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<document ID-DOI="http://doi.org/10.5281/zenodo.4758396" ID-GBIF-Dataset="d0f0400d-eb9b-4114-9a4e-a7f7f680e3a3" ID-GBIF-Taxon="182067635" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758396" checkinTime="1620844596556" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2008" docId="03BB0738FFE1FFC1FEB4FEC5FA96FBA3" docLanguage="en" docName="Illiesia.4.20.208-225.pdf" docOrigin="Illiesia 4 (20)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Togoperla limbata" docType="treatment" docVersion="6" lastPageNumber="215" masterDocId="FF827F40FFE6FFC6FFF1FFDAFFB7FFF1" masterDocTitle="The Genus Togoperla Klapálek (Plecoptera: Perlidae)" masterLastPageNumber="225" masterPageNumber="208" pageId="7" pageNumber="215" updateTime="1632022738459" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Genus Togoperla Klapálek (Plecoptera: Perlidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>20</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>208</mods:start>
<mods:end>225</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4754843</mods:identifier>
<mods:identifier type="GBIF-Dataset">d0f0400d-eb9b-4114-9a4e-a7f7f680e3a3</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4754843</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4758396" ID-GBIF-Taxon="182067635" ID-Zenodo-Dep="4758396" LSID="urn:lsid:plazi:treatment:03BB0738FFE1FFC1FEB4FEC5FA96FBA3" httpUri="http://treatment.plazi.org/id/03BB0738FFE1FFC1FEB4FEC5FA96FBA3" lastPageNumber="215" pageId="7" pageNumber="215">
<subSubSection box="[325,634,287,311]" pageId="7" pageNumber="215" type="nomenclature">
<paragraph blockId="7.[325,634,287,311]" box="[325,634,287,311]" pageId="7" pageNumber="215">
<heading bold="true" box="[325,634,287,311]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="215" reason="2">
<taxonomicName authority="(Pictet)" baseAuthorityName="Pictet" box="[325,634,287,311]" class="Insecta" family="Perlidae" genus="Togoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="limbata">
<emphasis bold="true" box="[325,634,287,311]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[325,538,287,311]" italics="true" pageId="7" pageNumber="215">Togoperla limbata</emphasis>
(Pictet)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[411,549,322,346]" pageId="7" pageNumber="215" type="description">
<paragraph blockId="7.[411,549,322,346]" box="[411,549,322,346]" pageId="7" pageNumber="215">
(
<figureCitation box="[419,541,322,346]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Figs. 21‐27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="215" type="reference_group">
<paragraph blockId="7.[221,776,394,635]" box="[221,541,394,416]" pageId="7" pageNumber="215">
<treatmentCitationGroup box="[221,541,394,416]" pageId="7" pageNumber="215">
<treatmentCitation author="Pictet, F. J." box="[221,537,394,416]" page="219" pageId="7" pageNumber="215" year="1841">
<taxonomicName authority="Pictet, 1841: 219" authorityName="Pictet" authorityPageNumber="219" authorityYear="1841" box="[221,537,394,416]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="limbata">
<emphasis box="[221,355,394,415]" italics="true" pageId="7" pageNumber="215">Perla limbata</emphasis>
<bibRefCitation author="Pictet, F. J." box="[369,537,394,416]" pageId="7" pageNumber="215" refId="ref8634" refString="Pictet, F. J. 1841. Histoire naturelle generale et particuliere des insectes Nevropteres. Famille des Perlides. 1. Partie. 423 pp." type="book" year="1841">Pictet, 1841:219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[221,776,394,635]" pageId="7" pageNumber="215">
<materialsCitation ID-GBIF-Occurrence="3111850309" country="Japan" location="Zwick" pageId="7" pageNumber="215" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus box="[554,656,395,416]" pageId="7" pageNumber="215">Lectotype</typeStatus>
<specimenCount box="[670,688,394,416]" pageId="7" pageNumber="215" type="male">♂</specimenCount>
(ZMB),
<collectingCountry box="[221,282,426,447]" name="Japan" pageId="7" pageNumber="215">Japan</collectingCountry>
, desig.
