treatments-xml/data/03/AB/DE/03ABDE62CC07B5104AD69F7BFEAFFC2D.xml

<document id="113CCEF4CBB48A187367C0CBB8B71E84" ID-DOI="10.35463/j.apr.2021.01.02" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520760" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705450788631" checkinUser="felipe" docAuthor="Akkemik, Unal" docDate="2021" docId="03ABDE62CC07B5104AD69F7BFEAFFC2D" docLanguage="en" docName="ActaPalaeontolRom.17.1.15-26.pdf" docOrigin="Acta Palaeontologica Romaniae 17 (1)" docSource="http://dx.doi.org/10.35463/j.apr.2021.01.02" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Juniperoxylon acarcae Akkemik 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="25" masterDocId="FF92A61ACC04B51A4A419E58FFA1FFFD" masterDocTitle="A New Species Of Juniperoxylon From The Early Miocene Of Northwestern Turkey" masterLastPageNumber="26" masterPageNumber="15" pageNumber="18" updateTime="1708527701142" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="966978A6A213EA24B3C1DB6BF952002E">A New Species Of Juniperoxylon From The Early Miocene Of Northwestern Turkey</mods:title>
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<mods:namePart id="FAB07E76AA7F79313910BC5CB0F18EC7">Akkemik, Unal</mods:namePart>
<mods:affiliation id="48CD85AA6F5AB6EADC2803CDDF89572C">Istanbul University-Cerrahpasa, Forestry Faculty, Forest Botany Department, 34473 Bahceköy, Istanbul, Turkey</mods:affiliation>
<mods:nameIdentifier id="A342EADEFF2F49533D5E5CB41BA0E3FB" type="email">uakkemik@istanbul.edu.tr</mods:nameIdentifier>
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<mods:title id="CEADF168A909092B34190603D48B0159">Acta Palaeontologica Romaniae</mods:title>
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<mods:date id="A47A4372051F4BBB4E016ABF9C2D173A">2021</mods:date>
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<treatment id="03ABDE62CC07B5104AD69F7BFEAFFC2D" LSID="urn:lsid:plazi:treatment:03ABDE62CC07B5104AD69F7BFEAFFC2D" httpUri="http://treatment.plazi.org/id/03ABDE62CC07B5104AD69F7BFEAFFC2D" lastPageId="10" lastPageNumber="25" pageId="3" pageNumber="18">
<subSubSection id="C3183CFFCC07B5194AD69F7BFDC0FEC6" box="[151,609,291,315]" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="8BBD6F74CC07B5194AD69F7BFDC0FEC6" blockId="3.[151,710,158,347]" box="[151,609,291,315]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69F7BFDC0FEC6" bold="true" box="[151,609,291,315]" pageId="3" pageNumber="18">
<taxonomicName id="4C0214F7CC07B5194AD69F7BFDA5FEC6" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[151,516,291,315]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC07B5194AD69F7BFE35FEC6" bold="true" box="[151,404,291,315]" italics="true" pageId="3" pageNumber="18">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC07B519484B9F7BFDC0FEC6" box="[522,609,291,315]" pageId="3" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69F1BFF57FEA6" box="[151,246,323,347]" pageId="3" pageNumber="18" type="description">
<paragraph id="8BBD6F74CC07B5194AD69F1BFF57FEA6" blockId="3.[151,710,158,347]" box="[151,246,323,347]" pageId="3" pageNumber="18">
<figureCitation id="133973F1CC07B5194AD69F1BFF57FEA6" box="[151,246,323,347]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[267,302,1960,1981]" captionStartId-1="5.[234,267,1792,1813]" captionStartId-2="6.[215,248,1853,1874]" captionStartId-3="7.[151,184,1947,1968]" captionTargetBox-0="[481,1144,237,1878]" captionTargetBox-1="[251,1342,151,1750]" captionTargetBox-2="[230,1363,188,1812]" captionTargetBox-3="[156,1365,168,1905]" captionTargetId-0="figure-264@2.[397,1190,155,1952]" captionTargetId-1="figure-33@5.[246,1342,151,1750]" captionTargetId-2="figure-48@6.[225,1363,183,1812]" captionTargetId-3="figure-16@7.[152,1365,163,1905]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 2 The type specimen (AGUD08). These are remaining pieces from AGUD08 after taking three thin sec- tions and storied at the Department of Forest Botany, Faculty of Forestry, Istanbul University-Cerrahpasa." captionText-1="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." captionText-2="Fig. 4 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of trache- ids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows)." captionText-3="Fig. 5 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field." figureDoi-0="http://doi.org/10.5281/zenodo.10520764" figureDoi-1="http://doi.org/10.5281/zenodo.10520766" figureDoi-2="http://doi.org/10.5281/zenodo.10520768" figureDoi-3="http://doi.org/10.5281/zenodo.10520770" httpUri-0="https://zenodo.org/record/10520764/files/figure.png" httpUri-1="https://zenodo.org/record/10520766/files/figure.png" httpUri-2="https://zenodo.org/record/10520768/files/figure.png" httpUri-3="https://zenodo.org/record/10520770/files/figure.png" pageId="3" pageNumber="18">Figs. 2-5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69FDEFEB0FDFC" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BBD6F74CC07B5194AD69FDEFEDAFE60" blockId="3.[151,772,389,1572]" box="[151,379,389,414]" pageId="3" pageNumber="18">
<materialsCitation id="3B6A6529CC07B5194AD69FDEFEDAFE60" box="[151,379,389,414]" collectionCode="AGUD" country="Turkey" county="Asagiguney Village" location="Galatian Volcanic Province" municipality="Faculty of Forestry" pageId="3" pageNumber="18" specimenCode="AGUD08, PFN001814" specimenCount="1" stateProvince="Istanbul" typeStatus="holotype">
<emphasis id="B976B366CC07B5194AD69FDEFEA7FE63" bold="true" box="[151,262,390,414]" pageId="3" pageNumber="18">
<typeStatus id="54B9D1D6CC07B5194AD69FDEFEA0FE63" box="[151,257,390,414]" pageId="3" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBA4C70FCC07B5194B4C9FDDFED5FE60" box="[269,372,389,413]" pageId="3" pageNumber="18">AGUD08</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD6F74CC07B5194AD69FFFFEB0FDFC" blockId="3.[151,772,389,1572]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69FFFFEB2FE42" bold="true" box="[151,275,423,447]" pageId="3" pageNumber="18">Repository</emphasis>
. The Department of Forest Botany, Faculty of Forestry, Istanbul University-Cerrahpasa, Istanbul, Turkey.
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69C52FD90FD5B" pageId="3" pageNumber="18" type="etymology">
<paragraph id="8BBD6F74CC07B5194AD69C52FD90FD5B" blockId="3.[151,772,389,1572]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69C52FEB8FDDF" bold="true" box="[151,281,522,546]" pageId="3" pageNumber="18">Etymology.</emphasis>
The epithet 
<taxonomicName id="4C0214F7CC07B5194BF09C52FDB2FDDF" authorityName="Akkemik" authorityYear="2021" box="[433,531,522,546]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="acarcae">
“ 
<emphasis id="B976B366CC07B5194BFC9C53FDB2FDDF" box="[445,531,523,546]" italics="true" pageId="3" pageNumber="18">acarcae</emphasis>
</taxonomicName>
” originates from the surname of Dr. N. Neslihan Acarca Bayam who worked on the petrified woods in the Galatian Volcanic Province for her PhD in 
<collectingCountry id="F3152FE4CC07B519484B9C35FDFAFD78" box="[522,603,621,645]" name="Turkey" pageId="3" pageNumber="18">Turkey</collectingCountry>
. It was the first PhD on petrified woods in 
<collectingCountry id="F3152FE4CC07B5194B9C9CD6FD8FFD5B" box="[477,558,654,678]" name="Turkey" pageId="3" pageNumber="18">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69CE8FD4FFCD7" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BBD6F74CC07B5194AD69CE8FD6AFD3A" blockId="3.[151,772,389,1572]" box="[151,715,687,712]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69CE8FD9DFD35" bold="true" box="[151,572,688,712]" pageId="3" pageNumber="18">Plant Fossil Names Registry Number</emphasis>
: 
<specimenCode id="DBA4C70FCC07B519480B9CF7FD6AFD3A" box="[586,715,687,711]" pageId="3" pageNumber="18">PFN001814</specimenCode>
</paragraph>
<paragraph id="8BBD6F74CC07B5194AD69C89FD4FFCD7" blockId="3.[151,772,389,1572]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69C89FE96FD14" bold="true" box="[151,311,721,745]" pageId="3" pageNumber="18">Type locality.</emphasis>
<collectingCounty id="62DC17F8CC07B5194B029C89FDBDFD14" box="[323,540,721,745]" pageId="3" pageNumber="18">Aşağıgüney Village</collectingCounty>
of the city of 
<collectingCounty id="62DC17F8CC07B519488E9C89FF5FFCF4" pageId="3" pageNumber="18">Beypazarı</collectingCounty>
in the province of 
<collectingRegion id="49C6A196CC07B5194B929CAAFD80FCF7" box="[467,545,754,778]" country="Turkey" name="Ankara" pageId="3" pageNumber="18">Ankara</collectingRegion>
(
<collectionCode id="ED13F7B1CC07B51948759CAAFDCEFCF7" box="[564,623,754,778]" pageId="3" pageNumber="18">AGU</collectionCode>
). This region is called 
<location id="8EDD39AFCC07B5194B5A9D4BFDE3FCD6" LSID="urn:lsid:plazi:treatment:03ABDE62CC07B5104AD69F7BFEAFFC2D:8EDD39AFCC07B5194B5A9D4BFDE3FCD6" box="[283,578,787,811]" county="Asagiguney Village" municipality="Beypazari" name="Galatian Volcanic Province" pageId="3" pageNumber="18" stateProvince="Ankara">Galatian Volcanic Province</location>
(GPV) (
<figureCitation id="133973F1CC07B51948DE9D4BFD7CFCD6" box="[671,733,787,811]" captionStart="Fig" captionStartId="1.[230,263,1449,1470]" captionTargetBox="[265,1326,226,1402]" captionTargetId="figure-343@1.[230,1358,187,1440]" captionTargetPageId="1" captionText="Fig. 1 The location map of the studies samples in Turkey, and the generalized stratigraphic section of the region covering the western parts of the Galatia Massif (Akbaş et al., 2002) and zone of fossils (Hançili Formation). The detailed geological structure of the area was given in Akkemik et al. (2016) and Acarca Bayam et al. (2018)." figureDoi="http://doi.org/10.5281/zenodo.10520762" httpUri="https://zenodo.org/record/10520762/files/figure.png" pageId="3" pageNumber="18">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69D6CFDA9FC90" pageId="3" pageNumber="18" type="description">
<paragraph id="8BBD6F74CC07B5194AD69D6CFED7FCB1" blockId="3.[151,772,389,1572]" box="[151,374,820,844]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69D6CFF6BFCB1" bold="true" box="[151,202,820,844]" pageId="3" pageNumber="18">Age.</emphasis>
Early Miocene.
</paragraph>
<paragraph id="8BBD6F74CC07B5194AD69D0DFDA9FC90" blockId="3.[151,772,389,1572]" box="[151,520,853,877]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69D0DFE97FC90" bold="true" box="[151,310,853,877]" pageId="3" pageNumber="18">
<typeStatus id="54B9D1D6CC07B5194AD69D0DFF70FC90" box="[151,209,853,877]" pageId="3" pageNumber="18">Type</typeStatus>
horizon.
</emphasis>
Hançili Formation.
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B5194AD69D2EFDBFF9D9" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="8BBD6F74CC07B5194AD69D2EFDBFF9D9" blockId="3.[151,772,389,1572]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B5194AD69D2EFEAFFC73" bold="true" box="[151,270,886,910]" pageId="3" pageNumber="18">Diagnosis:</emphasis>
Growth ring boundaries distinct with 2-3 rows of flattened latewood tracheids. False rings commonly present. Transition from earlywood to latewood indistinct. Tracheids generally circular, and intercellular spaces clearly obvious and occur throughout the wood. Normal axial resin canals absent. Axial parenchyma appears both diffuse (in the transition from earlywood to latewood and within the latewood itself) and tangentially zonate. Rays uniseriate, and sometimes partly biseriate. Ray height is 1-6 (max.16) cells. Tracheidal pits on tangential walls common and 1-2 seriate, irregular or slightly alternate. The rays with one cell height conspicuously longer. Transverse end walls of axial parenchyma cells are nodular and smooth. Ray tracheids absent. End walls of ray parenchyma cells nodular and smooth, horizontal walls of rays sparsely pitted. Tracheid pitting in radial walls of the earlywood is uniseriate and biseriate, opposite, frequent, contiguous, and sometimes spaced. Cross-field pitting is cupressoid with 2–3 (-5) pits per cross-field.
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B51949719EC6FB78FBA9" pageId="3" pageNumber="18" type="description">
<paragraph id="8BBD6F74CC07B51949719EC6FACEFF2A" blockId="3.[816,1437,158,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719EC6FC1CFF4B" bold="true" box="[816,957,158,182]" pageId="3" pageNumber="18">Description:</emphasis>
Wood anatomical characteristics of the specimen of 
<taxonomicName id="4C0214F7CC07B519499E9EE7FBD6FF2A" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" box="[991,1143,191,215]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC07B519499E9EE7FBD6FF2A" box="[991,1143,191,215]" italics="true" pageId="3" pageNumber="18">Juniperoxylon</emphasis>
</taxonomicName>
were given as follows:
</paragraph>
<paragraph id="8BBD6F74CC07B51949719EB8FB2AFDDF" blockId="3.[816,1437,158,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719EB8FBB8FF05" bold="true" box="[816,1049,224,248]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719EB8FBB1FF05" bold="true" box="[816,1040,224,248]" italics="true" pageId="3" pageNumber="18">Transversal section</emphasis>
:
</emphasis>
Growth ring boundaries distinct with 2-3 rows of flattened latewood tracheids. False rings commonly present (
<figureCitation id="133973F1CC07B5194E2A9F7AFB0CFEC7" box="[1131,1197,290,314]" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: a-c). Transition from earlywood to latewood gradual and indistinct. Tracheids generally circular, and intercellular spaces clearly obvious and occur throughout the wood. Normal axial resin canals absent. Axial parenchyma presents and both diffuse (in the transition from earlywood to latewood and within the latewood itself) and tangentially zonate (
<figureCitation id="133973F1CC07B51949EA9C52FC4DFDDF" box="[939,1004,522,546]" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: b (arrows)-c).
</paragraph>
<paragraph id="8BBD6F74CC07B51949719C73FA29FCD6" blockId="3.[816,1437,158,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719C73FBB0FDBE" bold="true" box="[816,1041,555,579]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719C73FBA9FDBE" bold="true" box="[816,1032,555,579]" italics="true" pageId="3" pageNumber="18">Tangential section</emphasis>
:
</emphasis>
Rays exclusively uniseriate, and sometimes partly biseriate (
<figureCitation id="133973F1CC07B5194E3E9C14FB60FD99" box="[1151,1217,588,612]" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: d). Ray height is 2- 6 (max.16) cells (
<figureCitation id="133973F1CC07B5194E529C35FBF7FD78" box="[1043,1110,621,645]" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: d-e and 
<figureCitation id="133973F1CC07B5194EFB9C35FB5DFD78" box="[1210,1276,621,645]" captionStart="Fig" captionStartId="6.[215,248,1853,1874]" captionTargetBox="[230,1363,188,1812]" captionTargetId="figure-48@6.[225,1363,183,1812]" captionTargetPageId="6" captionText="Fig. 4 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of trache- ids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows)." figureDoi="http://doi.org/10.5281/zenodo.10520768" httpUri="https://zenodo.org/record/10520768/files/figure.png" pageId="3" pageNumber="18">Fig. 4</figureCitation>
: a). Tracheidal pits on tangential walls commonly present, and uni- to biseriate, irregularly spaced or slightly alternate (
<figureCitation id="133973F1CC07B5194F309CF7FCC7FD14" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: e-f). The rays with one cell height conspicuously longer (
<figureCitation id="133973F1CC07B51949EA9CAAFC4FFCF7" box="[939,1006,754,778]" captionStart="Fig" captionStartId="5.[234,267,1792,1813]" captionTargetBox="[251,1342,151,1750]" captionTargetId="figure-33@5.[246,1342,151,1750]" captionTargetPageId="5" captionText="Fig. 3 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids." figureDoi="http://doi.org/10.5281/zenodo.10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="3" pageNumber="18">Fig. 3</figureCitation>
: d-e). Transverse end walls of axial parenchyma cells are nodular and smooth (
<figureCitation id="133973F1CC07B5194F499D4BFAEBFCD6" box="[1288,1354,787,811]" captionStart="Fig" captionStartId="6.[215,248,1853,1874]" captionTargetBox="[230,1363,188,1812]" captionTargetId="figure-48@6.[225,1363,183,1812]" captionTargetPageId="6" captionText="Fig. 4 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of trache- ids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows)." figureDoi="http://doi.org/10.5281/zenodo.10520768" httpUri="https://zenodo.org/record/10520768/files/figure.png" pageId="3" pageNumber="18">Fig. 4</figureCitation>
: b-d).
</paragraph>
<paragraph id="8BBD6F74CC07B51949719D6CFB78FBA9" blockId="3.[816,1437,158,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719D6CFC79FCB1" bold="true" box="[816,984,820,844]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719D6CFC6EFCB1" bold="true" box="[816,975,820,844]" italics="true" pageId="3" pageNumber="18">Radial section</emphasis>
:
</emphasis>
Ray tracheids absent. Tracheid pitting in radial walls of the earlywood is uniseriate and biseriate, opposite, frequent, contiguous (
<figureCitation id="133973F1CC07B5194EA69D2EFA8FFC73" box="[1255,1326,886,910]" captionStart="Fig" captionStartId="6.[215,248,1853,1874]" captionTargetBox="[230,1363,188,1812]" captionTargetId="figure-48@6.[225,1363,183,1812]" captionTargetPageId="6" captionText="Fig. 4 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of trache- ids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows)." figureDoi="http://doi.org/10.5281/zenodo.10520768" httpUri="https://zenodo.org/record/10520768/files/figure.png" pageId="3" pageNumber="18">Fig. 4</figureCitation>
: e-f), and sometimes spaced (
<figureCitation id="133973F1CC07B5194E729DCFFBDAFC52" box="[1075,1147,919,943]" captionStart="Fig" captionStartId="7.[151,184,1947,1968]" captionTargetBox="[156,1365,168,1905]" captionTargetId="figure-16@7.[152,1365,163,1905]" captionTargetPageId="7" captionText="Fig. 5 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field." figureDoi="http://doi.org/10.5281/zenodo.10520770" httpUri="https://zenodo.org/record/10520770/files/figure.png" pageId="3" pageNumber="18">Fig. 5</figureCitation>
: c). Crassulae sometimes presents (
<figureCitation id="133973F1CC07B51949FE9DE0FB83FC2D" box="[959,1058,952,976]" captionStart="Fig" captionStartId="6.[215,248,1853,1874]" captionTargetBox="[230,1363,188,1812]" captionTargetId="figure-48@6.[225,1363,183,1812]" captionTargetPageId="6" captionText="Fig. 4 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of trache- ids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows)." figureDoi="http://doi.org/10.5281/zenodo.10520768" httpUri="https://zenodo.org/record/10520768/files/figure.png" pageId="3" pageNumber="18">Figure 4</figureCitation>
: f). End walls of ray parenchyma cells nodular and smooth, and horizontal walls of rays sparsely pitted (
<figureCitation id="133973F1CC07B5194E419DA2FBFFFBEF" box="[1024,1118,1018,1042]" captionStart="Fig" captionStartId="7.[151,184,1947,1968]" captionTargetBox="[156,1365,168,1905]" captionTargetId="figure-16@7.[152,1365,163,1905]" captionTargetPageId="7" captionText="Fig. 5 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field." figureDoi="http://doi.org/10.5281/zenodo.10520770" httpUri="https://zenodo.org/record/10520770/files/figure.png" pageId="3" pageNumber="18">Figure 5</figureCitation>
: a-d). Indenture rarely visible (
<figureCitation id="133973F1CC07B519491C9A43FC66FBCE" box="[861,967,1051,1075]" captionStart="Fig" captionStartId="7.[151,184,1947,1968]" captionTargetBox="[156,1365,168,1905]" captionTargetId="figure-16@7.[152,1365,163,1905]" captionTargetPageId="7" captionText="Fig. 5 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field." figureDoi="http://doi.org/10.5281/zenodo.10520770" httpUri="https://zenodo.org/record/10520770/files/figure.png" pageId="3" pageNumber="18">Figure 5d</figureCitation>
). Cross-field pitting is cupressoid with 2–3 (-5) pits per cross-field (
<figureCitation id="133973F1CC07B5194E1B9A64FB38FBA9" box="[1114,1177,1084,1108]" captionStart="Fig" captionStartId="7.[151,184,1947,1968]" captionTargetBox="[156,1365,168,1905]" captionTargetId="figure-16@7.[152,1365,163,1905]" captionTargetPageId="7" captionText="Fig. 5 The thin sections of Juniperoxylon acarcae Akkemik sp. nov. a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field." figureDoi="http://doi.org/10.5281/zenodo.10520770" httpUri="https://zenodo.org/record/10520770/files/figure.png" pageId="3" pageNumber="18">Fig. 5</figureCitation>
: e-g).
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B51949719A06FBEAF99B" pageId="3" pageNumber="18" type="discussion">
<paragraph id="8BBD6F74CC07B51949719A06FBEAF99B" blockId="3.[816,1437,158,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949719A06FC10FB8B" bold="true" box="[816,945,1118,1142]" pageId="3" pageNumber="18">Discussion:</emphasis>
According to the discussion by 
<bibRefCitation id="EF931285CC07B5194F749A05FC5AFB6A" author="Ruiz, D. P. &amp; Bodnar, J." pageId="3" pageNumber="18" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
, the features of distinct growth ring boundary, narrow latewood band, common spaces among tracheid, presence of pits on tangential walls of tracheids, presence of diffuse and/or zonate axial parenchyma, cupressoid or taxodioid 
<typeStatus id="54B9D1D6CC07B5194F449B5CFA92FAE1" box="[1285,1331,1284,1308]" pageId="3" pageNumber="18">type</typeStatus>
of cross field pits, low height of rays, generally nodular end walls of axial parenchyma, and homocellular rays are the main characteristics of 
<taxonomicName id="4C0214F7CC07B5194E369B3EFAAEFA83" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" box="[1143,1295,1382,1406]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC07B5194E369B3EFAAEFA83" box="[1143,1295,1382,1406]" italics="true" pageId="3" pageNumber="18">Juniperoxylon</emphasis>
</taxonomicName>
. Due to having very close wood anatomical characteristics with 
<bibRefCitation id="EF931285CC07B51949179BF1FBD6FA3C" author="Ruiz, D. P. &amp; Bodnar, J." box="[854,1143,1449,1473]" pageId="3" pageNumber="18" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
’s description, the present fossil specimen was identified as 
<taxonomicName id="4C0214F7CC07B5194EBC9B92FA34FA1F" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" box="[1277,1429,1482,1506]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC07B5194EBC9B92FA34FA1F" box="[1277,1429,1482,1506]" italics="true" pageId="3" pageNumber="18">Juniperoxylon</emphasis>
</taxonomicName>
. Based on the differences in the characteristics of the fossil species (
<tableCitation id="C6805ACFCC07B51949BD9854FBF3F9D9" box="[1020,1106,1548,1572]" captionStart="Table 1" captionStartId="8.[521,586,418,442]" captionTargetBox="[112,1970,497,1772]" captionTargetId="graphics-77@8.[97,1987,1049,1152]" captionTargetPageId="8" captionText="Table 1 Comparison of the new species with the former fossil species (Ruiz and Bodnar, 2019)" pageId="3" pageNumber="18">Table 1</tableCitation>
), the following identification key was prepared to find the exact place of the present 
<taxonomicName id="4C0214F7CC07B51949179816FBE6F99B" box="[854,1095,1614,1638]" pageId="3" pageNumber="18">
<emphasis id="B976B366CC07B51949179816FC4EF99B" box="[854,1007,1614,1638]" italics="true" pageId="3" pageNumber="18">Juniperoxylon</emphasis>
species
</taxonomicName>
:
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC07B51E4A8798CDFB4BFB69" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="key">
<paragraph id="8BBD6F74CC07B5194A8798CDFE78F950" blockId="3.[198,1420,1685,1960]" box="[198,473,1685,1710]" pageId="3" pageNumber="18">1A. Ray tracheids present</paragraph>
<paragraph id="8BBD6F74CC07B5194A8798ECFABEF916" blockId="3.[198,1420,1685,1960]" pageId="3" pageNumber="18">
2A. Cross-field pits 4-6 cupressoid; end walls of axial parenchyma cells smooth; radial tracheidal pitting spaced and contiguous
<emphasis id="B976B366CC07B5194B2E988BFABEF916" bold="true" box="[367,1311,1747,1771]" pageId="3" pageNumber="18">
&gt; 
<taxonomicName id="4C0214F7CC07B5194BC5988BFABEF916" authority="Bonetti, 1966 emend. Bodnar &amp; Artabe, 2007" authorityName="Bonetti, 1966 emend. Bodnar &amp; Artabe" authorityYear="2007" box="[388,1311,1747,1771]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="ischigualastense">
<emphasis id="B976B366CC07B5194BC5988BFCB7F916" bold="true" box="[388,790,1747,1771]" italics="true" pageId="3" pageNumber="18">Protojuniperoxylon ischigualastense</emphasis>
<bibRefCitation id="EF931285CC07B519495F988BFC12F916" author="Bonetti, M. I. R." box="[798,947,1747,1771]" pageId="3" pageNumber="18" pagination="211 - 218" refId="ref5786" refString="Bonetti, M. I. R., 1966. Protojuniperoxylon ischigualastensis sp. nov. Del Triasico de Ischigualasto (San Juan). Ameghiniana 4: 211 - 218." type="journal article" year="1966">Bonetti, 1966</bibRefCitation>
emend. 
<bibRefCitation id="EF931285CC07B5194E52988BFABEF916" author="Bodnar, J. &amp; Artabe, A. E." box="[1043,1311,1747,1771]" pageId="3" pageNumber="18" pagination="303 - 319" refId="ref5743" refString="Bodnar, J. &amp; Artabe, A. E., 2007. Estudio sistematico y paleodendrologico del leno de una Cupressaceae triasica de la Formacion Ischigualasto provincia de San Juan, Argentina. Ameghiniana, 44: 303 - 319." type="journal article" year="2007">Bodnar &amp; Artabe, 2007</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC07B5194A8798A9FC80F8D4" blockId="3.[198,1420,1685,1960]" pageId="3" pageNumber="18">
2B. Cross-field pits 1-3 cupressoid and taxodioid; end walls of axial parenchyma nodular; radial tracheidal pits sparse
<emphasis id="B976B366CC07B5194B499949FC80F8D4" bold="true" box="[264,801,1809,1833]" pageId="3" pageNumber="18">
&gt; 
<taxonomicName id="4C0214F7CC07B5194B5F9949FC80F8D4" authority="van der Burgh 1973" authorityName="van der Burgh" authorityYear="1973" box="[286,801,1809,1833]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="rhenanum">
<emphasis id="B976B366CC07B5194B5F9949FD99F8D4" bold="true" box="[286,568,1809,1833]" italics="true" pageId="3" pageNumber="18">Juniperoxylon rhenanum</emphasis>
<bibRefCitation id="EF931285CC07B519487E9949FC80F8D4" author="van der Burgh, J." box="[575,801,1809,1833]" pageId="3" pageNumber="18" pagination="73 - 275" refId="ref6680" refString="van der Burgh, J., 1973. Holzer der niederrheinischen braunkohlenformation, 2. Holzer der braunkohlengruben &quot; Maria Theresia &quot; zu herzogenrath, &quot; Zukunft West &quot; zu eschweiler und &quot; Victor &quot; (Zulpich mitte) zu Zulpich. Nebst einer systematisch-anatomischen bearbeitung der gattung Pinus L. Review of Palaeobotany and Palynology, 15 / 2: 73 - 275." type="journal article" year="1973">van der Burgh 1973</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC07B5194A879977FE6EF8BA" blockId="3.[198,1420,1685,1960]" box="[198,463,1839,1864]" pageId="3" pageNumber="18">1B. Ray tracheids absent</paragraph>
<paragraph id="8BBD6F74CC07B5194A879916FEC9F855" blockId="3.[198,1420,1685,1960]" pageId="3" pageNumber="18">
3A. Axial parenchyma absent or scarce, if present, diffuse; Ray parenchyma horizontal walls smooth; 4-6 pits per cross-field; radial tracheidal pitting spaced and contiguous
<emphasis id="B976B366CC07B51949109937FEC9F855" bold="true" pageId="3" pageNumber="18">
&gt; 
<taxonomicName id="4C0214F7CC07B519492B9937FEC9F855" authority="(Stopes, 1915) Eckhold, 1923" authorityName="Eckhold" authorityYear="1923" baseAuthorityName="Stopes" baseAuthorityYear="1915" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="maidstonense">
<emphasis id="B976B366CC07B519492B9937FB41F87A" bold="true" box="[874,1248,1903,1927]" italics="true" pageId="3" pageNumber="18">Protojuniperoxylon maidstonense</emphasis>
(
<bibRefCitation id="EF931285CC07B5194EB09937FA25F87A" author="Stopes, M. C." box="[1265,1412,1903,1927]" pageId="3" pageNumber="18" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) 
<bibRefCitation id="EF931285CC07B5194A8799C8FEC9F855" author="Eckhold, W." box="[198,360,1936,1960]" pageId="3" pageNumber="18" pagination="472 - 505" refId="ref5859" refString="Eckhold, W., 1923. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch der preussischen geologischen Landesanstalt zu Berlin, 42: 472 - 505." type="journal article" year="1923">Eckhold, 1923</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AA19EFCFD9DFF46" blockId="4.[224,1426,164,1172]" box="[224,572,164,188]" pageId="4" pageNumber="19">3B. Axial parenchyma common.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B479E9AFD37FF27" blockId="4.[224,1426,164,1172]" box="[262,662,194,218]" pageId="4" pageNumber="19">4A. Zonate axial parenchyma present</paragraph>
<paragraph id="8BBD6F74CC00B51E4B6D9EB9FCFDFECA" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
5A. Cross-field pits only taxodioid; 2-6 pits per cross-field; end walls of axial parenchyma smooth or irregularly pitted; ray height 1-14 cells; horizontal end walls of rays smooth or rarely pitted
<emphasis id="B976B366CC00B51E4EB79F58FA30FEE5" bold="true" box="[1270,1425,256,280]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E4F4D9F58FCFDFECA" authority="Watari &amp; Nishida 1973" authorityName="Watari &amp; Nishida" authorityYear="1973" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="breviparenchmatosum">
<emphasis id="B976B366CC00B51E4B6D9F47FCFDFECA" bold="true" box="[300,860,287,311]" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E4F4D9F58FA30FEE5" bold="true" box="[1292,1425,256,280]" italics="true" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E4B6D9F47FEF1FECA" bold="true" box="[300,336,287,311]" italics="true" pageId="4" pageNumber="19">Juniperoxylon</emphasis>
</emphasis>
<emphasis id="B976B366CC00B51E4B179F47FDEDFECA" bold="true" box="[342,588,287,311]" italics="true" pageId="4" pageNumber="19">breviparenchmatosum</emphasis>
<bibRefCitation id="EF931285CC00B51E48129F47FCFDFECA" author="Watari, S. &amp; Nishida, M." box="[595,860,287,311]" pageId="4" pageNumber="19" pagination="154 - 159" refId="ref6746" refString="Watari, S. &amp; Nishida, M., 1973. A Juniperoxylon from the Tertiary of Hokkaido. Journal of Japanese Botany 48: 154 - 159." type="journal article" year="1973">Watari &amp; Nishida 1973</bibRefCitation>
</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B6D9F65FC11FEA8" blockId="4.[224,1426,164,1172]" box="[300,944,317,341]" pageId="4" pageNumber="19">5B. Cross-field pits only cupressoid, and 1-6 per cross-field.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B109F04FBD0FE4F" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
6A. Horizontal walls of rays smooth; ray width uniseriate and rarely partly biseriate; end walls of axial parenchyma smooth or nodular; radial wall pitting uniseriate, rarely bi-to triseriate pits opposite, and spaced
<emphasis id="B976B366CC00B51E4B8F9FC2FBD0FE4F" bold="true" box="[462,1137,410,434]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E4BA59FC2FBD0FE4F" authority="(Goppert, 1850) Krausel, 1949" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Goppert" baseAuthorityYear="1850" box="[484,1137,410,434]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="pachyderma">
<emphasis id="B976B366CC00B51E4BA59FC2FCB1FE4F" bold="true" box="[484,784,410,434]" italics="true" pageId="4" pageNumber="19">Juniperoxylon pachyderma</emphasis>
(Göppert, 1850) 
<bibRefCitation id="EF931285CC00B51E49929FC2FBD0FE4F" author="Krausel, R." box="[979,1137,410,434]" pageId="4" pageNumber="19" pagination="83 - 203" refId="ref6132" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungen zur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica B, 89: 83 - 203." type="journal article" year="1949">Kräusel, 1949</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B109FE0FC95FE2D" blockId="4.[224,1426,164,1172]" box="[337,820,440,465]" pageId="4" pageNumber="19">6B. Horizontal walls of rays distinctly pitted.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B369F8FFD18FDD1" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
7A. Radial wall pitting exclusively uniseriate, horizontal walls of rays distinctly pitted; ray width uniseriate; transversal end walls of axial parenchyma smooth
<emphasis id="B976B366CC00B51E4E099FAEFA30FDF3" bold="true" box="[1096,1425,502,526]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E4E249FAEFDB7FDD1" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E4E249FAEFA30FDF3" bold="true" box="[1125,1425,502,526]" italics="true" pageId="4" pageNumber="19">Juniperoxylon pottoniense</emphasis>
<emphasis id="B976B366CC00B51E4B369C4CFD18FDD1" bold="true" box="[375,697,532,556]" pageId="4" pageNumber="19">
(
<bibRefCitation id="EF931285CC00B51E4B3E9C4CFDAEFDD1" author="Stopes, M. C." box="[383,527,532,556]" pageId="4" pageNumber="19" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) 
<bibRefCitation id="EF931285CC00B51E485C9C4CFD18FDD1" author="Krausel, R." box="[541,697,532,556]" pageId="4" pageNumber="19" pagination="83 - 203" refId="ref6132" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungen zur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica B, 89: 83 - 203." type="journal article" year="1949">Kräusel, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B369C6BFA96FDB7" blockId="4.[224,1426,164,1172]" box="[375,1335,562,586]" pageId="4" pageNumber="19">7B. Radial wall pitting uni-to biseriate; transversal end walls of axial parenchyma nodular</paragraph>
<paragraph id="8BBD6F74CC00B51E4BDC9C09FDFFFD3B" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
8A. Tracheidal pits on radial walls often frequent and contiguous; rays uni- to biseriate; crassulae rarely present, end walls of axial parenchyma nodular and smooth; end walls of rays nodular and smooth; horizontal walls of rays sparsely pitted
<emphasis id="B976B366CC00B51E4E2F9CD7FA30FD5A" bold="true" box="[1134,1425,655,679]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E4ECD9CD7FDA7FD3B" authority="Akkemik, 2021" authorityName="Akkemik" authorityYear="2021" pageId="4" pageNumber="19" status="sp.nov.">
<emphasis id="B976B366CC00B51E4ECD9CD7FA30FD5A" bold="true" box="[1164,1425,655,679]" italics="true" pageId="4" pageNumber="19">Juniperoxylon acarcae</emphasis>
<emphasis id="B976B366CC00B51E4BDC9CF6FDFFFD3B" bold="true" box="[413,606,686,710]" pageId="4" pageNumber="19">
Akkemik 
<taxonomicNameLabel id="A2450E1DCC00B51E484F9CF6FDFFFD3B" box="[526,606,686,710]" pageId="4" pageNumber="19" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4BDC9C94FA2CFCFE" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
8B. Spacing of radial tracheidal pits sparse; rays uniseriate; crassulae present; ray parenchyma horizontal walls distinctly pitted
<emphasis id="B976B366CC00B51E48BC9CB3FA2CFCFE" bold="true" box="[765,1421,747,771]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E49539CB3FA2CFCFE" authority="(Kownas, 1951) Huard, 1966" authorityName="Huard" authorityYear="1966" baseAuthorityName="Kownas" baseAuthorityYear="1951" box="[786,1421,747,771]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="juniperoides">
<emphasis id="B976B366CC00B51E49539CB3FBE0FCFE" bold="true" box="[786,1089,747,771]" italics="true" pageId="4" pageNumber="19">Juniperoxylon juniperoides</emphasis>
(Kownas, 1951) 
<bibRefCitation id="EF931285CC00B51E4EBE9CB3FA2CFCFE" author="Huard, J." box="[1279,1421,747,771]" pageId="4" pageNumber="19" pagination="1 - 85" refId="ref5943" refString="Huard, J., 1966. Etude anatomique des bois de coniferes des couches a lignite neogenes des Landes. Memoires de la Societe geologique de France 105: 1 - 85." type="journal article" year="1966">Huard, 1966</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B479D51FC42FCDC" blockId="4.[224,1426,164,1172]" box="[262,995,777,801]" pageId="4" pageNumber="19">4B. Zonate axial parenchyma absent, axial parenchyma only diffuse.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B6D9D70FCEBFC83" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
9A. Axial parenchyma scarce; ray heights 2-34 cells; ray parenchyma horizontal walls distinctly pitted; 1-2 (3-4) pits per cross-field; radial tracheidal pitting spaced but not contiguous
<emphasis id="B976B366CC00B51E4ECE9D1FFA33FCA2" bold="true" box="[1167,1426,839,863]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E4EE79D1FFDF9FC83" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC00B51E4EE79D1FFA33FCA2" bold="true" box="[1190,1426,839,863]" italics="true" pageId="4" pageNumber="19">
Juniperoxylon 
<emphasis id="B976B366CC00B51E4B6D9D3EFCEBFC83" bold="true" box="[300,842,870,894]" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E4B6D9D3EFECBFC83" bold="true" box="[300,362,870,894]" italics="true" pageId="4" pageNumber="19">zamunerae</emphasis>
(Bodnar et al., 2015) 
<bibRefCitation id="EF931285CC00B51E481E9D3EFCEBFC83" author="Ruiz, D. P. &amp; Bodnar, J." box="[607,842,870,894]" pageId="4" pageNumber="19" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz &amp; Bodnar 2019</bibRefCitation>
</emphasis>
</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B6D9DDCFBDBFC61" blockId="4.[224,1426,164,1172]" box="[300,1146,900,925]" pageId="4" pageNumber="19">9B. Axial parenchyma present, abundant, and diffuse; ray height up to 18 cells.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B109DFBFBA4FC27" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
10A. Ray parenchyma horizontal walls smooth; end walls of axial parenchyma cells nodular; ray heights 1-5 cells
<emphasis id="B976B366CC00B51E48469D9AFBA4FC27" bold="true" box="[519,1029,962,986]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E485C9D9AFBA4FC27" authority="Suss &amp; Rather 1998" authorityName="Suss &amp; Rather" authorityYear="1998" box="[541,1029,962,986]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="wagneri">
<emphasis id="B976B366CC00B51E485C9D9AFCBDFC27" bold="true" box="[541,796,962,986]" italics="true" pageId="4" pageNumber="19">Juniperoxylon wagneri</emphasis>
Süss &amp; Rather 1998
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B109DB8FAE3FC05" blockId="4.[224,1426,164,1172]" box="[337,1346,992,1017]" pageId="4" pageNumber="19">10B. