,1226,1250]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE375ABFB36FAEBFB71" box="[1207,1344,1226,1250]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
). Other characters given by Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE370E3FB11FD18FA95" author="Erk'akan, F. &amp; Ozdemir, F." box="[511,691,1261,1286]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
do not allow distinguishing these two 
<taxonomicName id="4C12C78D1F2AFFE37579FB11FB1DFA96" box="[1125,1206,1261,1285]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE37579FB11FB1DFA96" box="[1125,1206,1261,1285]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
species. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE3718BFAEDFED6FAB9" author="Erk'akan, F. &amp; Ozdemir, F." box="[151,381,1297,1322]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014:278)</bibRefCitation>
distinguish 
<taxonomicName id="4C12C78D1F2AFFE37314FAEEFDD6FABA" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[520,637,1297,1321]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE37314FAEEFDD6FABA" box="[520,637,1297,1321]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C12C78D1F2AFFE373DFFAEEFCF5FABA" authorityName="Bacescu" authorityYear="1962" box="[707,862,1297,1321]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F2AFFE373DFFAEEFCF5FABA" box="[707,862,1297,1321]" italics="true" pageId="18" pageNumber="19">C. battalgilae</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C12C78D1F2AFFE3729BFAEEFBA7FABA" authorityName="Battalgazi" authorityYear="1944" box="[903,1036,1297,1321]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE3729BFAEEFBA7FABA" box="[903,1036,1297,1321]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
) by having 6½ branched dorsal-fin rays (vs. 8) but the individual of 
<taxonomicName id="4C12C78D1F2AFFE3731FFAC9FD23FADE" authorityName="Battalgazi" authorityYear="1944" box="[515,648,1333,1357]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE3731FFAC9FD23FADE" box="[515,648,1333,1357]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
shown by Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE37299FAC8FB9FFADE" author="Erk'akan, F. &amp; Ozdemir, F." box="[901,1076,1332,1357]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014</bibRefCitation>
: 
<figureCitation id="1329A08B1F2AFFE37522FAC8FBD4FADE" box="[1086,1151,1332,1357]" captionStart="FIGURE 2" captionStartId="8.[151,250,1957,1979]" captionTargetBox="[151,1436,1010,1918]" captionTargetId="figure@8.[151,1436,1010,1918]" captionTargetPageId="8" captionText="FIGURE 2. Site scale records of Cobitis species in the Middle East based on materials examined for this study and additional records listed in Table 10." httpUri="https://zenodo.org/record/2615779/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
[a drawing by T. Nalbant already published by Erk'akan 
<emphasis id="B966601C1F2AFFE370E3FAA5FD9FFAE2" box="[511,564,1369,1393]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 1999: Fig. 8]) has 6½ branched dorsal-fin rays (indeed it has 6 branched dorsal-fin rays as T. Nalbant always ignored the ½ ray on his drawings) and all 
<taxonomicName id="4C12C78D1F2AFFE3752FFA81FB2FFA06" box="[1075,1156,1405,1429]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE3752FFA81FB2FFA06" box="[1075,1156,1405,1429]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
examined for this study have 6½ or 7½ branched dorsal-fin rays. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE373E7FA5CFC48FA2B" author="Erk'akan, F. &amp; Ozdemir, F." box="[763,995,1440,1465]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014:279)</bibRefCitation>
further distinguish 
<taxonomicName id="4C12C78D1F2AFFE375D8FA5CFA91FA2B" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[1220,1338,1440,1464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE375D8FA5CFA91FA2B" box="[1220,1338,1440,1464]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C12C78D1F2AFFE3749EFA5CFEBAFA4F" authorityName="Bacescu" authorityYear="1962" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F2AFFE3749EFA5CFEBAFA4F" italics="true" pageId="18" pageNumber="19">C. battalgilae</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C12C78D1F2AFFE37027FA38FE69FA4F" authorityName="Battalgazi" authorityYear="1944" box="[315,450,1476,1500]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE37027FA38FE69FA4F" box="[315,450,1476,1500]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
) by having 8+8 branched caudal-fin rays (vs. 7+7) and 6½ branched pectoral-fin rays (vs. 7 
<emphasis id="B966601C1F2AFFE371C9FA1BFF48F993" bold="true" box="[213,227,1511,1536]" pageId="18" pageNumber="19">–</emphasis>
8). A range of 7+7, 8+7 or 8+8 is common in populations of the 
<taxonomicName id="4C12C78D1F2AFFE372A9FA14FBADF993" box="[949,1030,1512,1536]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE372A9FA14FBADF993" box="[949,1030,1512,1536]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
species examined for this study and 
<taxonomicName id="4C12C78D1F2AFFE3718BF9F0FEBEF9B0" box="[151,277,1547,1571]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3718BF9F0FEBEF9B0" box="[151,277,1547,1571]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
caught by us at the type locality of 
<taxonomicName id="4C12C78D1F2AFFE373B6F9F0FCB5F9B0" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[682,798,1547,1571]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE373B6F9F0FCB5F9B0" box="[682,798,1547,1571]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
have 7+7 branched caudal-fin rays. No details are given how the pectoral-fin rays are counted. Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE373CFF9D3FC2CF9D4" author="Erk'akan, F. &amp; Ozdemir, F." box="[723,903,1583,1608]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
refer to 
<bibRefCitation id="EF83C1FF1F2AFFE372F5F9D3FAAAF9DB" author="Economidis, P. S. &amp; Nalbant, T. T." box="[1001,1281,1583,1608]" pageId="18" pageNumber="19" pagination="295 - 347" refId="ref46572" refString="Economidis, P. S. &amp; Nalbant, T. T. (1996) A study of the loaches of the genera Cobitis and Sabanejewia (Pisces, Cobitidae) of Greece, with description of six new taxa. Travaux du Museum d'Histoire Naturelle &quot; Grigore Antipa &quot;, 36, 295 - 347" type="journal article" year="1996">
Economidis 
<emphasis id="B966601C1F2AFFE3756AF9CCFB03F9D4" box="[1142,1192,1583,1607]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1996)
</bibRefCitation>
and Erk'akan 
<emphasis id="B966601C1F2AFFE3718BF9A8FF62F9F8" box="[151,201,1619,1643]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1998) for methods of count. Neither 
<bibRefCitation id="EF83C1FF1F2AFFE37363F9AFFC33F9FF" author="Economidis, P. S. &amp; Nalbant, T. T." box="[639,920,1619,1644]" pageId="18" pageNumber="19" pagination="295 - 347" refId="ref46572" refString="Economidis, P. S. &amp; Nalbant, T. T. (1996) A study of the loaches of the genera Cobitis and Sabanejewia (Pisces, Cobitidae) of Greece, with description of six new taxa. Travaux du Museum d'Histoire Naturelle &quot; Grigore Antipa &quot;, 36, 295 - 347" type="journal article" year="1996">
Economidis 
<emphasis id="B966601C1F2AFFE37211F9A8FC94F9F8" box="[781,831,1619,1643]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1996)
</bibRefCitation>
nor Erk'akan 
<emphasis id="B966601C1F2AFFE3752AF9A8FBC3F9F8" box="[1078,1128,1619,1643]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1998) count the pectoralfin rays as ½ and we can only suspect, that there are 7 branched pectoral-fin rays in the type of 
<taxonomicName id="4C12C78D1F2AFFE375DBF98BFA90F91D" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[1223,1339,1654,1678]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE375DBF98BFA90F91D" box="[1223,1339,1654,1678]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
(vs. 
<quantity id="4CEA11EB1F2AFFE3746EF98BFF06F920" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="18" pageNumber="19" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">
7 
<emphasis id="B966601C1F2AFFE3749CF98AFA25F91C" bold="true" box="[1408,1422,1654,1679]" pageId="18" pageNumber="19">–</emphasis>
8 in
</quantity>
<taxonomicName id="4C12C78D1F2AFFE371AFF967FEE5F921" authorityName="Bacescu" authorityYear="1962" box="[179,334,1690,1714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F2AFFE371AFF967FEE5F921" box="[179,334,1690,1714]" italics="true" pageId="18" pageNumber="19">C. battalgilae</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C12C78D1F2AFFE37076F967FE44F921" authorityName="Battalgazi" authorityYear="1944" box="[362,495,1690,1714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE37076F967FE44F921" box="[362,495,1690,1714]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
). In the 
<taxonomicName id="4C12C78D1F2AFFE37350F967FD62F921" box="[588,713,1690,1714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE37350F967FD62F921" box="[588,713,1690,1714]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
from the type locality of 
<taxonomicName id="4C12C78D1F2AFFE372F9F967FBF3F921" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[997,1112,1690,1714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE372F9F967FBF3F921" box="[997,1112,1690,1714]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
as well as in 
<taxonomicName id="4C12C78D1F2AFFE375F2F967FAD9F921" authorityName="Battalgazi" authorityYear="1944" box="[1262,1394,1690,1714]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE375F2F967FAD9F921" box="[1262,1394,1690,1714]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
, 6– 10 branched pectoral-fin rays are very common and a range of 2–3 rays is typical in all species. The only indication given is the shape of the external part of the suborbital spine, which is simple in 
<taxonomicName id="4C12C78D1F2AFFE37524F91EFB15F969" authorityName="Battalgazi" authorityYear="1944" box="[1080,1214,1762,1786]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE37524F91EFB15F969" box="[1080,1214,1762,1786]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
and bifurcate in 
<taxonomicName id="4C12C78D1F2AFFE3749EF91EFF58F88E" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3749EF91EFF58F88E" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
. In the type of 
<taxonomicName id="4C12C78D1F2AFFE370BFF8FAFDBCF88E" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[419,535,1797,1821]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE370BFF8FAFDBCF88E" box="[419,535,1797,1821]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
, the spine is bifurcate (Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE372B7F8F9FBF9F88D" author="Erk'akan, F. &amp; Ozdemir, F." box="[939,1106,1797,1822]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir 2014</bibRefCitation>
). Furthermore, the type of 
<taxonomicName id="4C12C78D1F2AFFE3749EF8FAFF41F8D2" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE3749EF8FAFF41F8D2" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
is a fully ripe female individual of only 
<quantity id="4CEA11EB1F2AFFE373B2F8D5FD56F8D2" box="[686,765,1833,1858]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="18" pageNumber="19" unit="mm" value="64.0">64 mm</quantity>
SL. Females of 
<taxonomicName id="4C12C78D1F2AFFE372A8F8D6FB92F8D2" authorityName="Battalgazi" authorityYear="1944" box="[948,1081,1833,1857]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F2AFFE372A8F8D6FB92F8D2" box="[948,1081,1833,1857]" italics="true" pageId="18" pageNumber="19">C. phrygica</emphasis>
</taxonomicName>
grow much larger and reach up to 
<quantity id="4CEA11EB1F2AFFE371A8F8B1FEBAF8F6" box="[180,273,1869,1893]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="18" pageNumber="19" unit="mm" value="119.0">119 mm</quantity>
SL. As the type of 
<taxonomicName id="4C12C78D1F2AFFE370F3F8B1FDCFF8F6" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[495,612,1869,1893]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE370F3F8B1FDCFF8F6" box="[495,612,1869,1893]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
has a bifurcate suborbital spine, is relatively small and as we found only 
<taxonomicName id="4C12C78D1F2AFFE3718BF88DFEBEF81A" box="[151,277,1905,1929]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE3718BF88DFEBEF81A" box="[151,277,1905,1929]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
at the type locality, we are unable to distinguish 
