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<document id="38E62EC26BF4F8204DE393C5C8BFC204" ID-DOI="10.5852/ejt.2019.558" ID-GBIF-Dataset="5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59" ID-ISSN="2118-9773" ID-Zenodo-Dep="3474974" ID-ZooBank="5B9DC474-1F26-4732-B7B8-32C285FE0099" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1570442388906" checkinUser="plazi" docAuthor="Fehér, Zoltán, Deli, Tamás, Erőss, Zoltán P. &amp; Lika, Romilda" docDate="2019" docId="03B17A5AF718FC51FE7CF3FB6F12FB89" docLanguage="en" docName="ejt-558_feher_deli_eross.pdf.imf" docOrigin="European Journal of Taxonomy 558" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Virpazaria (Virpazaria) gittenbergeri Fehér &amp; Deli &amp; Erőss &amp; Lika 2019, sp. nov." docType="treatment" docUuid="C987B11A-418B-48B2-A5D5-B71966765600" docUuidSource="ZooBank" docVersion="17" lastPageNumber="13" masterDocId="FF880222F712FC5DFFCFF2516A65FFED" masterDocTitle="Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="11" updateTime="1716581079825" updateUser="carolina">
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<mods:title id="677142BB18F1146A58017549CD2E6A5B">Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)</mods:title>
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<mods:namePart id="73BEB363C089A36FEA6C537ECDE2736F">Fehér, Zoltán</mods:namePart>
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<mods:affiliation id="D0A3A4726FB91541DF062220AB35A5A3">Department of Zoology, Hungarian Natural History Museum, Baross u. 13, Budapest, H- 1088, Hungary.</mods:affiliation>
<mods:nameIdentifier id="5746FBF02DF5BD5BE112A79AE3D801CB" type="email">feher.zoltan@nhmus.hu</mods:nameIdentifier>
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<mods:namePart id="F7789298951B708F091AE9D2F95CA00F">Deli, Tamás</mods:namePart>
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<mods:affiliation id="1DEF9A2F87AB33A91DFCB94A7138D2E7">Móricz Zsigmond u. 2, Gyomaendrőd, H- 5500, Hungary.</mods:affiliation>
<mods:nameIdentifier id="6541AEAF5E1AE34308AE3CFADB5023B6" type="email">virpazaria@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F4D600A879B9ACBD7D28ED4C6AD6C447">Erőss, Zoltán P.</mods:namePart>
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<mods:affiliation id="C95C8B57DB650510DEC3BBF54B6384FC">Bem u. 36., Budapest, H- 1151, Hungary.</mods:affiliation>
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<mods:namePart id="E0EE9C362BCCF75222924658F3342A7B">Lika, Romilda</mods:namePart>
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<mods:affiliation id="7660C8454FAE63771DC29A50F759363D">Departament of Biology, Faculty of Natural Sciences, Tirana University, Bulevardi Zogu i Parë, Tiranë, AL- 1001, Albania.</mods:affiliation>
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<treatment id="03B17A5AF718FC51FE7CF3FB6F12FB89" ID-DOI="http://doi.org/10.5281/zenodo.3475253" ID-GBIF-Taxon="159654645" ID-Zenodo-Dep="3475253" LSID="urn:lsid:zoobank.org:act:C987B11A-418B-48B2-A5D5-B71966765600" httpUri="http://treatment.plazi.org/id/03B17A5AF718FC51FE7CF3FB6F12FB89" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<heading id="D0EF7C20F718FC57FE7CF3FB6EE4FE29" box="[435,1153,425,452]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="4C18B0CFF718FC57FE7CF3FB6E7AFE29" ID-CoL="7TM29" authority="Feher &amp; Eross" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[435,1055,425,452]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="gittenbergeri" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF718FC57FE7CF3FB690BFE29" bold="true" box="[435,878,425,452]" italics="true" pageId="10" pageNumber="11">Virpazaria (Virpazaria) gittenbergeri</emphasis>
Fehér &amp; Erőss
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<taxonomicNameLabel id="A25FAA25F718FC57FBE9F3FB6EE4FE29" box="[1062,1153,426,452]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<figureCitation id="1323D7C9F718FC57FD12F3A16945FDE7" box="[733,800,496,522]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[211,1376,265,1904]" captionTargetId="figure@11.[211,1376,265,1904]" captionTargetPageId="11" captionText="Fig. 4. Shells of species of Virpazaria Gittenberger, 1969. AE. Virpazaria (Virpazaria) gittenbergeri Fehér &amp; Erőss sp. nov., holotype (HNHM 103411). FJ. Virpazaria (Virpazaria) pesici Fehér &amp; Deli sp. nov., holotype (HNHM 103403). Scale bars = 1 mm (note that the latter species is less magnified)." figureDoi="http://doi.org/10.5281/zenodo.3474982" httpUri="https://zenodo.org/record/3474982/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
AE
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<taxonomicName id="4C18B0CFF718FC57FF72F0636B06FDA1" ID-CoL="7PH5P" authorityName="Brusina" authorityYear="1886" box="[189,355,562,588]" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF718FC57FF72F0636B06FDA1" box="[189,355,562,588]" italics="true" pageId="10" pageNumber="11">Spelaeodiscus</emphasis>
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sp. (juv.)
