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<document id="F6436CD932C0A34598026B9213B57FAA" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A60E0617FD96FB97FE51E587" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba vethi" docType="treatment" docVersion="9" lastPageNumber="62" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="60" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7ABAC4A7DFE06795CBA6E835888C3933">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="2946450CFC619DD2D5037542A7F156FE">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="12BD0F0A494FE6FCCAAD7F96F3EC0EDA">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="17DAD596F15B072C34E6C966BA179309">Coca-Abia, María Milagro</mods:namePart>
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<mods:title id="3124BC67B4B259CB27418DF9A6ABE0F0">European Journal of Taxonomy</mods:title>
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<treatment id="03B56743A60E0617FD96FB97FE51E587" ID-DOI="http://doi.org/10.5281/zenodo.4770378" ID-GBIF-Taxon="182237248" ID-Zenodo-Dep="4770378" LSID="urn:lsid:plazi:treatment:03B56743A60E0617FD96FB97FE51E587" httpUri="http://treatment.plazi.org/id/03B56743A60E0617FD96FB97FE51E587" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
<subSubSection id="C30685DEA60E0611FD96FB97FC24E068" box="[621,966,1086,1113]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="8BA3D655A60E0611FD96FB97FC24E068" blockId="59.[621,966,1086,1148]" box="[621,966,1086,1113]" pageId="59" pageNumber="60">
<heading id="D0EB6139A60E0611FD96FB97FC24E068" box="[621,966,1086,1113]" centered="true" fontSize="11" level="2" pageId="59" pageNumber="60" reason="2">
<taxonomicName id="4C1CADD6A60E0611FD96FB97FC24E068" authority="(Moser, 1917)" baseAuthorityName="Moser" baseAuthorityYear="1917" box="[621,966,1086,1113]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A60E0611FD96FB97FCFBE068" bold="true" box="[621,793,1086,1113]" italics="true" pageId="59" pageNumber="60">Miridiba vethi</emphasis>
(
<bibRefCitation id="EF8DABA4A60E0611FCD3FB96FC5CE068" author="Moser J." box="[808,958,1087,1113]" pageId="59" pageNumber="60" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA60E0611FD23FBCBFCB9E04D" box="[728,859,1122,1148]" pageId="59" pageNumber="60" type="description">
<paragraph id="8BA3D655A60E0611FD23FBCBFCB9E04D" blockId="59.[621,966,1086,1148]" box="[728,859,1122,1148]" pageId="59" pageNumber="60">
<figureCitation id="1327CAD0A60E0611FD23FBCBFCB9E04D" box="[728,859,1122,1148]" captionStart="Figs 77–78" captionStartId="60.[191,243,1956,1982]" captionTargetBox="[284,1322,1149,1894]" captionTargetId="figure-218@60.[264,1323,1129,1916]" captionTargetPageId="60" captionText="Figs 77–78. Miridiba vethi (Moser, 1917), lectotype, ♂ (MFNB). 77. Habitus in dorsal view. 78. Parameres in dorsal (A) and lateral (B) views. Scale bars: 77 = 5.0 mm; 78 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770392" httpUri="https://zenodo.org/record/4770392/files/figure.png" pageId="59" pageNumber="60">Figs 77–78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA60E0611FF46FB0AFC69E131" pageId="59" pageNumber="60" type="reference_group">
<paragraph id="8BA3D655A60E0611FF46FB0AFC3AE08F" blockId="59.[189,984,1187,1214]" box="[189,984,1187,1214]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB0CF17BA60E0611FF46FB0AFC3AE08F" box="[189,984,1187,1214]" pageId="59" pageNumber="60">
<treatmentCitation id="0ABDF044A60E0611FF46FB0AFDAFE08F" author="Moser J." box="[189,589,1187,1214]" page="90" pageId="59" pageNumber="60" year="1917">
<taxonomicName id="4C1CADD6A60E0611FF46FB0AFDAFE08F" authority="Moser, 1917: 90" authorityName="Moser" authorityPageNumber="90" authorityYear="1917" box="[189,589,1187,1214]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A60E0611FF46FB0AFE64E08C" box="[189,390,1187,1213]" italics="true" pageId="59" pageNumber="60">Holotrichia vethi</emphasis>
