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<mods:title id="C0645A7ED17D0F39AB589CD974111297">New and little known species of the genus Martiodrilus Michaelsen, 1936 from French Guiana (Oligochaeta, Glossoscolecidae)</mods:title>
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<taxonomicName id="4C184D42FF9BFFF51BBFF8BAFD94F8EF" authority="Michaelsen, 1936" authorityName="Michaelsen" authorityYear="1936" box="[151,527,1879,1905]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="0" pageNumber="57" phylum="Annelida" rank="genus">
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<emphasis id="B96CEAD3FF9BFFF51BBFF8BAFEABF8EF" bold="true" box="[151,304,1879,1905]" italics="true" pageId="0" pageNumber="57">Martiodrilus</emphasis>
<bibRefCitation id="EF894B30FF9BFFF51A1FF8BAFD94F8EF" author="Michaelsen" box="[311,527,1879,1905]" pageId="0" pageNumber="57" refString="Michaelsen, W. (1936) On the genus Thamnodrilus Beddard. Proceedings of the Zoological Society of London, 1936, 1171 - 1173." type="proceedings paper" year="1936">Michaelsen, 1936</bibRefCitation>
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<taxonomicName id="4C184D42FF9BFFF51BBFF873FD82F82A" authority="Michaelsen, 1936: 1172" authorityName="Michaelsen" authorityPageNumber="1172" authorityYear="1936" box="[151,537,1950,1972]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="0" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9BFFF51BBFF873FE82F82A" box="[151,281,1950,1972]" italics="true" pageId="0" pageNumber="57">Martiodrilus</emphasis>
<treatmentCitation id="0AB910D0FF9BFFF51A37F873FD82F82A" author="Michaelsen" box="[287,537,1950,1972]" page="1172" pageId="0" pageNumber="57" year="1936">
<bibRefCitation id="EF894B30FF9BFFF51A37F873FE43F82A" author="Michaelsen" box="[287,472,1950,1972]" pageId="0" pageNumber="57" refString="Michaelsen, W. (1936) On the genus Thamnodrilus Beddard. Proceedings of the Zoological Society of London, 1936, 1171 - 1173." type="proceedings paper" year="1936">Michaelsen, 1936</bibRefCitation>
: 1172
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.,
<treatmentCitation id="0AB910D0FF9BFFF51900F873FD4FF82A" author="Righi" box="[552,724,1950,1972]" page="512" pageId="0" pageNumber="57" year="1995">
<bibRefCitation id="EF894B30FF9BFFF51900F873FD05F82A" author="Righi" box="[552,670,1950,1972]" pageId="0" pageNumber="57" refString="Righi, G. (1995) Colombian earthworms. Studies on Tropical Andean Ecosystems, 4, 485 - 607." type="journal article" year="1995">Righi 1995</bibRefCitation>
: 512
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.,
<treatmentCitation id="0AB910D0FF9BFFF519CBF873FC7EF82A" author="Zicsi" box="[739,997,1950,1972]" page="82" pageId="0" pageNumber="57" year="1997">
<bibRefCitation id="EF894B30FF9BFFF519CBF873FC27F82A" author="Zicsi" box="[739,956,1950,1972]" pageId="0" pageNumber="57" refString="Zicsi, A. &amp; Csuzdi, Cs. (1997) Uber wietere Riesenregenwurmer aus Ekuador. Regenwurmer aus Sudamerika, 28. (Oligochaeta). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 84, 81 - 103." type="journal article" year="1997">Zicsi &amp; Csuzdi 1997</bibRefCitation>
: 82
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.,
<treatmentCitation id="0AB910D0FF9BFFF518DCF873FB0DF82A" author="Zicsi" box="[1012,1174,1950,1972]" page="113" pageId="0" pageNumber="57" year="2001">
<bibRefCitation id="EF894B30FF9BFFF518DCF873FBFFF82A" author="Zicsi" box="[1012,1124,1950,1972]" pageId="0" pageNumber="57" refString="Zicsi, A. (2001) Revision der Untergattung Martiodrilus (Maipure Righi, 1995) (Oligochaeta: Glossoscolecidae). Regenwurm- er aus Sudamerika 33. Opuscula Zoologica Budapest, 33, 113 - 131." type="journal article" year="2001">Zicsi 2001</bibRefCitation>
: 113
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, Fragoso &amp; Brown 2007: 53.