<taxonomicName id="4C12C78D1F2AFFE37258F88DFC13F81A" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[836,952,1905,1929]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE37258F88DFC13F81A" box="[836,952,1905,1929]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C12C78D1F2AFFE372E3F88DFBD6F81A" box="[1023,1149,1905,1929]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE372E3F88DFBD6F81A" box="[1023,1149,1905,1929]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
. Our molecular data also place 
<taxonomicName id="4C12C78D1F2AFFE371C5F869FE81F83E" box="[217,298,1941,1965]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2AFFE371C5F869FE81F83E" box="[217,298,1941,1965]" italics="true" pageId="18" pageNumber="19">Cobitis</emphasis>
</taxonomicName>
from the type locality of 
<taxonomicName id="4C12C78D1F2AFFE37357F869FD14F83E" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[587,703,1941,1965]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE37357F869FD14F83E" box="[587,703,1941,1965]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
close to 
<taxonomicName id="4C12C78D1F2AFFE37238F869FC09F83E" box="[804,930,1941,1965]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE37238F869FC09F83E" box="[804,930,1941,1965]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
and we found no morphological character to distinguish these fishes from 
<taxonomicName id="4C12C78D1F2AFFE370F8F844FDC8F843" box="[484,611,1976,2000]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE370F8F844FDC8F843" box="[484,611,1976,2000]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
based on our materials examined and details given in the description by Erk'akan &amp; 
<bibRefCitation id="EF83C1FF1F2AFFE37001F820FE7FF867" author="Erk'akan, F. &amp; Ozdemir, F." box="[285,468,2012,2037]" pageId="18" pageNumber="19" pagination="275 - 279" refId="ref46838" refString="Erk'akan, F. &amp; Ozdemir, F. (2014) The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2), 275 - 279. https: // doi. org / 10.15671 / HJBC. 20144210863" type="journal article" year="2014">Özdemir (2014)</bibRefCitation>
. Therefore, we treat 
<taxonomicName id="4C12C78D1F2AFFE373A7F820FC84F867" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[699,815,2012,2036]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2AFFE373A7F820FC84F867" box="[699,815,2012,2036]" italics="true" pageId="18" pageNumber="19">C. damlae</emphasis>
</taxonomicName>
as a synonym of 
<taxonomicName id="4C12C78D1F2AFFE372EFF820FBDBF867" box="[1011,1136,2012,2036]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2AFFE372EFF820FBDBF867" box="[1011,1136,2012,2036]" italics="true" pageId="18" pageNumber="19">C. fahireae</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF6DEC861F2BFFE2718BF84AFAB1F878" httpUri="https://zenodo.org/record/2615797/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1974,1996]" targetBox="[189,1404,193,1957]" targetPageId="19">
<paragraph id="8BADBC0E1F2BFFE2718BF84AFAB1F878" blockId="19.[151,1436,1974,2027]" pageId="19" pageNumber="20">
<emphasis id="B966601C1F2BFFE2718BF84AFEB4F85F" bold="true" box="[151,287,1974,1996]" pageId="19" pageNumber="20">FIGURE 11.</emphasis>
<taxonomicName id="4C12C78D1F2BFFE27039F84AFE61F85F" box="[293,458,1974,1996]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2BFFE27039F84AFE61F85F" box="[293,458,1974,1996]" italics="true" pageId="19" pageNumber="20">Cobitis fahireae</emphasis>
</taxonomicName>
, FSJF 1975, male, 46 mm SL; female, 62 mm SL; Turkey: Bakır River; FSJF 3703, female, 57 mm SL; Turkey: stream Develi; FSJF 3692, female, 51 mm SL, male 40 mm SL; Turkey: Büyük Menderes River.
</paragraph>
</caption>
<caption id="DF6DEC861F2CFFE5718BFE30FABBFE71" box="[151,1296,460,482]" httpUri="https://zenodo.org/record/2615799/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,460,482]" targetBox="[202,1430,194,414]" targetPageId="20">
<paragraph id="8BADBC0E1F2CFFE5718BFE30FABBFE71" blockId="20.[151,1296,460,482]" box="[151,1296,460,482]" pageId="20" pageNumber="21">
<emphasis id="B966601C1F2CFFE5718BFE30FEB4FE71" bold="true" box="[151,287,460,482]" pageId="20" pageNumber="21">FIGURE 12.</emphasis>
<taxonomicName id="4C12C78D1F2CFFE57039FE30FE63FE71" box="[293,456,460,482]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE57039FE30FE63FE71" box="[293,456,460,482]" italics="true" pageId="20" pageNumber="21">Cobitis fahireae</emphasis>
</taxonomicName>
, HUIC uncat., holotype of 
<taxonomicName id="4C12C78D1F2CFFE573C1FE31FC9BFE71" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[733,816,460,482]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B966601C1F2CFFE573C1FE31FC9BFE71" box="[733,816,460,482]" italics="true" pageId="20" pageNumber="21">C. kurui</emphasis>
</taxonomicName>
, 44.5 mm SL; Turkey: Küçük Menderes River.
</paragraph>
</caption>
<caption id="DF6DEC861F2CFFE5718BFAF9FB84FAA9" httpUri="https://zenodo.org/record/2615801/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1285,1307]" targetBox="[150,1436,532,1264]" targetPageId="20">
<paragraph id="8BADBC0E1F2CFFE5718BFAF9FB84FAA9" blockId="20.[151,1435,1285,1338]" pageId="20" pageNumber="21">
<emphasis id="B966601C1F2CFFE5718BFAF9FE8FFA88" bold="true" box="[151,292,1285,1307]" pageId="20" pageNumber="21">FIGURE 13.</emphasis>
<taxonomicName id="4C12C78D1F2CFFE57033FAF9FE7CFA88" box="[303,471,1285,1307]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE57033FAF9FE7CFA88" box="[303,471,1285,1307]" italics="true" pageId="20" pageNumber="21">Cobitis fahireae</emphasis>
</taxonomicName>
, from the top: HUIC, uncat., holotype of 
<taxonomicName id="4C12C78D1F2CFFE572BFFAFAFBBBFA88" authorityName="Erkakan &amp; Ozmedir" authorityYear="2014" box="[931,1040,1285,1307]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="damlae">
<emphasis id="B966601C1F2CFFE572BFFAFAFBBBFA88" box="[931,1040,1285,1307]" italics="true" pageId="20" pageNumber="21">C. damlae</emphasis>
</taxonomicName>
, female, 64 mm SL; Turkey: stream Bayındır. Photo: Füsun Erk'akan; FSJF 3692, female, 55 mm SL; Turkey: stream Bayındır.
</paragraph>
</caption>
<paragraph id="8BADBC0E1F2CFFE571DBFA9EFAC6F971" blockId="20.[151,1437,1377,2014]" pageId="20" pageNumber="21">
Our molecular data suggest that there is substantial diversity within the species we call 
<taxonomicName id="4C12C78D1F2CFFE5758EFA9EFAA5FAEA" box="[1170,1294,1377,1401]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE5758EFA9EFAA5FAEA" box="[1170,1294,1377,1401]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
. We found a minimum COI K2P distance of 1.0% between 
<taxonomicName id="4C12C78D1F2CFFE573B1FA79FD55FA0E" box="[685,766,1413,1437]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE573B1FA79FD55FA0E" box="[685,766,1413,1437]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
from the Dalaman / Gediz Rivers and the Şaşal River and 1.3% K2P distance between 
<taxonomicName id="4C12C78D1F2CFFE570C9FA55FD8DFA52" box="[469,550,1449,1473]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE570C9FA55FD8DFA52" box="[469,550,1449,1473]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
from the Dalaman / Gediz Rivers and the Madra River. 
<taxonomicName id="4C12C78D1F2CFFE5758AFA55FB4CFA52" box="[1174,1255,1449,1473]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE5758AFA55FB4CFA52" box="[1174,1255,1449,1473]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
from the Madra River are distinguished from those from the Şaşal River by 1.8%. 
<bibRefCitation id="EF83C1FF1F2CFFE57266FA31FBC1FA75" author="Perdices, A. &amp; Ozeren, C. S. &amp; Erk'akan, F. &amp; Freyhof, J." box="[890,1130,1485,1510]" pageId="20" pageNumber="21" pagination="0205678" refId="ref48738" refString="Perdices, A., Ozeren, C. S., Erk'akan, F. &amp; Freyhof, J. (2018) Diversity of spined loaches from Asia Minor in a phylogenetic context (Teleostei: Cobitidae). PLoS ONE, 13 (10), e 0205678. https: // doi. org / 10.1371 / journal. pone. 0205678" type="journal article" year="2018">
Perdices 
<emphasis id="B966601C1F2CFFE572FCFA32FBBAFA76" box="[992,1041,1485,1509]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. (2018)
</bibRefCitation>
found very little molecular difference between 
<taxonomicName id="4C12C78D1F2CFFE5706FFA0DFE6FF99A" box="[371,452,1521,1545]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE5706FFA0DFE6FF99A" box="[371,452,1521,1545]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
populations from the Gediz and Bakir Rivers. We found no differences between 
<taxonomicName id="4C12C78D1F2CFFE57457FA0DFA37F99A" box="[1355,1436,1521,1545]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE57457FA0DFA37F99A" box="[1355,1436,1521,1545]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
populations from the Gediz and Dalaman Rivers. As the Küçük Menderes River is situated between the Gediz and the Dalaman Rivers, we could speculate, that this group is the one the 
<typeStatus id="54A902AC1F2CFFE572ABF9C6FC4CF9C1" box="[951,999,1594,1618]" pageId="20" pageNumber="21">type</typeStatus>
of 
<taxonomicName id="4C12C78D1F2CFFE57512F9C6FB27F9C2" box="[1038,1164,1593,1617]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE57512F9C6FB27F9C2" box="[1038,1164,1593,1617]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
was collected from. On the other hand, the Küçük Menderes River might have (had) an own fish fauna and its estuary is only 
<quantity id="4CEA11EB1F2CFFE5744EF9A1FA37F9E5" box="[1362,1436,1629,1654]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="km" value="25.0">25 km</quantity>
southwest of the Şaşal River estuary. Both rivers are very likely to have flown together during the last glaciation. Therefore, it is likely that 
<taxonomicName id="4C12C78D1F2CFFE570DAF959FDBCF92E" box="[454,535,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE570DAF959FDBCF92E" box="[454,535,1701,1725]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
from the Şaşal River are identical with 
<taxonomicName id="4C12C78D1F2CFFE572FAF95AFBCDF92E" box="[998,1126,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE572FAF95AFBCDF92E" box="[998,1126,1701,1725]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
from the Küçük Menderes River. More fieldwork is urgently needed, especially in the Küçük Menderes River, to locate 
<taxonomicName id="4C12C78D1F2CFFE575B5F936FA8CF972" box="[1193,1319,1737,1761]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE575B5F936FA8CF972" box="[1193,1319,1737,1761]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
there.
</paragraph>
<paragraph id="8BADBC0E1F2CFFE471DBF912FD09FF46" blockId="20.[151,1437,1377,2014]" lastBlockId="21.[151,1436,151,213]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
Our own molecular data (
<figureCitation id="1329A08B1F2CFFE570FFF911FD88F895" box="[483,547,1773,1798]" captionStart="FIGURE 1" captionStartId="6.[151,249,1665,1687]" captionTargetBox="[164,1417,206,1642]" captionTargetId="figure@6.[155,1431,190,1647]" captionTargetPageId="6" captionText="FIGURE 1. Maximum likelihood estimation of the phylogenetic relationships based on the mitochondrial COI barcode region (TN93 model, discrete Gamma distribution for rate differences with 3 categories +G parameter = 0.3792. Nucleotide positions with less than 95% site coverage were eliminated, resulting in 590 analysed positions. Numbers of major nodes indicate bootstrap values from 1000 pseudoreplicates from the ML, NJ and MP method. A star denotes highest possible support from all three methods. Solid bars right to the specimen labels indicate species delimitation results from PTP followed by the results of the mPTP approach as dashed bars. Except to the outgroups and the branch to the right part of the figure, the tree is drawn to scale with branch lengths depicting number of substitutions per site." httpUri="https://zenodo.org/record/2615777/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
) and 
<bibRefCitation id="EF83C1FF1F2CFFE5737FF911FCFAF895" author="Perdices, A. &amp; Ozeren, C. S. &amp; Erk'akan, F. &amp; Freyhof, J." box="[611,849,1773,1798]" pageId="20" pageNumber="21" pagination="0205678" refId="ref48738" refString="Perdices, A., Ozeren, C. S., Erk'akan, F. &amp; Freyhof, J. (2018) Diversity of spined loaches from Asia Minor in a phylogenetic context (Teleostei: Cobitidae). PLoS ONE, 13 (10), e 0205678. https: // doi. org / 10.1371 / journal. pone. 0205678" type="journal article" year="2018">
Perdices 
<emphasis id="B966601C1F2CFFE573D4F912FD52F896" box="[712,761,1773,1797]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. (2018)
</bibRefCitation>
place 
<taxonomicName id="4C12C78D1F2CFFE57284F911FBE7F896" box="[920,1100,1773,1797]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE57284F911FBE7F896" box="[920,1100,1773,1797]" italics="true" pageId="20" pageNumber="21">Cobitis fahireae</emphasis>
</taxonomicName>
in the 
<taxonomicName id="4C12C78D1F2CFFE5758BF912FB56F896" authorityName="Linnaeus" authorityYear="1758" box="[1175,1277,1774,1797]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="taenia">
<emphasis id="B966601C1F2CFFE5758BF912FB56F896" box="[1175,1277,1774,1797]" italics="true" pageId="20" pageNumber="21">C. taenia</emphasis>
</taxonomicName>
species group (
<taxonomicName id="4C12C78D1F2CFFE571BCF8EEFE9EF8BA" authorityName="Mousavi-Sabet, Vatandoust, Esmaeili, Geiger &amp; Freyhof" authorityYear="2015" box="[160,309,1810,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="avicennae">
<emphasis id="B966601C1F2CFFE571BCF8EEFE9EF8BA" box="[160,309,1810,1833]" italics="true" pageId="20" pageNumber="21">C. avicennae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE5705AF8EEFE52F8BA" box="[326,505,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="elongatoides">
<emphasis id="B966601C1F2CFFE5705AF8EEFE52F8BA" box="[326,505,1809,1833]" italics="true" pageId="20" pageNumber="21">C. elongatoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE57316F8EEFDC7F8BA" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[522,620,1810,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="emrei">
<emphasis id="B966601C1F2CFFE57316F8EEFDC7F8BA" box="[522,620,1810,1833]" italics="true" pageId="20" pageNumber="21">C. emrei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE57361F8EEFD56F8BA" box="[637,765,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE57361F8EEFD56F8BA" box="[637,765,1809,1833]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE57212F8EEFC33F8BA" box="[782,920,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="faridpaki">
<emphasis id="B966601C1F2CFFE57212F8EEFC33F8BA" box="[782,920,1809,1833]" italics="true" pageId="20" pageNumber="21">C. faridpaki</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE572B6F8EEFB8AF8BA" authorityName="Vasil'eva &amp; Vasil'ev" authorityYear="2006" box="[938,1057,1810,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pontica">
<emphasis id="B966601C1F2CFFE572B6F8EEFB8AF8BA" box="[938,1057,1810,1833]" italics="true" pageId="20" pageNumber="21">C. pontica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE5752FF8EEFB49F8BA" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[1075,1250,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="puncticulata">
<emphasis id="B966601C1F2CFFE5752FF8EEFB49F8BA" box="[1075,1250,1809,1833]" italics="true" pageId="20" pageNumber="21">C. puncticulata</emphasis>
</taxonomicName>
, 
<emphasis id="B966601C1F2CFFE575EFF8EEFE2BF8DE" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C12C78D1F2CFFE575EFF8EEFADFF8BA" authorityName="Gladkov" authorityYear="1935" box="[1267,1396,1810,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="satunini">C. satunini</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE5749EF8EEFF48F8DE" authorityName="Eagderi, Jouladeh-Roudbar, Jalili, Sayyadzadeh &amp; Esmaeili" authorityYear="2017" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="saniae">C. saniae</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE571ECF8CAFE2BF8DE" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[240,384,1845,1869]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="splendens">C. splendens</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C12C78D1F2CFFE5708CF8CAFDBCF8DE" box="[400,535,1846,1869]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tanaitica">
<emphasis id="B966601C1F2CFFE5708CF8CAFDBCF8DE" box="[400,535,1846,1869]" italics="true" pageId="20" pageNumber="21">C. tanaitica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2CFFE5733AF8CAFD31F8DE" authorityName="Vasil'eva, Vasil'ev, Janko, Ráb &amp; Rábová" authorityYear="2005" box="[550,666,1846,1869]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="taurica">
<emphasis id="B966601C1F2CFFE5733AF8CAFD31F8DE" box="[550,666,1846,1869]" italics="true" pageId="20" pageNumber="21">C. taurica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F2CFFE573C8F8CAFCD3F8DE" authorityName="Karaman" authorityYear="1928" box="[724,888,1845,1869]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="vardarensis">
<emphasis id="B966601C1F2CFFE573C8F8CAFCD3F8DE" box="[724,888,1845,1869]" italics="true" pageId="20" pageNumber="21">C. vardarensis</emphasis>
</taxonomicName>
). Based on DNA barcoding, 
<taxonomicName id="4C12C78D1F2CFFE575D5F8CAFAECF8DE" box="[1225,1351,1845,1869]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE575D5F8CAFAECF8DE" box="[1225,1351,1845,1869]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
is well separated from all other 
<taxonomicName id="4C12C78D1F2CFFE570B6F8A5FE50F8E2" box="[426,507,1881,1905]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2CFFE570B6F8A5FE50F8E2" box="[426,507,1881,1905]" italics="true" pageId="20" pageNumber="21">Cobitis</emphasis>
</taxonomicName>
included in this study and it is distinguished from its closest relative, 
<emphasis id="B966601C1F2CFFE5740CF8A6FA37F8E2" box="[1296,1436,1882,1905]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C12C78D1F2CFFE5740CF8A6FA3CF8E2" box="[1296,1431,1882,1905]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tanaitica">C. tanaitica</taxonomicName>
,
</emphasis>
by a minimum K2P distance of 4.5%, also supported as single PTP entity. It is further distinguished from 
<taxonomicName id="4C12C78D1F2CFFE5749EF882FE8DF82A" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="elongatoides">
<emphasis id="B966601C1F2CFFE5749EF882FE8DF82A" italics="true" pageId="20" pageNumber="21">C. elongatoides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C12C78D1F2CFFE5707DF85EFE73F82A" authorityName="Vasil'eva &amp; Vasil'ev" authorityYear="2006" box="[353,472,1954,1977]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pontica">
<emphasis id="B966601C1F2CFFE5707DF85EFE73F82A" box="[353,472,1954,1977]" italics="true" pageId="20" pageNumber="21">C. pontica</emphasis>
</taxonomicName>
by having one line or a narrow band of brown spots in Z3 (vs. many small, brown spots the in Z3 giving it a &quot;sandy&quot; pattern). See below for details to distinguish 
<taxonomicName id="4C12C78D1F2CFFE5751BF83AFB28F84E" box="[1031,1155,1989,2013]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fahireae">
<emphasis id="B966601C1F2CFFE5751BF83AFB28F84E" box="[1031,1155,1989,2013]" italics="true" pageId="20" pageNumber="21">C. fahireae</emphasis>
</taxonomicName>
from other species in the 
<taxonomicName id="4C12C78D1F2DFFE4718BFF64FF54FF3C" authorityName="Linnaeus" authorityYear="1758" box="[151,255,152,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="taenia">
<emphasis id="B966601C1F2DFFE4718BFF64FF54FF3C" box="[151,255,152,175]" italics="true" pageId="21" pageNumber="22">C. taenia</emphasis>
</taxonomicName>
species group in 
<collectingCountry id="F305FC9E1F2DFFE470DBFF6BFD8CFF23" box="[455,551,151,176]" name="Turkey" pageId="21" pageNumber="22">Anatolia</collectingCountry>
(
<taxonomicName id="4C12C78D1F2DFFE47325FF64FD4CFF23" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[569,743,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="puncticulata">
<emphasis id="B966601C1F2DFFE47325FF64FD4CFF23" box="[569,743,152,176]" italics="true" pageId="21" pageNumber="22">C. puncticulata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2DFFE473EAFF64FC2DFF23" authorityName="Erk'akan, Atalay-Ekmekçi &amp; Nalbant" authorityYear="1998" box="[758,902,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="splendens">
<emphasis id="B966601C1F2DFFE473EAFF64FC2DFF23" box="[758,902,152,176]" italics="true" pageId="21" pageNumber="22">C. splendens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2DFFE47289FF64FC56FF3C" authorityName="Linnaeus" authorityYear="1758" box="[917,1021,152,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="taenia">
<emphasis id="B966601C1F2DFFE47289FF64FC56FF3C" box="[917,1021,152,175]" italics="true" pageId="21" pageNumber="22">C. taenia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C12C78D1F2DFFE47510FF64FB22FF3C" authorityName="Gladkov" authorityYear="1935" box="[1036,1161,152,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="satunini">
<emphasis id="B966601C1F2DFFE47510FF64FB22FF3C" box="[1036,1161,152,175]" italics="true" pageId="21" pageNumber="22">C. satunini</emphasis>
</taxonomicName>
) and from other 
<taxonomicName id="4C12C78D1F2DFFE47457FF64FA37FF23" box="[1355,1436,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F2DFFE47457FF64FA37FF23" box="[1355,1436,152,176]" italics="true" pageId="21" pageNumber="22">Cobitis</emphasis>
</taxonomicName>
species found in the eastern Aegean Sea basin.
</paragraph>
</subSubSection>
</treatment>
</document>