<treatmentCitation id="0AB9ED5DF718FC57FE20F06368D5FDA1" author="Subai P." box="[495,688,562,588]" page="111" pageId="10" pageNumber="11" year="2009">
<bibRefCitation id="EF89B6BDF718FC57FE20F06368D5FDA1" author="Subai P." box="[495,688,562,588]" firstAuthor="Subai" journalOrPublisher="Archiv fur Molluskenkunde" pageId="10" pageNumber="11" pagination="103 - 112" part="138" refId="ref13557" refString="Subai P. 2009. New land snails from Montenegro and Albania. Archiv fur Molluskenkunde 138 (1): 103 - 112. https: // doi. org / 10.1127 / arch. moll / 0003 - 9284 / 138 / 103 - 112" title="New land snails from Montenegro and Albania" type="journal article" year="2009">Subai 2009: 111</bibRefCitation>
</treatmentCitation>
.
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<emphasis id="B96C175EF718FC57FF72F0226BACFD60" bold="true" box="[189,457,627,653]" pageId="10" pageNumber="11">Differential diagnosis</emphasis>
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This species can be distinguished from congeners by its small size and by its having one knob on the palatal peristome, and from species of the related
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<emphasis id="B96C175EF718FC57FCD0F09469A0FD32" box="[799,965,709,735]" italics="true" pageId="10" pageNumber="11">Spelaeodiscus</emphasis>
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, which have one knob/tooth on the peristome by its inferiorly broader rounded body whorl and narrower aperture.
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<emphasis id="B96C175EF718FC57FF72F17B6B21FCA9" bold="true" box="[189,324,810,836]" pageId="10" pageNumber="11">Etymology</emphasis>
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This species is dedicated to and named after Edmund Gittenberger (Leiden,
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), a prominent malacologist who did pioneering work on the subterranean terrestrial gastropods of the Balkans. The name is used as the genitive of a noun of male gender.
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<emphasis id="B96C175EF718FC57FF72F1B16BCFFC17" bold="true" box="[189,426,992,1018]" pageId="10" pageNumber="11">Material examined</emphasis>
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<typeStatus id="54A375EEF718FC57FF25F65E6B39FBC4" box="[234,348,1039,1065]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
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<collectingCountry id="F30F8BDCF718FC57FF72F6636BEFFBA1" box="[189,394,1074,1100]" name="Serbia" pageId="10" pageNumber="11">MONTENEGRO</collectingCountry>
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,
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S of Štegvaš Pass on OstrosUlcinj
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road (
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S of Arbnež
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);
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,
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alt.;
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;
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,
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and
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leg.;
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.
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</emphasis>
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF72F6EB6E8DFA27" blockId="10.[189,1399,1174,1482]" pageId="10" pageNumber="11">
<materialsCitation id="3B70C111F718FC57FF72F6EB6EC0FB39" ID-GBIF-Occurrence="2424146979" box="[189,1189,1209,1236]" collectingDate="2015-05-28" collectionCode="HNHM" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="MONTENEGRO" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCode="HNHM 103413" specimenCount="20" typeStatus="paratype">
<collectingCountry id="F30F8BDCF718FC57FF72F6EB6BEFFB39" box="[189,394,1210,1236]" name="Serbia" pageId="10" pageNumber="11">MONTENEGRO</collectingCountry>
<specimenCount id="9D1E00C5F718FC57FE65F6EB6875FB39" box="[426,528,1210,1236]" count="20" pageId="10" pageNumber="11" type="generic">20 spec.</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBBE6337F718FC57FC26F6EB6EC0FB39" box="[1001,1189,1210,1236]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM 103413</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FB0BF6EB688AFB1B" ID-GBIF-Occurrence="2424151010" collectingDate="2015-05-28" collectionCode="NHMW" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="Rumija Mts" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCode="111671" specimenCount="20" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FB0BF6EB6F4EFB39" box="[1220,1323,1210,1236]" count="20" pageId="10" pageNumber="11" type="generic">20 spec.</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED095389F718FC57FDFEF68C68F4FB1A" box="[561,657,1245,1271]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="11" type="Museum">NHMW</collectionCode>
<specimenCode id="DBBE6337F718FC57FD56F68D688AFB1B" box="[665,751,1244,1270]" pageId="10" pageNumber="11">111671</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FCC7F68C6A81FAF7" ID-GBIF-Occurrence="2424146990" collectingDate="2015-05-28" collectionCode="DT" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="Rumija Mts" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCount="20" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FCC7F68C690EFB1A" box="[776,875,1245,1271]" count="20" pageId="10" pageNumber="11" type="generic">20 spec.</specimenCount>
, 38 fr.; same collection data as for holotype;
<collectionCode id="ED095389F718FC57FF72F7516A81FAF7" box="[189,228,1280,1306]" pageId="10" pageNumber="11">DT</collectionCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FF35F7516F13FAF6" ID-GBIF-Occurrence="2424146980" box="[250,1398,1280,1307]" collectingDate="2017-07-16" collectionCode="HNHM" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="Rumija Mts" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCode="HNHM 103451" specimenCount="16" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FF35F7516B51FAF7" box="[250,308,1280,1306]" count="8" pageId="10" pageNumber="11" type="generic">8 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FEF0F7516BE6FAF7" box="[319,387,1280,1306]" count="8" pageId="10" pageNumber="11" type="generic">8 juv.</specimenCount>
; same locality as for holotype;
<date id="FFA6ED8CF718FC57FD25F751691EFAF7" box="[746,891,1280,1306]" pageId="10" pageNumber="11" value="2017-07-16">
<collectingDate id="EFE21464F718FC57FD25F751691EFAF7" box="[746,891,1280,1306]" pageId="10" pageNumber="11" value="2017-07-16">16 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26EDAE9AF718FC57FC49F7516983FAF7" box="[902,998,1280,1306]" pageId="10" pageNumber="11">Z. Erőss</collectorName>
and
<collectorName id="26EDAE9AF718FC57FBD2F7516E1AFAF7" box="[1053,1151,1280,1306]" pageId="10" pageNumber="11">Z. Fehér</collectorName>
leg.;
<specimenCode id="DBBE6337F718FC57FB70F7516F13FAF6" box="[1215,1398,1280,1307]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM 103451</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FF00F7726F12FAD0" ID-GBIF-Occurrence="2424151035" box="[207,1399,1315,1342]" collectingDate="2017-07-16" collectingDateMax="74-11-16" collectingDateMin="2017-07-16" collectionCode="NHMW" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="Rumija Mts" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCode="111674" specimenCount="17" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FF00F7726B6EFAD0" box="[207,267,1315,1341]" count="8" pageId="10" pageNumber="11" type="generic">8 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FED6F7726B3BFAD0" box="[281,350,1315,1341]" count="9" pageId="10" pageNumber="11" type="generic">9 juv.</specimenCount>
; same locality as for holotype;
<date id="FFA6ED8CF718FC57FD1EF7726902FAD0" box="[721,871,1315,1341]" pageId="10" pageNumber="11" value="2017-07-16">
<collectingDate id="EFE21464F718FC57FD1EF7726902FAD0" box="[721,871,1315,1341]" pageId="10" pageNumber="11" value="2017-07-16">16 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26EDAE9AF718FC57FCBBF77569B3FAD0" box="[884,982,1315,1342]" pageId="10" pageNumber="11">Z. Erőss</collectorName>
and
<collectorName id="26EDAE9AF718FC57FBDEF7756E10FAD0" box="[1041,1141,1315,1342]" pageId="10" pageNumber="11">Z. Fehér</collectorName>
leg.;
<collectionCode id="ED095389F718FC57FB76F7756F7CFAD3" box="[1209,1305,1316,1342]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="11" type="Museum">NHMW</collectionCode>
<specimenCode id="DBBE6337F718FC57FAEEF7726F12FAD0" box="[1313,1399,1315,1341]" pageId="10" pageNumber="11">111674</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FF1EF7176F71FA8C" ID-GBIF-Occurrence="2424151039" box="[209,1300,1350,1377]" collectingDate="2017-07-16" collectionCode="ER" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="350" latitude="42.0582" location="Rumija Mts" longLatPrecision="6" longitude="19.3535" pageId="10" pageNumber="11" specimenCount="25" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FF1EF7176B7BFA8D" box="[209,286,1350,1376]" count="16" pageId="10" pageNumber="11" type="generic">16 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FEE1F7176B10FA8D" box="[302,373,1350,1376]" count="9" pageId="10" pageNumber="11" type="generic">9 juv.</specimenCount>
; same locality as for holotype;
<date id="FFA6ED8CF718FC57FD3AF71769EBFA8D" box="[757,910,1350,1376]" pageId="10" pageNumber="11" value="2017-07-16">
<collectingDate id="EFE21464F718FC57FD3AF71769EBFA8D" box="[757,910,1350,1376]" pageId="10" pageNumber="11" value="2017-07-16">16 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26EDAE9AF718FC57FC52F7166E67FA8D" box="[925,1026,1350,1377]" pageId="10" pageNumber="11">Z. Erőss</collectorName>
and
<collectorName id="26EDAE9AF718FC57FB8EF7166EC2FA8D" box="[1089,1191,1350,1377]" pageId="10" pageNumber="11">Z. Fehér</collectorName>
leg.;
<collectionCode id="ED095389F718FC57FB20F7166F71FA8C" box="[1263,1300,1351,1377]" pageId="10" pageNumber="11">ER</collectionCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FAFCF71769BDFA4A" ID-GBIF-Occurrence="2424151057" collectingDate="2017-07-16" collectionCode="HNHM" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="480" latitude="42.0636" location="SW of Stegvas Pass" longLatPrecision="6" longitude="19.371" pageId="10" pageNumber="11" specimenCode="HNHM 103445" specimenCount="3" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FAFCF7176F14FA8D" box="[1331,1393,1350,1376]" count="2" pageId="10" pageNumber="11" type="generic">2 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FF72F73B6B6DFA6E" box="[189,264,1385,1412]" count="1" pageId="10" pageNumber="11" type="generic">1 juv.</specimenCount>
;
<location id="8EC79D97F718FC57FED6F73B6872FA69" LSID="urn:lsid:plazi:treatment:03B17A5AF718FC51FE7CF3FB6F12FB89:8EC79D97F718FC57FED6F73B6872FA69" box="[281,535,1384,1412]" country="Serbia" latitude="42.0636" longLatPrecision="6" longitude="19.371" name="SW of Stegvas Pass" pageId="10" pageNumber="11">SW of Štegvaš Pass</location>
,
<locationDeviation id="9A15D26EF718FC57FDE6F73B6989FA6E" box="[553,1004,1385,1412]" location="Arbnez on Ostros-Ulcinj" pageId="10" pageNumber="11">
<quantity id="4CE066A9F718FC57FDE6F73B680CFA6E" box="[553,617,1385,1412]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="2.0">2 km</quantity>
S of Arbnež on OstrosUlcinj
</locationDeviation>
road;
<geoCoordinate id="EE2CAD8BF718FC57FB8DF7386EB7FA69" box="[1090,1234,1385,1412]" degrees="42.0636" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="42.0636">42.0636° N</geoCoordinate>
,
<geoCoordinate id="EE2CAD8BF718FC57FB2BF73B6F14FA69" box="[1252,1393,1386,1412]" degrees="19.3710" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="19.371">19.3710° E</geoCoordinate>
;
<quantity id="4CE066A9F718FC57FF72F7DC6B62FA4A" box="[189,263,1421,1447]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="11" unit="m" value="480.0">
<elevation id="00352C7FF718FC57FF72F7DC6B62FA4A" box="[189,263,1421,1447]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="11" unit="m" value="480.0">480 m</elevation>
</quantity>
alt.;
<date id="FFA6ED8CF718FC57FE8DF7DD6BB0FA4A" box="[322,469,1420,1447]" pageId="10" pageNumber="11" value="2017-07-16">
<collectingDate id="EFE21464F718FC57FE8DF7DD6BB0FA4A" box="[322,469,1420,1447]" pageId="10" pageNumber="11" value="2017-07-16">16 Jul. 2017</collectingDate>
</date>
;
<collectorName id="26EDAE9AF718FC57FE2EF7DC6827FA4A" box="[481,578,1421,1447]" pageId="10" pageNumber="11">Z. Erőss</collectorName>
and
<collectorName id="26EDAE9AF718FC57FDB4F7DC68BBFA4A" box="[635,734,1421,1447]" pageId="10" pageNumber="11">Z. Fehér</collectorName>
leg.;
<specimenCode id="DBBE6337F718FC57FCD0F7DC69BDFA4A" box="[799,984,1420,1447]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM 103445</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FC3FF7DC6823FA27" ID-GBIF-Occurrence="2424151026" collectingDate="2017-07-16" collectionCode="NHMW" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="480" latitude="42.0636" location="Stegvas Pass" longLatPrecision="6" longitude="19.371" pageId="10" pageNumber="11" specimenCode="111673" specimenCount="4" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FC3FF7DC6E4EFA4A" box="[1008,1067,1421,1447]" count="3" pageId="10" pageNumber="11" type="generic">3 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FBF7F7DC6E19FA4A" box="[1080,1148,1421,1447]" count="1" pageId="10" pageNumber="11" type="generic">1 juv.</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED095389F718FC57FE46F7E16B8CFA27" box="[393,489,1456,1482]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="11" type="Museum">NHMW</collectionCode>
<specimenCode id="DBBE6337F718FC57FE3FF7E16823FA27" box="[496,582,1456,1482]" pageId="10" pageNumber="11">111673</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FD90F7E16E86FA27" ID-GBIF-Occurrence="2424146968" box="[607,1251,1456,1482]" collectingDate="2017-07-16" collectionCode="ER" collectorName="Z. Eross &amp; Z. Feher" country="Serbia" elevation="480" latitude="42.0636" location="Stegvas Pass" longLatPrecision="6" longitude="19.371" pageId="10" pageNumber="11" specimenCount="4" typeStatus="paratype">
<specimenCount id="9D1E00C5F718FC57FD90F7E168FFFA27" box="[607,666,1456,1482]" count="3" pageId="10" pageNumber="11" type="generic">3 ad.</specimenCount>
,
<specimenCount id="9D1E00C5F718FC57FD68F7E16889FA27" box="[679,748,1456,1482]" count="1" pageId="10" pageNumber="11" type="generic">1 juv.</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED095389F718FC57FB75F7E16E86FA27" box="[1210,1251,1456,1482]" pageId="10" pageNumber="11">ER</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF25F7A06BC3F9E6" blockId="10.[189,1398,1521,1653]" box="[234,422,1521,1547]" pageId="10" pageNumber="11">
<emphasis id="B96C175EF718FC57FF25F7A06BC3F9E6" bold="true" box="[234,422,1521,1547]" pageId="10" pageNumber="11">Other material</emphasis>
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF72F444699BF998" blockId="10.[189,1398,1521,1653]" pageId="10" pageNumber="11">
<materialsCitation id="3B70C111F718FC57FF72F44469C8F9BF" ID-GBIF-Occurrence="2424146965" collectingDate="2015-05-27" collectionCode="HNHM" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Albania" elevation="190" latitude="42.0506" location="Vallas" longLatPrecision="6" longitude="19.4162" municipality="Mt Tarabosh" pageId="10" pageNumber="11" specimenCode="HNHM 103415" specimenCount="1" stateProvince="Shkoder district">
<collectingCountry id="F30F8BDCF718FC57FF72F4446B5AF9C2" box="[189,319,1557,1583]" name="Albania" pageId="10" pageNumber="11">ALBANIA</collectingCountry>
<collectingRegion id="49DC05AEF718FC57FE95F445687CF9C3" box="[346,537,1556,1582]" country="Albania" name="Shkoder" pageId="10" pageNumber="11">Shkodër district</collectingRegion>
,
<collectingMunicipality id="6BC35136F718FC57FDE9F44468DBF9C3" box="[550,702,1556,1583]" pageId="10" pageNumber="11">Mt Tarabosh</collectingMunicipality>
,
<location id="8EC79D97F718FC57FD05F4456970F9C3" LSID="urn:lsid:plazi:treatment:03B17A5AF718FC51FE7CF3FB6F12FB89:8EC79D97F718FC57FD05F4456970F9C3" box="[714,789,1556,1582]" country="Albania" latitude="42.0506" longLatPrecision="6" longitude="19.4162" municipality="Mt Tarabosh" name="Vallas" pageId="10" pageNumber="11" stateProvince="Shkoder district">Vallas</location>
,
<location id="8EC79D97F718FC57FCEEF44469DEF9C2" LSID="urn:lsid:plazi:treatment:03B17A5AF718FC51FE7CF3FB6F12FB89:8EC79D97F718FC57FCEEF44469DEF9C2" box="[801,955,1557,1583]" country="Albania" latitude="42.0506" longLatPrecision="6" longitude="19.4162" municipality="Mt Tarabosh" name="near mosque" pageId="10" pageNumber="11" stateProvince="Shkoder district">near mosque</location>
;
<geoCoordinate id="EE2CAD8BF718FC57FC07F4456E36F9C2" box="[968,1107,1556,1583]" degrees="42.0506" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="42.0506">42.0506° N</geoCoordinate>
,
<geoCoordinate id="EE2CAD8BF718FC57FBAFF4456E8CF9C2" box="[1120,1257,1556,1583]" degrees="19.4162" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="19.4162">19.4162° E</geoCoordinate>
;
<quantity id="4CE066A9F718FC57FB39F4446F24F9C2" box="[1270,1345,1557,1583]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="11" unit="m" value="190.0">
<elevation id="00352C7FF718FC57FB39F4446F24F9C2" box="[1270,1345,1557,1583]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="11" unit="m" value="190.0">190 m</elevation>
</quantity>
alt.;
<date id="FFA6ED8CF718FC57FF72F4696B3CF9BF" box="[189,345,1591,1618]" pageId="10" pageNumber="11" value="2015-05-27">
<collectingDate id="EFE21464F718FC57FF72F4696B3CF9BF" box="[189,345,1591,1618]" pageId="10" pageNumber="11" value="2015-05-27">27 May 2015</collectingDate>
</date>
;
<collectorName id="26EDAE9AF718FC57FEACF4696BD0F9BF" box="[355,437,1592,1618]" pageId="10" pageNumber="11">T. Deli</collectorName>
,
<collectorName id="26EDAE9AF718FC57FE0FF4696845F9BF" box="[448,544,1592,1618]" pageId="10" pageNumber="11">Z. Erőss</collectorName>
and
<collectorName id="26EDAE9AF718FC57FD9AF46968D2F9BF" box="[597,695,1592,1618]" pageId="10" pageNumber="11">Z. Fehér</collectorName>
leg.;
<specimenCode id="DBBE6337F718FC57FD39F46969C8F9BF" box="[758,941,1591,1618]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM 103415</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FC0CF4696B90F998" ID-GBIF-Occurrence="2424151041" collectingDate="2015-05-27" collectionCode="NHMW" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Albania" elevation="190" latitude="42.0506" location="Vallas" longLatPrecision="6" longitude="19.4162" municipality="Mt Tarabosh" pageId="10" pageNumber="11" specimenCode="110430/MN/0986" specimenCount="1" stateProvince="Shkoder">
same collection data as for preceding;
<collectionCode id="ED095389F718FC57FF72F40A6B78F998" box="[189,285,1627,1653]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="11" type="Museum">NHMW</collectionCode>
<specimenCode id="DBBE6337F718FC57FEEBF40A6B90F998" box="[292,501,1627,1653]" pageId="10" pageNumber="11">110430/MN/0986</specimenCode>
</materialsCitation>
<materialsCitation id="3B70C111F718FC57FDC1F40A699FF998" ID-GBIF-Occurrence="2424146996" box="[526,1018,1627,1653]" collectingDate="2015-05-27" collectionCode="DT" collectorName="T. Deli &amp; Z. Eross &amp; Z. Feher" country="Albania" elevation="190" latitude="42.0506" location="Vallas" longLatPrecision="6" longitude="19.4162" municipality="Mt Tarabosh" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Shkoder">
same collection data as for preceding;
<collectionCode id="ED095389F718FC57FC1EF40A699FF998" box="[977,1018,1627,1653]" pageId="10" pageNumber="11">DT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F718FC51FF72F4CD684AFDAC" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="8BA7CB4CF718FC57FF72F4CD6B2BF95B" blockId="10.[189,334,1692,1718]" box="[189,334,1692,1718]" pageId="10" pageNumber="11">
<heading id="D0EF7C20F718FC57FF72F4CD6B2BF95B" bold="true" box="[189,334,1692,1718]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B96C175EF718FC57FF72F4CD6B2BF95B" bold="true" box="[189,334,1692,1718]" pageId="10" pageNumber="11">Dimensions</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF72F49A6F5DF8E4" blockId="10.[189,1399,1739,1801]" pageId="10" pageNumber="11">
<typeStatus id="54A375EEF718FC57FF72F49A6B4AF908" box="[189,303,1739,1765]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
: H
<quantity id="4CE066A9F718FC57FE9CF49A6BDBF908" box="[339,446,1739,1766]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="10" pageNumber="11" unit="mm" value="1.55">1.55 mm</quantity>
; W
<quantity id="4CE066A9F718FC57FE26F49A6820F908" box="[489,581,1739,1766]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
; Ha 1.0 mm; Wa
<quantity id="4CE066A9F718FC57FCC5F49A6903F908" box="[778,870,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="11" unit="mm" value="1.1">1.1 mm</quantity>
.
<typeStatus id="54A375EEF718FC57FCA0F49A6984F908" box="[879,993,1739,1765]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
(HNHM 103413, n =5): H
<quantity id="4CE066A9F718FC57FAD1F49A6A8BF8E4" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="10" pageNumber="11" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
; W
<quantity id="4CE066A9F718FC57FED3F4BF6BDEF8E4" box="[284,443,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="2.425" metricValueMax="2.5" metricValueMin="2.35" pageId="10" pageNumber="11" unit="mm" value="2.425" valueMax="2.5" valueMin="2.35">2.352.5 mm</quantity>
. Vallas population (HNHM 103415, n= 5): H
<quantity id="4CE066A9F718FC57FC19F4BF6E10F8E4" box="[982,1141,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.5" metricValueMin="1.25" pageId="10" pageNumber="11" unit="mm" value="1.375" valueMax="1.5" valueMin="1.25">1.251.5 mm</quantity>
; W
<quantity id="4CE066A9F718FC57FB6CF4BF6F56F8E4" box="[1187,1331,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="10" pageNumber="11" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
.
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF72F57E6B2BF8A4" blockId="10.[189,334,1839,1865]" box="[189,334,1839,1865]" pageId="10" pageNumber="11">
<heading id="D0EF7C20F718FC57FF72F57E6B2BF8A4" bold="true" box="[189,334,1839,1865]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B96C175EF718FC57FF72F57E6B2BF8A4" bold="true" box="[189,334,1839,1865]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF718FC57FF72F50F6EB4F853" blockId="10.[189,1399,1884,1983]" pageId="10" pageNumber="11">Shell discoid, spire barely raised, consists of 4¼4¾ regularly increasing whorls, of which 1½2 are protoconch whorls. Fresh shells glassy translucent, older shells whitish. Protoconch grained. Teleoconch whorls equidistantly ribbed: 5969 ribs on body whorl and fine, regular lines in between.</paragraph>
<caption id="DF679BC4F719FC56FF72F5DB6F0DF801" ID-DOI="http://doi.org/10.5281/zenodo.3474982" ID-Zenodo-Dep="3474982" httpUri="https://zenodo.org/record/3474982/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1930,1956]" targetBox="[211,1376,265,1904]" targetPageId="11">
<paragraph id="8BA7CB4CF719FC56FF72F5DB6F0DF801" blockId="11.[189,1399,1930,2028]" pageId="11" pageNumber="12">
<emphasis id="B96C175EF719FC56FF72F5DB6B69F849" bold="true" box="[189,268,1930,1956]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
Shells of species of
<taxonomicName id="4C18B0CFF719FC56FDCAF5DA690EF849" authority="Gittenberger, 1969" authorityName="Gittenberger" authorityYear="1969" box="[517,875,1930,1956]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF719FC56FDCAF5DA68E7F849" box="[517,642,1931,1956]" italics="true" pageId="11" pageNumber="12">Virpazaria</emphasis>
<bibRefCitation id="EF89B6BDF719FC56FD44F5DB690EF849" author="Gittenberger E." box="[651,875,1930,1956]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="11" pageNumber="12" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
.
<emphasis id="B96C175EF719FC56FCB7F5DB69CAF849" bold="true" box="[888,943,1930,1956]" pageId="11" pageNumber="12">AE</emphasis>
.
<taxonomicName id="4C18B0CFF719FC56FC70F5DA6B0FF825" authority="Feher &amp; Eross" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="gittenbergeri" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF719FC56FC70F5DA6F13F849" box="[959,1398,1930,1957]" italics="true" pageId="11" pageNumber="12">Virpazaria (Virpazaria) gittenbergeri</emphasis>
Fehér &amp; Erőss
</taxonomicName>
<taxonomicNameLabel id="A25FAA25F719FC56FEBCF5FF6BB4F825" box="[371,465,1966,1992]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (HNHM 103411).
<emphasis id="B96C175EF719FC56FCE3F5FF693EF825" bold="true" box="[812,859,1966,1992]" pageId="11" pageNumber="12">FJ</emphasis>
.
<taxonomicName id="4C18B0CFF719FC56FCA3F5FF6F12F82A" authority="Feher &amp; Deli" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[876,1399,1965,1992]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="pesici" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF719FC56FCA3F5FF6EAAF82A" box="[876,1231,1966,1992]" italics="true" pageId="11" pageNumber="12">Virpazaria (Virpazaria) pesici</emphasis>
Fehér &amp; Deli
</taxonomicName>
<taxonomicNameLabel id="A25FAA25F719FC56FF72F5806B7FF806" box="[189,282,2001,2027]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (HNHM 103403). Scale bars = 1 mm (note that the latter species is less magnified).
</paragraph>
</caption>
<paragraph id="8BA7CB4CF71EFC51FF72F35C684AFDAC" blockId="12.[189,1399,268,577]" pageId="12" pageNumber="13">In bottom view, umbilicus wide, 1/41/5 of total shell width, a small part of it overlaid by columellar part of aperture. In frontal view, silhouette of body whorl asymmetrically rounded (shouldered): broader curved in bottom, more angulated on upper side and near suture horizontally flattened. Suture barely depressed.Aperture narrow crescent-shaped. Instead of a flap, with an outstretched swelling (a thickened and emerged callus) on parietal side. One knob on palatal peristome, ca at one-third of way between its angular and columellar junctions. Peristome somewhat reflected between columellar junction and palatal knob, appearing thickened in frontal view. Above knob, peristome simple, delineating an apertural sinus. In side view, parietal swelling oblique (inferior part stands forward) and aperture sigmoidally inclined (inferior part stands backward).</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71EFC51FF72F03668F4FCF4" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BA7CB4CF71EFC51FF72F0366B33FD6C" blockId="12.[189,342,615,641]" box="[189,342,615,641]" pageId="12" pageNumber="13">
<heading id="D0EF7C20F71EFC51FF72F0366B33FD6C" bold="true" box="[189,342,615,641]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96C175EF71EFC51FF72F0366B33FD6C" bold="true" box="[189,342,615,641]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71EFC51FF72F0C768F4FCF4" blockId="12.[189,1399,662,793]" pageId="12" pageNumber="13">
This species is known from the southeastern part of the Rumija Mts (also known as Mt Tarabosh). The Štegvaš Pass (
<collectingCountry id="F30F8BDCF71EFC51FE54F0E8684BFD3E" box="[411,558,697,723]" name="Serbia" pageId="12" pageNumber="13">Montenegro</collectingCountry>
) and Vallas (
<collectingCountry id="F30F8BDCF71EFC51FD07F0E8694DFD39" box="[712,808,697,724]" name="Albania" pageId="12" pageNumber="13">Albania</collectingCountry>
) populations are within a distance of
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of one another. Both sites are on UpperMiddle Triassic limestone. At the
<typeStatus id="54A375EEF71EFC51FC1DF08C6E60FD1A" box="[978,1029,733,759]" pageId="12" pageNumber="13">type</typeStatus>
locality, this species was found together with
<taxonomicName id="4C18B0CFF71EFC51FEAFF1516B8AFCF4" box="[352,495,767,793]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="stojaspali">
<emphasis id="B96C175EF71EFC51FEAFF1516B8AFCF4" box="[352,495,767,793]" italics="true" pageId="12" pageNumber="13">V. stojaspali</emphasis>
</taxonomicName>
and
<emphasis id="B96C175EF71EFC51FDE7F15168F4FCF4" box="[552,657,768,793]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C18B0CFF71EFC51FDE7F15168E8FCF4" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[552,653,768,793]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pesici">V. pesici</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71EFC51FF72F1106E03FC1E" pageId="12" pageNumber="13" type="conservation">
<paragraph id="8BA7CB4CF71EFC51FF72F1106BD0FCB6" blockId="12.[189,437,833,859]" box="[189,437,833,859]" pageId="12" pageNumber="13">
<heading id="D0EF7C20F71EFC51FF72F1106BD0FCB6" bold="true" box="[189,437,833,859]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96C175EF71EFC51FF72F1106BD0FCB6" bold="true" box="[189,437,833,859]" pageId="12" pageNumber="13">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71EFC51FF72F1216E03FC1E" blockId="12.[189,1399,879,1012]" pageId="12" pageNumber="13">
To our present knowledge, this is a narrow-range endemic species with an area of occurrence (AOO) smaller than
<quantity id="4CE066A9F71EFC51FE96F1C26BC1FC40" box="[345,420,915,941]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="km" value="20.0">20 km</quantity>
<superScript id="7C6D6604F71EFC51FE6AF1C06BCBFC72" attach="right" box="[421,430,913,927]" fontSize="6" pageId="12" pageNumber="13">2</superScript>
. However, there is no reason to suppose that AOO, extent of occurrence (EOO), number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71EFC51FF72F64A6F12FB89" box="[189,1399,1051,1124]" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BA7CB4CF71EFC51FF72F64A6B4AFBD8" blockId="12.[189,303,1051,1077]" box="[189,303,1051,1077]" pageId="12" pageNumber="13">
<heading id="D0EF7C20F71EFC51FF72F64A6B4AFBD8" bold="true" box="[189,303,1051,1077]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96C175EF71EFC51FF72F64A6B4AFBD8" bold="true" box="[189,303,1051,1077]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71EFC51FF72F6186F12FB89" blockId="12.[189,1399,1097,1124]" box="[189,1399,1097,1124]" pageId="12" pageNumber="13">The Vallas population differs from the typical form by its denser ribbing (7688 ribs on the body whorl).</paragraph>
</subSubSection>
</treatment>
</document>