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<taxonomicName id="4C184D42FF9BFFF51BBFF836FE5AF86F" authority="Beddard, 1887" authorityName="Beddard" authorityYear="1887" box="[151,449,2011,2033]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodrilus" kingdom="Animalia" order="Haplotaxida" pageId="0" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9BFFF51BBFF836FEBEF86F" box="[151,293,2011,2033]" italics="true" pageId="0" pageNumber="57">Thamnodrilus</emphasis>
<bibRefCitation id="EF894B30FF9BFFF51A03F836FE5AF86F" author="Beddard" box="[299,449,2011,2033]" pageId="0" pageNumber="57" refString="Beddard, F. E. (1887) On the structure of a new Genus of Lumbricidae (Thamnodrilus gulielmi). Proceedings of the Zoological Society of London, 1887, 154 - 163." type="proceedings paper" year="1887">Beddard, 1887</bibRefCitation>
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(part.):
<treatmentCitation id="0AB910D0FF9BFFF5193AF836FD63F86F" author="Michaelsen" box="[530,760,2011,2033]" page="434" pageId="0" pageNumber="57" year="1900">
<bibRefCitation id="EF894B30FF9BFFF5193AF836FD58F86F" author="Michaelsen" box="[530,707,2011,2033]" pageId="0" pageNumber="57" refString="Michaelsen, W. (1900) Oligochaeta. Das Tierreich, 11, 1 - 575." type="journal article" year="1900">Michaelsen 1900</bibRefCitation>
: 434
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.,
<treatmentCitation id="0AB910D0FF9BFFF5182EF83AFC7FF86F" author="Cernosvitov" box="[774,996,2007,2034]" page="18" pageId="0" pageNumber="57" year="1935">
<bibRefCitation id="EF894B30FF9BFFF5182EF83AFC24F86F" author="Cernosvitov" box="[774,959,2007,2034]" pageId="0" pageNumber="57" refString="Cernosvitov, L. (1935) Oligochaeten aus dem tropischen Sud-America. Capita Zoologica, 6, 3 - 37." type="journal article" year="1935">Černosvitov 1935</bibRefCitation>
: 18
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.
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<paragraph id="8BA736C1FF9AFFF41BBFFF75FE6BFF52" blockId="1.[151,1295,147,205]" pageId="1" pageNumber="58">
<treatmentCitationGroup id="AB0811EFFF9AFFF41BBFFF75FE6BFF52" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41BBFFF75FE51FF30" box="[151,458,152,174]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41BBFFF75FE51FF30" box="[151,458,152,174]" italics="true" pageId="1" pageNumber="58">Thamnodrilus (Thamnodrilus)</emphasis>
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(part.):
<treatmentCitation id="0AB910D0FF9AFFF41934FF75FD6EFF30" author="Michaelsen" box="[540,757,152,174]" page="86" pageId="1" pageNumber="58" year="1918">
<bibRefCitation id="EF894B30FF9AFFF41934FF75FD56FF33" author="Michaelsen" box="[540,717,152,174]" pageId="1" pageNumber="58" refString="Michaelsen, W. (1918) Die Lumbriciden, mit besonderer Berucksichtigung der bisher als Familie Glossoscolecidae zusammengefassten Unterfamilien. Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 41, 1 - 398." type="journal article" year="1918">Michaelsen 1918</bibRefCitation>
: 86
</treatmentCitation>
.,
<treatmentCitation id="0AB910D0FF9AFFF4182CFF7EFC7FFF30" author="Cernosvitov" box="[772,996,147,174]" page="53" pageId="1" pageNumber="58" year="1934">
<bibRefCitation id="EF894B30FF9AFFF4182CFF7EFC27FF33" author="Cernosvitov" box="[772,956,147,174]" pageId="1" pageNumber="58" refString="Cernosvitov, L. (1934) Les Oligochaetes de la Guyane Francaise et d'autres pays de l'Amerique du Sud. Bulletin de la Musee d'Histoire Naturelle de Paris, 6, 47 - 59." type="journal article" year="1934">Černosvitov 1934</bibRefCitation>
: 53
</treatmentCitation>
., Fragoso &amp; Brown 2007: 59.
<taxonomicName id="4C184D42FF9AFFF41BBFFF5AFE77FF52" authority="Gates, 1968: 14" authorityName="Gates" authorityPageNumber="14" authorityYear="1968" box="[151,492,182,205]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodriloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41BBFFF5AFEDFFF53" box="[151,324,183,205]" italics="true" pageId="1" pageNumber="58">Thamnodriloides</emphasis>
<treatmentCitation id="0AB910D0FF9AFFF41A62FF5AFE77FF52" author="Gates" box="[330,492,182,204]" page="14" pageId="1" pageNumber="58" year="1968">
<bibRefCitation id="EF894B30FF9AFFF41A62FF5AFE5DFF52" author="Gates" box="[330,454,182,204]" pageId="1" pageNumber="58" refString="Gates, G. E. (1968) On a glossoscolecid earthworm from Panama and its genus. Megadrilogica, 1 (1), 1 - 15." type="journal article" year="1968">Gates, 1968</bibRefCitation>
: 14
</treatmentCitation>
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.