<bibRefCitation author="Zwick, P." box="[358,482,426,448]" pageId="7" pageNumber="215" pagination="1123 - 1194" refId="ref8903" refString="Zwick, P. 1972. Die Plecopteren Pictets und Burmeisters, mit Angaben uber weitere Arten (Insecta). Revue Suisse de Zoologie, 78: 1123 - 1194." type="journal article" year="1972">
<location LSID="urn:lsid:plazi:treatment:03BB0738FFE1FFC1FEB4FEC5FA96FBA3:8ECDE0F5FFE1FFC1FE97FE70FE1EFE31" box="[358,425,426,448]" country="Japan" name="Zwick" pageId="7" pageNumber="215">Zwick</location>
, 1972
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph blockId="7.[221,776,394,635]" box="[221,776,457,479]" pageId="7" pageNumber="215">
<treatmentCitationGroup box="[221,776,457,479]" pageId="7" pageNumber="215">
<treatmentCitation author="Okamoto, H." box="[221,772,457,479]" page="123" pageId="7" pageNumber="215" year="1912">
<taxonomicName authority="Okamoto, 1912: 123" authorityName="Okamoto" authorityPageNumber="123" authorityYear="1912" box="[221,772,457,479]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="kawamurae" subGenus="Togoperla">
<emphasis box="[221,270,457,478]" italics="true" pageId="7" pageNumber="215">Perla</emphasis>
(
<emphasis box="[301,394,457,478]" italics="true" pageId="7" pageNumber="215">Togoperla</emphasis>
)
<emphasis box="[425,533,457,478]" italics="true" pageId="7" pageNumber="215">kawamurae</emphasis>
<bibRefCitation author="Okamoto, H." box="[556,772,457,479]" pageId="7" pageNumber="215" pagination="105 - 170" refId="ref8606" refString="Okamoto, H. 1912. Erster Beitrag zur Kenntnis der Japanischen Plecopteren. Transactions of the Sapporo Natural History Society, 4: 105 - 170." type="journal article" year="1912">Okamoto, 1912:123</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[221,776,394,635]" pageId="7" pageNumber="215">
<materialsCitation ID-GBIF-Occurrence="3111850331" county="Japan" location="Japan" pageId="7" pageNumber="215" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus box="[221,319,488,510]" pageId="7" pageNumber="215">Holotype</typeStatus>
<specimenCount box="[332,350,488,510]" pageId="7" pageNumber="215" type="female">♀</specimenCount>
(Hokkaido University, Sapporo, lost),
<collectingCounty box="[221,282,520,541]" pageId="7" pageNumber="215">Japan</collectingCounty>
, syn.
<bibRefCitation author="Sivec, I., B. P." box="[340,509,519,541]" pageId="7" pageNumber="215" pagination="1 - 66" refId="ref8696" refString="Sivec, I., B. P. Stark &amp; S. Uchida. 1988. Synopsis of the world genera of Perlinae (Plecoptera: Perlidae). Scopolia, 16: 1 - 66." type="journal article" year="1988">Sivec et al., 1988</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph blockId="7.[221,776,394,635]" box="[221,671,551,573]" pageId="7" pageNumber="215">
<taxonomicName authority="Okamoto, 1912" authorityName="Okamoto, S. Uchida" authorityYear="1912" box="[221,668,551,573]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="matsumurae" subGenus="Togoperla">
<emphasis box="[221,270,551,572]" italics="true" pageId="7" pageNumber="215">Perla</emphasis>
(
<emphasis box="[282,375,551,572]" italics="true" pageId="7" pageNumber="215">Togoperla</emphasis>
)
<emphasis box="[388,506,552,573]" italics="true" pageId="7" pageNumber="215">matsumurae</emphasis>
<bibRefCitation author="Okamoto, H." box="[511,668,551,573]" pageId="7" pageNumber="215" pagination="105 - 170" refId="ref8606" refString="Okamoto, H. 1912. Erster Beitrag zur Kenntnis der Japanischen Plecopteren. Transactions of the Sapporo Natural History Society, 4: 105 - 170." type="journal article" year="1912">Okamoto, 1912</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="7.[221,776,394,635]" pageId="7" pageNumber="215">
<materialsCitation ID-GBIF-Occurrence="3111850321" pageId="7" pageNumber="215">
<typeStatus box="[675,776,551,572]" pageId="7" pageNumber="215">Lectotype</typeStatus>
<specimenCount box="[221,239,582,604]" pageId="7" pageNumber="215" type="female">♀</specimenCount>
, (
<collectingRegion box="[268,373,582,604]" country="Japan" name="Hokkaido" pageId="7" pageNumber="215">Hokkaido</collectingRegion>
University, Sapporo), Haki,
<collectingCountry box="[711,772,583,604]" name="Japan" pageId="7" pageNumber="215">Japan</collectingCountry>
, design.
<bibRefCitation author="Uchida, S." box="[301,437,613,635]" pageId="7" pageNumber="215" refId="ref8766" refString="Uchida, S. 1990. A revision of the Japanese Perlidae (Insecta: Plecoptera), with special reference to their phylogeny. Unpublished PhD Dissertation, Tokyo Metropolitan University. 228 pp." type="book" year="1990">Uchida, 1990</bibRefCitation>
, syn.
<bibRefCitation author="Sivec, I., B. P." box="[495,666,613,635]" pageId="7" pageNumber="215" pagination="1 - 66" refId="ref8696" refString="Sivec, I., B. P. Stark &amp; S. Uchida. 1988. Synopsis of the world genera of Perlinae (Plecoptera: Perlidae). Scopolia, 16: 1 - 66." type="journal article" year="1988">Sivec et al., 1988</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="215" type="materials_examined">
<paragraph blockId="7.[184,776,684,1953]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[184,411,684,708]" pageId="7" pageNumber="215">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111200312" collectingDate="1981-07-04" collectionCode="PMSL" collectorName="S. Uchida" country="Japan" location="Unknown" pageId="7" pageNumber="215" specimenCount="1" specimenCount-male="1">
<collectingCountry box="[417,485,685,708]" name="Japan" pageId="7" pageNumber="215">Japan</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03BB0738FFE1FFC1FEB4FEC5FA96FBA3:8ECDE0F5FFE1FFC1FE1FFD76FDD6FD35" box="[494,609,684,708]" country="Japan" name="Unknown" pageId="7" pageNumber="215">Unknown</location>
locality,
<date pageId="7" pageNumber="215" value="1981-07-04">
<collectingDate pageId="7" pageNumber="215" value="1981-07-04">4 July 1981</collectingDate>
</date>
,
<collectorName box="[252,366,719,743]" pageId="7" pageNumber="215">S. Uchida</collectorName>
,
<specimenCount box="[380,422,718,743]" pageId="7" pageNumber="215" type="male">1 ♂</specimenCount>
(
<collectionCode box="[440,515,719,743]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="7" pageNumber="215">PMSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111200309" collectingDate="1897-06" country="Japan" county="Tamba" location="Hozuyama" municipality="Kuwada Gori" pageId="7" pageNumber="215" specimenCount="3" specimenCount-female="3">
<location LSID="urn:lsid:plazi:treatment:03BB0738FFE1FFC1FEB4FEC5FA96FBA3:8ECDE0F5FFE1FFC1FDE6FD15FD2FFD16" box="[535,664,719,743]" country="Japan" county="Tamba" municipality="Kuwada Gori" name="Hozuyama" pageId="7" pageNumber="215">Hozuyama</location>
,
<collectingMunicipality pageId="7" pageNumber="215">Kuwada Gori</collectingMunicipality>
,
<collectingCounty box="[251,332,753,777]" pageId="7" pageNumber="215">Tamba</collectingCounty>
,
<date box="[345,460,753,777]" pageId="7" pageNumber="215" value="1897-06">
<collectingDate box="[345,460,753,777]" pageId="7" pageNumber="215" value="1897-06">June 1897</collectingDate>
</date>
,
<specimenCount box="[473,513,753,777]" pageId="7" pageNumber="215" type="female">3 ♀</specimenCount>
(??)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111200317" collectingDate="1926-05" collectionCode="CSU" country="Japan" location="Honshu" municipality="Obama" pageId="7" pageNumber="215" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:03BB0738FFE1FFC1FEB4FEC5FA96FBA3:8ECDE0F5FFE1FFC1FDB3FD2BFD17FCF8" box="[578,672,753,777]" country="Japan" municipality="Obama" name="Honshu" pageId="7" pageNumber="215">Honshu</location>
,
<collectingMunicipality box="[685,770,753,777]" pageId="7" pageNumber="215">Obama</collectingMunicipality>
,
<date box="[184,296,788,812]" pageId="7" pageNumber="215" value="1926-05">
<collectingDate box="[184,296,788,812]" pageId="7" pageNumber="215" value="1926-05">May 1926</collectingDate>
</date>
,
<specimenCount box="[307,345,787,812]" pageId="7" pageNumber="215" type="male">1 ♂</specimenCount>
(
<collectionCode box="[360,417,788,812]" country="USA" httpUri="http://grbio.org/cool/hsbf-qixz" name="Colorado State University" pageId="7" pageNumber="215">CSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="7.[184,776,684,1953]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[184,353,822,846]" pageId="7" pageNumber="215">Adult habitus.</emphasis>
General color brown. Head with dark brown pigment over ocelli and forward of M‐ line; occiput dusky brown. Pronotum brown but usually with pale anterolateral areas on disc (
<figureCitation box="[606,684,926,950]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 21</figureCitation>
). Wings brown except costal area pale. Legs entirely brown, without distinctive banding.
</paragraph>
<paragraph blockId="7.[184,776,684,1953]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[184,250,1030,1054]" pageId="7" pageNumber="215">Male.</emphasis>
Forewing length
<quantity box="[444,557,1030,1054]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="7" pageNumber="215" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19‐22 mm</quantity>
.
<collectionCode box="[565,582,1030,1054]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="215">T</collectionCode>
5 produced into a notched or emarginate lobe;
<collectionCode box="[499,515,1065,1089]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="215">T</collectionCode>
6‐
<collectionCode box="[535,551,1065,1089]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="215">T</collectionCode>
9 typical; membrane of
<collectionCode box="[211,227,1099,1123]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="215">T</collectionCode>
6‐
<collectionCode box="[248,264,1099,1123]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="215">T</collectionCode>
9 bearing patches of mixed setae and scattered sensilla basiconica. Dorsal aspect of hemiterga moderately long and attenuated from basal callus to tip; length from basal callus to hemitergal tip about 1.3‐1.5 times as long as basal callus (
<figureCitation box="[604,684,1237,1262]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 22</figureCitation>
); lateral aspect of hemiterga slender across basal curve and gradually narrowed to form a slender triangular structure (
<figureCitation box="[305,386,1341,1365]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 23</figureCitation>
); sensilla basiconica clustered on basal callus and sparse on hemitergal tip. Membranous aedeagal sac armed with a coarse band of variably sized spines from subapical lobes to about midlength of tube (
<figureCitation box="[404,482,1479,1503]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 25</figureCitation>
); largest spines occur in a band somewhat proximal to margin of spinous zone; apex of sac extends as a nose like lobe below a pair of membranous, unarmed dorsal lobes; apical lobe armed at tip with a zone of slender, long spines; tube with a pair of midlateral spiny lobes.
</paragraph>
<paragraph blockId="7.[184,776,684,1953]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[184,276,1687,1711]" pageId="7" pageNumber="215">Female.</emphasis>
Forewing length
<quantity box="[473,586,1687,1711]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="7" pageNumber="215" unit="mm" value="26.5" valueMax="29.0" valueMin="24.0">24‐29 mm</quantity>
. Subgenital plate parabolic without apical emargination or notch. Dorsal membrane of plate with a short sclerotized mesal band reaching less than halfway to base of plate (
<figureCitation box="[254,332,1825,1849]" captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 26</figureCitation>
). Vagina with essentially parallel sides, or sides slightly constricted at back of chamber (
<figureCitation captionStart="Figs" captionStartId="6.[184,234,1877,1901]" captionTargetBox="[307,1282,629,1750]" captionTargetId="figure-176@6.[297,1291,625,1769]" captionTargetPageId="6" captionText="Figs. 21‐27. Togoperla limbata. 21. Head and pronotum, 22. Male terminalia, 23. Male hemitergum, dorsal, 24. Male hemitergum, lateral, 25. Aedeagus, lateral aspect, 26. Female terminalia, 27. Vagina." figureDoi="http://doi.org/10.5281/zenodo.4754853" httpUri="https://zenodo.org/record/4754853/files/figure.png" pageId="7" pageNumber="215">Fig. 27</figureCitation>
); accessory glands about half as long as vagina.
</paragraph>
<paragraph blockId="7.[184,776,684,1953]" lastBlockId="7.[812,1404,287,1106]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[184,235,1929,1953]" pageId="7" pageNumber="215">Egg.</emphasis>
Length
<quantity box="[345,507,1929,1953]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="7" pageNumber="215" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38‐0.42 mm</quantity>
, width
<quantity box="[609,772,1929,1953]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="7" pageNumber="215" unit="mm" value="0.325" valueMax="0.34" valueMin="0.31">0.31‐0.34 mm</quantity>
. Collar expressed as a small dark brown disc. Micropyles oriented almost perpendicular to long axis of egg; micropylar row set almost halfway between equator and pole.
</paragraph>
</subSubSection>
<subSubSection box="[812,1256,425,450]" pageId="7" pageNumber="215" type="distribution">
<paragraph blockId="7.[812,1404,287,1106]" box="[812,1256,425,450]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[812,966,425,449]" pageId="7" pageNumber="215">Distribution.</emphasis>
Known only from
<collectingCountry box="[1184,1252,426,449]" name="Japan" pageId="7" pageNumber="215">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="215" type="discussion">
<paragraph blockId="7.[812,1404,287,1106]" pageId="7" pageNumber="215">
<emphasis bold="true" box="[812,924,461,485]" pageId="7" pageNumber="215">Remarks.</emphasis>
<bibRefCitation author="Sivec, I., B. P." box="[929,1122,461,485]" pageId="7" pageNumber="215" pagination="1 - 66" refId="ref8696" refString="Sivec, I., B. P. Stark &amp; S. Uchida. 1988. Synopsis of the world genera of Perlinae (Plecoptera: Perlidae). Scopolia, 16: 1 - 66." type="journal article" year="1988">Sivec et al. (1988)</bibRefCitation>
proposed the synonymy of
<taxonomicName authorityName="Okamoto" authorityYear="1912" box="[840,986,495,519]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="kawamurae">
<emphasis box="[840,986,495,519]" italics="true" pageId="7" pageNumber="215">T. kawamurae</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Okamoto, S. Uchida" authorityYear="1912" box="[1040,1196,496,519]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="matsumurae">
<emphasis box="[1040,1196,496,519]" italics="true" pageId="7" pageNumber="215">T. matsumurae</emphasis>
</taxonomicName>
with
<taxonomicName authorityName="Pictet" authorityYear="1841" box="[1257,1361,495,519]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="limbata">
<emphasis box="[1257,1361,495,519]" italics="true" pageId="7" pageNumber="215">T. limbata</emphasis>
</taxonomicName>
but omitted supporting data.
<bibRefCitation author="Uchida, S." box="[1114,1275,530,554]" pageId="7" pageNumber="215" refId="ref8766" refString="Uchida, S. 1990. A revision of the Japanese Perlidae (Insecta: Plecoptera), with special reference to their phylogeny. Unpublished PhD Dissertation, Tokyo Metropolitan University. 228 pp." type="book" year="1990">Uchida (1990)</bibRefCitation>
supported these synonymies in his thesis, but this has remained unpublished. We accept and validate the
<typeStatus box="[1299,1404,599,623]" pageId="7" pageNumber="215">lectotype</typeStatus>
for
<emphasis box="[857,1029,634,657]" italics="true" pageId="7" pageNumber="215">
<taxonomicName authorityName="Okamoto, S. Uchida" authorityYear="1912" box="[857,1025,634,657]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="matsumurae">T. matsumurae</taxonomicName>
,
</emphasis>
previously selected by
<bibRefCitation author="Uchida, S." pageId="7" pageNumber="215" refId="ref8766" refString="Uchida, S. 1990. A revision of the Japanese Perlidae (Insecta: Plecoptera), with special reference to their phylogeny. Unpublished PhD Dissertation, Tokyo Metropolitan University. 228 pp." type="book" year="1990">Uchida (1990)</bibRefCitation>
in his thesis, which bears label data of, Haki, 8/10,
<taxonomicName authority="Okam." authorityName="Okamoto, S. Uchida" authorityYear="1912" box="[877,1164,702,726]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="matsumurae">
<emphasis box="[877,1074,702,726]" italics="true" pageId="7" pageNumber="215">Perla matsumurae</emphasis>
Okam.
</taxonomicName>
, Sesuji‐ kawagera, &amp;
<typeStatus box="[812,927,738,762]" pageId="7" pageNumber="215">Lectotype</typeStatus>
,
<taxonomicName authority="Okamoto, S. Uchida" authorityName="Okamoto, S. Uchida" authorityYear="1912" box="[937,1356,737,761]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="matsumurae">
<emphasis box="[937,1127,737,761]" italics="true" pageId="7" pageNumber="215">Perla matsumurae</emphasis>
Okamoto, S. Uchida
</taxonomicName>
det. 1984, HU, and is in the
<collectingRegion box="[1149,1263,772,796]" country="Japan" name="Hokkaido" pageId="7" pageNumber="215">Hokkaido</collectingRegion>
University collection, Sapporo.
<bibRefCitation author="Uchida, S." box="[1048,1208,806,830]" pageId="7" pageNumber="215" refId="ref8766" refString="Uchida, S. 1990. A revision of the Japanese Perlidae (Insecta: Plecoptera), with special reference to their phylogeny. Unpublished PhD Dissertation, Tokyo Metropolitan University. 228 pp." type="book" year="1990">Uchida (1990)</bibRefCitation>
also described a second Japanese
<taxonomicName box="[1006,1199,841,865]" class="Insecta" family="Perlidae" genus="Togoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[1006,1109,841,865]" italics="true" pageId="7" pageNumber="215">Togoperla</emphasis>
species
</taxonomicName>
, generally similar to
<emphasis box="[847,967,875,899]" italics="true" pageId="7" pageNumber="215">
<taxonomicName authorityName="Pictet" authorityYear="1841" box="[847,963,875,899]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="limbata">T. limbata</taxonomicName>
,
</emphasis>
but differing in details of aedeagal armature, however this description has not been published and we have seen no material. Male hemiterga of this species are generally similar to those of
<emphasis box="[907,1165,1013,1038]" italics="true" pageId="7" pageNumber="215">
<taxonomicName baseAuthorityName="Navas" baseAuthorityYear="1926" box="[907,1031,1014,1038]" class="Insecta" family="Perlidae" genus="Togoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="fortunati">T. fortunati</taxonomicName>
,
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" baseAuthorityName="Navas" baseAuthorityYear="1934" box="[1040,1165,1013,1037]" class="Insecta" family="Perlidae" genus="Togoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="poilanina">T. poilanina</taxonomicName>
</emphasis>
and others, however it is easily distinguished by virtue of geography and by the pattern of armature on the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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