<bibRefCitation id="EF8DABA4A60E0611FE75FB0DFDAFE08F" author="Moser J." box="[398,589,1188,1214]" pageId="59" pageNumber="60" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917: 90</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A60E0611FDA6FB0DFD73E08F" box="[605,657,1188,1214]" pageId="59" pageNumber="60">type</typeStatus>
loc.: Batoe Bara,
<collectingRegion id="49D818B7A60E0611FC9EFB0DFC2EE08F" box="[869,972,1188,1214]" country="Indonesia" name="Sumatera Utara" pageId="59" pageNumber="60">Sumatra</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A60E0611FF46FB4CFC69E131" blockId="59.[189,907,1253,1280]" box="[189,907,1253,1280]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="AB0CF17BA60E0611FF46FB4CFC69E131" box="[189,907,1253,1280]" pageId="59" pageNumber="60">
<taxonomicName id="4C1CADD6A60E0611FF46FB4CFE84E0CE" baseAuthorityName="Moser" baseAuthorityYear="1917" box="[189,358,1253,1279]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A60E0611FF46FB4CFE84E0CE" box="[189,358,1253,1279]" italics="true" pageId="59" pageNumber="60">Miridiba vethi</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A60E0611FE78FB4CFD8BE0CE" author="Matsumoto T." box="[387,617,1253,1279]" page="1" pageId="59" pageNumber="60" year="2017">
<bibRefCitation id="EF8DABA4A60E0611FE78FB4CFD8BE0CE" author="Matsumoto T." box="[387,617,1253,1279]" pageId="59" pageNumber="60" pagination="1 - 4" refId="ref45176" refString="Matsumoto T. 2017. Taxonomic notes on Holotrichia vethi Moser from Sumaka (Scarabaeidae, Melolonthinae, Melolonthini). Kogane, Tokyo 19: 1 - 4." type="journal article" year="2017">Matsumoto 2017: 1</bibRefCitation>
</treatmentCitation>
, figs 1–6 (combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA60E0616FF46FA8FFC8AE576" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="8BA3D655A60E0611FF46FA8FFED4E171" blockId="59.[189,310,1318,1344]" box="[189,310,1318,1344]" pageId="59" pageNumber="60">
<heading id="D0EB6139A60E0611FF46FA8FFED4E171" bold="true" box="[189,310,1318,1344]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="B9680A47A60E0611FF46FA8FFED4E171" bold="true" box="[189,310,1318,1344]" pageId="59" pageNumber="60">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A60E0616FF46FAFCFC8AE576" blockId="59.[186,1401,1365,1990]" lastBlockId="60.[189,1399,265,327]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
External morphology of adult (
<figureCitation id="1327CAD0A60E0611FDE8FAFCFD87E15E" box="[531,613,1365,1391]" captionStart="Figs 77–78" captionStartId="60.[191,243,1956,1982]" captionTargetBox="[284,1322,1149,1894]" captionTargetId="figure-218@60.[264,1323,1129,1916]" captionTargetPageId="60" captionText="Figs 77–78. Miridiba vethi (Moser, 1917), lectotype, ♂ (MFNB). 77. Habitus in dorsal view. 78. Parameres in dorsal (A) and lateral (B) views. Scale bars: 77 = 5.0 mm; 78 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770392" httpUri="https://zenodo.org/record/4770392/files/figure.png" pageId="59" pageNumber="60">Fig. 77</figureCitation>
). Dorsal surface glabrous.Clypeus concave, shorter than frons, oblique laterally, anterior edge strongly emarginate at middle. Frons with sharp carina. Anterior pronotal margin glabrous, strongly flanged, narrowed at middle and at lateral ends; posterior margin glabrous, moderately flanged; lateral margins smooth, glabrous, moderately elevated at anterior half; anterior angles almost right and rounded, not projected forward; posterior angles obtuse and rounded. Pronotal surface densely punctate, each puncture with a tiny bulge hardly visible. Scutellum sparsely punctate. Elytral surface with punctures regularly distributed; each puncture with a tiny bulge hardly visible; apical callus well defined; epipleuron glabrous with membrane. Foretibia with a moderately developed dorsal carina; insertion of inner spur closer to third outer tooth than second one. Meso- and metatibia with transverse carina interrupted and complete respectively; inner margin of dorsal surface with few spines. Pygidium irregularly punctate; punctures thinner at anterior margin; surface without conspicuous pubescence, at most with tiny setae in each puncture hardly visible, apical margin with long pubescence. Ventrites 2, 3 and 4 without conspicuous pubescence, with small scales hardly visible. Ventrite 5 moderately depressed at posterior half with punctures uniformly distributed and long pubescence irregularly distributed on posterior half; anterior half without conspicuous pubescence. Ventrite 6 moderately bulging anteriorly; conspicuous pubescence irregularly distributed. Male genitalia (
<figureCitation id="1327CAD0A60E0611FED5F8CCFE64E34E" box="[302,390,1893,1919]" captionStart="Figs 77–78" captionStartId="60.[191,243,1956,1982]" captionTargetBox="[284,1322,1149,1894]" captionTargetId="figure-218@60.[264,1323,1129,1916]" captionTargetPageId="60" captionText="Figs 77–78. Miridiba vethi (Moser, 1917), lectotype, ♂ (MFNB). 77. Habitus in dorsal view. 78. Parameres in dorsal (A) and lateral (B) views. Scale bars: 77 = 5.0 mm; 78 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770392" httpUri="https://zenodo.org/record/4770392/files/figure.png" pageId="59" pageNumber="60">Fig. 78</figureCitation>
): dorsal branch of parameres thin and symmetrical, with sutural scar completely fused, triangular at proximal end, tapered with a rounded tip curving downward at distal end. Ventral branches with elevations as waves at outer margin, apices upward. Endophallus and temones not studied. Female genitalia: median oviduct with wrinkled and hardened epithelium; elongate plates of gonopore fold with sensillae; conspicuous vestigial sternites placed ventrally at gonopore.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6090616FF46FEC4FE46E723" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="8BA3D655A6090616FF46FEC4FE48E5B6" blockId="60.[189,426,365,391]" box="[189,426,365,391]" pageId="60" pageNumber="61">
<heading id="D0EB6139A6090616FF46FEC4FE48E5B6" bold="true" box="[189,426,365,391]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B9680A47A6090616FF46FEC4FE48E5B6" bold="true" box="[189,426,365,391]" pageId="60" pageNumber="61">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6090616FF11FE35FD9CE611" blockId="60.[189,1399,412,544]" pageId="60" pageNumber="61">
<materialsCitation id="3B74DC08A6090616FF11FE35FD9CE611" ID-GBIF-Occurrence="3114121306" collectionCode="MFNB" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="Sumatra on Batoe Bara" municipality="Sumatra on Batoe Bara" pageId="60" pageNumber="61" specimenCode="MNHUB 2016" specimenCount="1" specimenCount-male="1" stateProvince="SUMATRA" typeStatus="lectotype">
<emphasis id="B9680A47A6090616FF11FE35FE84E587" bold="true" box="[234,358,412,438]" pageId="60" pageNumber="61">
<typeStatus id="54A768F7A6090616FF11FE35FE84E587" box="[234,358,412,438]" pageId="60" pageNumber="61" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
<collectingRegion id="49D818B7A6090616FF46FE16FEAAE5E8" box="[189,328,447,473]" country="Indonesia" name="Sumatera Utara" pageId="60" pageNumber="61">SUMATRA</collectingRegion>
•
<specimenCount id="9D1A1DDCA6090616FE9EFE69FE9CE5EB" box="[357,382,448,474]" pageId="60" pageNumber="61" type="male">♂</specimenCount>
; “
<collectingMunicipality id="6BC74C2FA6090616FE6CFE16FD5AE5E8" box="[407,696,447,474]" pageId="60" pageNumber="61">Sumatra on Batoe Bara</collectingMunicipality>
”; “
<taxonomicName id="4C1CADD6A6090616FD25FE16FC05E5EB" authority="Mos" authorityName="Moser" authorityYear="1917" box="[734,999,447,474]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FD25FE16FC48E5E8" box="[734,938,447,473]" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
Mos
</taxonomicName>
Type ♂”; “Syntypus
<taxonomicName id="4C1CADD6A6090616FB15FE16FE7BE5CC" authority="Moser, 1917" authorityName="Moser" authorityYear="1917" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FB15FE16FF14E5CD" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
<bibRefCitation id="EF8DABA4A6090616FEFAFE4AFE7BE5CC" author="Moser J." box="[257,409,483,509]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA6090616FDCFFE4AFD0CE5CD" box="[564,750,482,509]" pageId="60" pageNumber="61">MNHUB 2016</specimenCode>
”; “Lectotype
<taxonomicName id="4C1CADD6A6090616FC6EFE4BFB15E5CC" authority="(Moser, 1917)" baseAuthorityName="Moser" baseAuthorityYear="1917" box="[917,1271,482,509]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FC6EFE4BFBA0E5CD" box="[917,1090,482,508]" italics="true" pageId="60" pageNumber="61">Miridiba vethi</emphasis>
(
<bibRefCitation id="EF8DABA4A6090616FBAEFE4AFB0DE5CC" author="Moser J." box="[1109,1263,483,509]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A6090616FAF8FE4BFF0CE611" pageId="60" pageNumber="61">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A6090616FEE8FDAFFE70E611" box="[275,402,518,544]" pageId="60" pageNumber="61">Coca-Abia</determinerName>
det. 2019”;
<collectionCode id="ED0D4E90A6090616FDD8FDAFFD98E611" box="[547,634,518,544]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="60" pageNumber="61">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A6090616FF11FDEEFE46E723" blockId="60.[188,1399,583,786]" pageId="60" pageNumber="61">
<materialsCitation id="3B74DC08A6090616FF11FDEEFDA8E6FA" ID-GBIF-Occurrence="3114121312" collectionCode="MFNB" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="SUMATRA" pageId="60" pageNumber="61" specimenCode="MNHUB 2016" specimenCount="1" specimenCount-male="1" stateProvince="SUMATRA" typeStatus="paralectotype">
<emphasis id="B9680A47A6090616FF11FDEEFE40E650" bold="true" box="[234,418,583,609]" pageId="60" pageNumber="61">
<typeStatus id="54A768F7A6090616FF11FDEEFE40E650" box="[234,418,583,609]" pageId="60" pageNumber="61" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
(here indicated)
<collectingRegion id="49D818B7A6090616FF46FDC3FEAAE6B5" box="[189,328,618,644]" country="Indonesia" name="Sumatera Utara" pageId="60" pageNumber="61">SUMATRA</collectingRegion>
•
<specimenCount id="9D1A1DDCA6090616FEA4FDC3FE69E6B4" box="[351,395,618,645]" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
(without head and pronotum); “Sumatra on Batoe Bara”; “Syntypus
<taxonomicName id="4C1CADD6A6090616FB55FDC3FEB3E699" authority="Moser, 1917" authorityName="Moser" authorityYear="1917" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FB55FDC3FA94E6B5" box="[1198,1398,618,644]" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
<bibRefCitation id="EF8DABA4A6090616FF47FD27FEB3E699" author="Moser J." box="[188,337,654,680]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA6090616FE1FFD27FD78E696" box="[484,666,653,680]" pageId="60" pageNumber="61">MNHUB 2016</specimenCode>
”; “Paralectotype
<taxonomicName id="4C1CADD6A6090616FC93FD24FB23E699" authority="(Moser, 1917)" baseAuthorityName="Moser" baseAuthorityYear="1917" box="[872,1217,653,680]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FC93FD24FBF0E696" box="[872,1042,653,679]" italics="true" pageId="60" pageNumber="61">Miridiba vethi</emphasis>
(
<bibRefCitation id="EF8DABA4A6090616FBDAFD27FB5BE699" author="Moser J." box="[1057,1209,654,680]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A6090616FB32FD24FA95E699" box="[1225,1399,653,680]" pageId="60" pageNumber="61">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A6090616FF24FD18FEBCE6FA" box="[223,350,689,715]" pageId="60" pageNumber="61">Coca-Abia</determinerName>
det. 2019”;
<collectionCode id="ED0D4E90A6090616FE0CFD18FDA8E6FA" box="[503,586,689,715]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="60" pageNumber="61">MFNB</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A6090616FD93FD18FE46E723" ID-GBIF-Occurrence="3114121383" collectionCode="MFNB" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="Holotrichia vethi Mos" municipality="Holotrichia vethi Mos" pageId="60" pageNumber="61" specimenCode="MNHUB 2016" specimenCount="1" specimenCount-female="1" stateProvince="Sumatra">
<specimenCount id="9D1A1DDCA6090616FD93FD18FD7AE6FA" box="[616,664,689,715]" pageId="60" pageNumber="61" type="female">1 ♀</specimenCount>
; “Coll Nonfried
<collectingRegion id="49D818B7A6090616FC9CFD18FC2DE6FA" box="[871,975,689,715]" country="Indonesia" name="Sumatera Utara" pageId="60" pageNumber="61">Sumatra</collectingRegion>
”; “
<taxonomicName id="4C1CADD6A6090616FC08FD18FB1FE6FA" authority="Mos" authorityName="Moser" authorityYear="1917" box="[1011,1277,689,715]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<collectingMunicipality id="6BC74C2FA6090616FC08FD18FB1FE6FA" box="[1011,1277,689,715]" pageId="60" pageNumber="61">
<emphasis id="B9680A47A6090616FC08FD18FB5DE6FA" box="[1011,1215,689,715]" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
Mos
</collectingMunicipality>
</taxonomicName>
Type ♀”; “Syntypus
<taxonomicName id="4C1CADD6A6090616FEBAFD7DFD49E6DF" authority="Moser, 1917" authorityName="Moser" authorityYear="1917" box="[321,683,724,751]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FEBAFD7DFDEEE6DF" box="[321,524,724,750]" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
<bibRefCitation id="EF8DABA4A6090616FDEDFD7CFD49E6DF" author="Moser J." box="[534,683,724,751]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA6090616FCB9FD7CFC18E6DF" box="[834,1018,724,751]" pageId="60" pageNumber="61">MNHUB 2016</specimenCode>
”; “
<determinerName id="7E7F3A91A6090616FBE7FD7DFB2FE6DF" box="[1052,1229,724,750]" pageId="60" pageNumber="61">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A6090616FB0CFD7DFA94E6DF" box="[1271,1398,724,750]" pageId="60" pageNumber="61">Coca-Abia</determinerName>
det. 2019”;
<collectionCode id="ED0D4E90A6090616FEB2FD51FE42E723" box="[329,416,760,786]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="60" pageNumber="61">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6090617FF46FC90FD11E577" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="discussion">
<paragraph id="8BA3D655A6090616FF46FC90FECDE762" blockId="60.[189,303,825,851]" box="[189,303,825,851]" pageId="60" pageNumber="61">
<heading id="D0EB6139A6090616FF46FC90FECDE762" bold="true" box="[189,303,825,851]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="B9680A47A6090616FF46FC90FECDE762" bold="true" box="[189,303,825,851]" pageId="60" pageNumber="61">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6090617FF46FCC1FD11E577" blockId="60.[189,1399,871,1038]" lastBlockId="61.[189,1398,265,327]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<bibRefCitation id="EF8DABA4A6090616FF46FCC1FE82E7B0" author="Moser J." box="[189,352,871,898]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser (1917)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A6090616FE1DFCCEFD53E7B0" authorityName="Moser" authorityYear="1917" box="[486,689,871,897]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FE1DFCCEFD53E7B0" box="[486,689,871,897]" italics="true" pageId="60" pageNumber="61">Holotrichia vethi</emphasis>
</taxonomicName>
from
<collectingRegion id="49D818B7A6090616FD00FCCEFCBFE7B0" box="[763,861,871,897]" country="Indonesia" name="Sumatera Utara" pageId="60" pageNumber="61">Sumatra</collectingRegion>
(Batoe Bara).
<bibRefCitation id="EF8DABA4A6090616FBEBFCC1FB0CE7B0" author="Matsumoto T." box="[1040,1262,871,898]" pageId="60" pageNumber="61" pagination="1 - 4" refId="ref45176" refString="Matsumoto T. 2017. Taxonomic notes on Holotrichia vethi Moser from Sumaka (Scarabaeidae, Melolonthinae, Melolonthini). Kogane, Tokyo 19: 1 - 4." type="journal article" year="2017">Matsumoto (2017)</bibRefCitation>
transferred this species to
<taxonomicName id="4C1CADD6A6090616FE89FC23FE38E795" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[370,474,906,932]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6090616FE89FC23FE38E795" box="[370,474,906,932]" italics="true" pageId="60" pageNumber="61">Miridiba</emphasis>
</taxonomicName>
after studying
<specimenCount id="9D1A1DDCA6090616FD6FFC23FCA2E794" box="[660,832,906,933]" pageId="60" pageNumber="61" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
(
<specimenCount id="9D1A1DDCA6090616FCA8FC23FC2DE795" box="[851,975,906,932]" pageId="60" pageNumber="61" type="male">two males</specimenCount>
and
<specimenCount id="9D1A1DDCA6090616FBF5FC22FB7AE794" box="[1038,1176,906,933]" pageId="60" pageNumber="61" type="female">one female</specimenCount>
), and no
<typeStatus id="54A768F7A6090616FAF0FC23FA95E795" box="[1291,1399,906,932]" pageId="60" pageNumber="61" type="lectotype">lectotype</typeStatus>
was designated. After studying these
<specimenCount id="9D1A1DDCA6090616FD8DFC04FCFDE7F9" box="[630,799,941,968]" pageId="60" pageNumber="61" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
(MFNB), we agree with the transfer of
<taxonomicName id="4C1CADD6A6090616FB00FC07FABAE7F6" authorityName="Moser" authorityYear="1917" box="[1275,1368,941,967]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FB00FC07FABAE7F6" box="[1275,1368,941,967]" italics="true" pageId="60" pageNumber="61">H. vethi</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A6090616FF45FC78FEC4E7DA" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[190,294,977,1003]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6090616FF45FC78FEC4E7DA" box="[190,294,977,1003]" italics="true" pageId="60" pageNumber="61">Miridiba</emphasis>
</taxonomicName>
and designate the
<typeStatus id="54A768F7A6090616FDF8FC78FD8DE7DA" box="[515,623,977,1003]" pageId="60" pageNumber="61" type="lectotype">lectotype</typeStatus>
and
<specimenCount id="9D1A1DDCA6090616FD51FC78FC69E7DA" box="[682,907,977,1003]" pageId="60" pageNumber="61" type="generic" typeStatus="paralectotypes">two paralectotypes</specimenCount>
. Moreover, we confirm the similarities of
<taxonomicName id="4C1CADD6A6090616FF46FC5CFEFDE03F" authorityName="Moser" authorityYear="1917" box="[189,287,1012,1038]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FF46FC5CFEFDE03F" box="[189,287,1012,1038]" italics="true" pageId="60" pageNumber="61">M. vethi</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A6090616FE9CFC5CFE09E03F" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[359,491,1012,1038]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="laosana">
<emphasis id="B9680A47A6090616FE9CFC5CFE09E03F" box="[359,491,1012,1038]" italics="true" pageId="60" pageNumber="61">M. laosana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1CADD6A6090616FE07FC5CFD76E03F" baseAuthorityName="Matsumoto" baseAuthorityYear="2016" box="[508,660,1012,1038]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="B9680A47A6090616FE07FC5CFD76E03F" box="[508,660,1012,1038]" italics="true" pageId="60" pageNumber="61">M. bidentata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1CADD6A6090616FD5FFC5CFCBEE03F" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[676,860,1012,1038]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="malaccensis">
<emphasis id="B9680A47A6090616FD5FFC5CFCBEE03F" box="[676,860,1012,1038]" italics="true" pageId="60" pageNumber="61">M. malaccensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1CADD6A6090616FC60FC5CFAD9E03F" authority="(Matsumoto 2017)" baseAuthorityName="Matsumoto" baseAuthorityYear="2017" box="[923,1339,1012,1038]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B9680A47A6090616FC60FC5CFBB1E03F" box="[923,1107,1012,1038]" italics="true" pageId="60" pageNumber="61">M. waterstradti</emphasis>
(
<bibRefCitation id="EF8DABA4A6090616FB98FC5DFAD6E03F" author="Matsumoto T." box="[1123,1332,1012,1038]" pageId="60" pageNumber="61" pagination="1 - 4" refId="ref45176" refString="Matsumoto T. 2017. Taxonomic notes on Holotrichia vethi Moser from Sumaka (Scarabaeidae, Melolonthinae, Melolonthini). Kogane, Tokyo 19: 1 - 4." type="journal article" year="2017">Matsumoto 2017</bibRefCitation>
)
</taxonomicName>
. Just like the four species,
<taxonomicName id="4C1CADD6A6080617FE44FEA3FDC3E512" authorityName="Moser" authorityYear="1917" box="[447,545,265,291]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6080617FE44FEA3FDC3E512" box="[447,545,265,291]" italics="true" pageId="61" pageNumber="62">M. vethi</emphasis>
</taxonomicName>
shows features that characterize morphotype IV “
<emphasis id="B9680A47A6080617FB8CFEA0FB3EE512" box="[1143,1244,265,291]" italics="true" pageId="61" pageNumber="62">Bilobata</emphasis>
”. Hence, we consider it to belong to this genital morphotype.
</paragraph>
</subSubSection>
<caption id="DF6386DDA6090616FF44F80DFB34E3D3" ID-DOI="http://doi.org/10.5281/zenodo.4770392" ID-Zenodo-Dep="4770392" httpUri="https://zenodo.org/record/4770392/files/figure.png" pageId="60" pageNumber="61" startId="60.[191,243,1956,1982]" targetBox="[284,1322,1149,1894]" targetPageId="60">
<paragraph id="8BA3D655A6090616FF44F80DFB34E3D3" blockId="60.[189,1399,1956,2018]" pageId="60" pageNumber="61">
<emphasis id="B9680A47A6090616FF44F80DFEB6E38F" bold="true" box="[191,340,1956,1982]" pageId="60" pageNumber="61">Figs 77–78.</emphasis>
<taxonomicName id="4C1CADD6A6090616FE98F80DFD3BE38F" authority="(Moser, 1917)" baseAuthorityName="Moser" baseAuthorityYear="1917" box="[355,729,1956,1982]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="vethi">
<emphasis id="B9680A47A6090616FE98F80DFDF7E38F" box="[355,533,1956,1982]" italics="true" pageId="60" pageNumber="61">Miridiba vethi</emphasis>
(
<bibRefCitation id="EF8DABA4A6090616FDD5F80DFD2DE38F" author="Moser J." box="[558,719,1956,1982]" pageId="60" pageNumber="61" pagination="54 - 97" refId="ref45427" refString="Moser J. 1917. Beitrag zur Kenntnis der Melolonthiden. (Col.) VII. Stettiner entomologische Zeitung 78: 54 - 97." type="journal article" year="1917">Moser, 1917</bibRefCitation>
)
</taxonomicName>
, lectotype, ♂ (MFNB).
<emphasis id="B9680A47A6090616FBECF80DFBD6E38F" bold="true" box="[1047,1076,1956,1982]" pageId="60" pageNumber="61">77</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A6090616FF46F86EFF38E3D0" bold="true" box="[189,218,1991,2017]" pageId="60" pageNumber="61">78</emphasis>
. Parameres in dorsal (A) and lateral (B) views. Scale bars: 77 = 5.0 mm; 78 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA6080617FF46FEC4FE51E587" box="[189,435,365,438]" pageId="61" pageNumber="62" type="distribution">
<paragraph id="8BA3D655A6080617FF46FEC4FEB4E5B6" blockId="61.[189,342,365,391]" box="[189,342,365,391]" pageId="61" pageNumber="62">
<heading id="D0EB6139A6080617FF46FEC4FEB4E5B6" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B9680A47A6080617FF46FEC4FEB4E5B6" bold="true" box="[189,342,365,391]" pageId="61" pageNumber="62">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6080617FF46FE35FE51E587" blockId="61.[189,435,412,438]" box="[189,435,412,438]" pageId="61" pageNumber="62">
<collectingCountry id="F30B96C5A6080617FF46FE35FECDE587" box="[189,303,412,438]" name="Indonesia" pageId="61" pageNumber="62">Indonesia</collectingCountry>
(
<collectingRegion id="49D818B7A6080617FEC4FE35FE4AE587" box="[319,424,412,438]" country="Indonesia" name="Sumatera Utara" pageId="61" pageNumber="62">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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