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<emphasis id="B96CEAD3FF9AFFF41BBFFF14FE90FE8F" bold="true" box="[151,267,249,273]" pageId="1" pageNumber="58">Remarks.</emphasis>
The genus
<taxonomicName id="4C184D42FF9AFFF41AB1FF14FDB3FE8F" box="[409,552,249,273]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41AB1FF14FDB3FE8F" box="[409,552,249,273]" italics="true" pageId="1" pageNumber="58">Martiodrilus</emphasis>
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was described by
<bibRefCitation id="EF894B30FF9AFFF41820FF14FC44FE8F" author="Michaelsen" box="[776,991,249,273]" pageId="1" pageNumber="58" refString="Michaelsen, W. (1936) On the genus Thamnodrilus Beddard. Proceedings of the Zoological Society of London, 1936, 1171 - 1173." type="proceedings paper" year="1936">Michaelsen (1936)</bibRefCitation>
to accommodate most of the species, which were previously classified in
<taxonomicName id="4C184D42FF9AFFF41900FEF0FCE8FEAB" authority="Beddard, 1887" authorityName="Beddard" authorityYear="1887" box="[552,883,285,309]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41900FEF0FD5EFEAB" box="[552,709,285,309]" italics="true" pageId="1" pageNumber="58">Thamnodrilus</emphasis>
<bibRefCitation id="EF894B30FF9AFFF419E4FEF0FCE8FEAB" author="Beddard" box="[716,883,285,309]" pageId="1" pageNumber="58" refString="Beddard, F. E. (1887) On the structure of a new Genus of Lumbricidae (Thamnodrilus gulielmi). Proceedings of the Zoological Society of London, 1887, 154 - 163." type="proceedings paper" year="1887">Beddard, 1887</bibRefCitation>
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, with eight pairs of composite-tubular calciferous glands (Kompositenschlauchtasche) in 714 and therefore different from the
<typeStatus id="54A38863FF9AFFF418D2FEAFFBB1FEC4" box="[1018,1066,322,346]" pageId="1" pageNumber="58">type</typeStatus>
species
<emphasis id="B96CEAD3FF9AFFF41FA2FEACFA83FEC7" box="[1162,1304,321,345]" italics="true" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41FA2FEACFA8FFEC7" box="[1162,1300,321,345]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="gulielmi">Th. gulielmi</taxonomicName>
,
</emphasis>
which possesses 6 pairs of panicled-tubular glands (Rispenschlauchtasche) in 914. The genus was thoroughly revised by
<bibRefCitation id="EF894B30FF9AFFF41BBFFE64FE80FE3F" author="Zicsi" box="[151,283,393,417]" pageId="1" pageNumber="58" refString="Zicsi, A. (1998) Revision weiterer Martiodrilus - Arten (Oligochaeta: Glossoscolecidae) Regenwurmer aus Sudamerika 30. Opuscula Zoologica Budapest, 31, 149 - 164." type="journal article" year="1998">Zicsi (1998</bibRefCitation>
,
<bibRefCitation id="EF894B30FF9AFFF41A0FFE64FEF9FE3F" author="Zicsi" box="[295,354,393,417]" pageId="1" pageNumber="58" refString="Zicsi, A. (2000) Die Untergattung Martiodrilus (Martiodrilus Michaelsen, 1936) (Oligochaeta: Glossoscolecidae). Regenwurmer aus Sudamerika, 29. Opuscula Zoologica Budapest, 32, 139 - 167." type="journal article" year="2000">2000</bibRefCitation>
,
<bibRefCitation id="EF894B30FF9AFFF41A47FE64FE33FE3F" author="Zicsi" box="[367,424,393,417]" pageId="1" pageNumber="58" refString="Zicsi, A. (2001) Revision der Untergattung Martiodrilus (Maipure Righi, 1995) (Oligochaeta: Glossoscolecidae). Regenwurm- er aus Sudamerika 33. Opuscula Zoologica Budapest, 33, 113 - 131." type="journal article" year="2001">2001</bibRefCitation>
) and
<bibRefCitation id="EF894B30FF9AFFF41AC1FE64FD73FE3F" author="Zicsi" box="[489,744,393,417]" pageId="1" pageNumber="58" refString="Zicsi, A. &amp; Csuzdi, Cs. (1997) Uber wietere Riesenregenwurmer aus Ekuador. Regenwurmer aus Sudamerika, 28. (Oligochaeta). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 84, 81 - 103." type="journal article" year="1997">Zicsi &amp; Csuzdi (1997)</bibRefCitation>
in a series of papers resulting in the recognition of four subgenera:
</paragraph>
</subSubSection>
<subSubSection id="C302654AFF9AFFF41BBFFE1BFBC3FB9B" pageId="1" pageNumber="58" type="reference_group">
<paragraph id="8BA736C1FF9AFFF41BBFFE1BFDD3FDAF" blockId="1.[151,1436,501,777]" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41BEFFE18FD4EFD93" authority="Michaelsen, 1936" authorityName="Michaelsen" authorityYear="1936" box="[199,725,501,525]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="subGenus" subGenus="Martiodrilus">
<emphasis id="B96CEAD3FF9AFFF41BEFFE18FD9AFD93" box="[199,513,501,525]" italics="true" pageId="1" pageNumber="58">Martiodrilus (Martiodrilus)</emphasis>
<bibRefCitation id="EF894B30FF9AFFF41924FE18FD4EFD93" author="Michaelsen" box="[524,725,501,525]" pageId="1" pageNumber="58" refString="Michaelsen, W. (1936) On the genus Thamnodrilus Beddard. Proceedings of the Zoological Society of London, 1936, 1171 - 1173." type="proceedings paper" year="1936">Michaelsen, 1936</bibRefCitation>
</taxonomicName>
with strongly thickened septa after the gizzard and nephridia with simple nephtostome (26 spp.)
</paragraph>
<paragraph id="8BA736C1FF9AFFF41BBFFDD3FD56FDE7" blockId="1.[151,1436,501,777]" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41BEFFDD0FCDAFDCB" authority="Zicsi &amp; Csuzdi, 1997" authorityName="Zicsi &amp; Csuzdi" authorityYear="1997" box="[199,833,573,597]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="subGenus" subGenus="Cordilleroscoloex">
<emphasis id="B96CEAD3FF9AFFF41BEFFDD0FDA0FDCB" box="[199,571,573,597]" italics="true" pageId="1" pageNumber="58">Martiodrilus (Cordilleroscoloex)</emphasis>
<bibRefCitation id="EF894B30FF9AFFF4196EFDD0FCDAFDCB" author="Zicsi" box="[582,833,573,597]" pageId="1" pageNumber="58" refString="Zicsi, A. &amp; Csuzdi, Cs. (1997) Uber wietere Riesenregenwurmer aus Ekuador. Regenwurmer aus Sudamerika, 28. (Oligochaeta). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 84, 81 - 103." type="journal article" year="1997">Zicsi &amp; Csuzdi, 1997</bibRefCitation>
</taxonomicName>
with strongly thickened septa after the gizzard and nephridia with multiple nephrostome (20 spp.)
</paragraph>
<paragraph id="8BA736C1FF9AFFF41BBFFD6BFDB8FD5F" blockId="1.[151,1436,501,777]" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41BEFFD68FDD1FD03" authority="Zicsi, 1998" authorityName="Zicsi" authorityYear="1998" box="[199,586,645,669]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="subGenus" subGenus="Botaria">
<emphasis id="B96CEAD3FF9AFFF41BEFFD68FE5EFD03" box="[199,453,645,669]" italics="true" pageId="1" pageNumber="58">Martiodrilus (Botaria)</emphasis>
<bibRefCitation id="EF894B30FF9AFFF41AE4FD68FDD1FD03" author="Zicsi" box="[460,586,645,669]" pageId="1" pageNumber="58" refString="Zicsi, A. (1998) Revision weiterer Martiodrilus - Arten (Oligochaeta: Glossoscolecidae) Regenwurmer aus Sudamerika 30. Opuscula Zoologica Budapest, 31, 149 - 164." type="journal article" year="1998">Zicsi, 1998</bibRefCitation>
</taxonomicName>
with simple nephrostome, membranous septa after gizzard and two to three pairs of spermathecae (20 spp.)
</paragraph>
<paragraph id="8BA736C1FF9AFFF41BBFFD23FE7DFC97" blockId="1.[151,1436,501,777]" pageId="1" pageNumber="58">
<taxonomicName id="4C184D42FF9AFFF41BEFFD20FDF1FD7B" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[199,618,717,741]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="subGenus" subGenus="Maipure">
<emphasis id="B96CEAD3FF9AFFF41BEFFD20FE55FD7B" box="[199,462,717,741]" italics="true" pageId="1" pageNumber="58">Martiodrilus (Maipure)</emphasis>
(
<bibRefCitation id="EF894B30FF9AFFF41AF4FD20FDF9FD7B" author="Righi" box="[476,610,717,741]" pageId="1" pageNumber="58" refString="Righi, G. (1995) Colombian earthworms. Studies on Tropical Andean Ecosystems, 4, 485 - 607." type="journal article" year="1995">Righi, 1995</bibRefCitation>
)
</taxonomicName>
with simple nephrostome, membranous septa after gizzard and four pairs of spermathecae (14 spp.)
</paragraph>
<paragraph id="8BA736C1FF9AFFF41BEDFCD4FBC3FB9B" blockId="1.[151,1436,825,1029]" pageId="1" pageNumber="58">
The species
<taxonomicName id="4C184D42FF9AFFF41A7BFCD7FD9CFCCF" box="[339,519,825,849]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="duodenarius">
<emphasis id="B96CEAD3FF9AFFF41A7BFCD7FD9CFCCF" box="[339,519,825,849]" italics="true" pageId="1" pageNumber="58">M. duodenarius</emphasis>
</taxonomicName>
with membranous septa and six pairs of spermathecae was overlooked by
<bibRefCitation id="EF894B30FF9AFFF41E4CFCD4FF7AFCEB" author="Zicsi" pageId="1" pageNumber="58" refString="Zicsi, A. (2001) Revision der Untergattung Martiodrilus (Maipure Righi, 1995) (Oligochaeta: Glossoscolecidae). Regenwurm- er aus Sudamerika 33. Opuscula Zoologica Budapest, 33, 113 - 131." type="journal article" year="2001">Zicsi (2001)</bibRefCitation>
during his revision of the subgenus
<emphasis id="B96CEAD3FF9AFFF419AAFCB3FD72FCEB" box="[642,745,862,885]" italics="true" pageId="1" pageNumber="58">Maipure.</emphasis>
This was probably a reason why Fragoso and Brown (2007) kept it in
<taxonomicName id="4C184D42FF9AFFF41A2BFC6CFE3BFC07" box="[259,416,897,921]" class="Clitellata" family="Glossoscolecidae" genus="Thamnodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41A2BFC6CFE3BFC07" box="[259,416,897,921]" italics="true" pageId="1" pageNumber="58">Thamnodrilus</emphasis>
</taxonomicName>
in spite of the presence of eight pairs of composite-tubular calciferous glands. In the present material we found another species,
<taxonomicName id="4C184D42FF9AFFF4197EFC4BFD7AFC23" box="[598,737,933,957]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="gaucheri" status="sp. nov.">
<emphasis id="B96CEAD3FF9AFFF4197EFC4BFD7AFC23" box="[598,737,933,957]" italics="true" pageId="1" pageNumber="58">M. gaucheri</emphasis>
</taxonomicName>
<emphasis id="B96CEAD3FF9AFFF419C2FC48FCDEFC23" bold="true" box="[746,837,933,957]" pageId="1" pageNumber="58">
<taxonomicNameLabel id="A25F57A8FF9AFFF419C2FC48FCDEFC23" box="[746,837,933,957]" pageId="1" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with five pairs of spermathecae that did not fit into any of the
<taxonomicName id="4C184D42FF9AFFF41A25FC24FE06FC7F" box="[269,413,969,993]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41A25FC24FE06FC7F" box="[269,413,969,993]" italics="true" pageId="1" pageNumber="58">Martiodrilus</emphasis>
</taxonomicName>
subgenera. Until the proper taxonomic positions of these species are determined, we modified the diagnosis of
<taxonomicName id="4C184D42FF9AFFF41AAAFC00FD11FB9B" authority="Maipure" authorityName="Maipure" box="[386,650,1005,1029]" class="Clitellata" family="Glossoscolecidae" genus="Martiodrilus" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="58" phylum="Annelida" rank="genus">
<emphasis id="B96CEAD3FF9AFFF41AAAFC00FD11FB9B" box="[386,650,1005,1029]" italics="true" pageId="1" pageNumber="58">Martiodrilus (Maipure)</emphasis>
</taxonomicName>
subgenus to accommodate these species.
</paragraph>
</subSubSection>
</treatment>
</document>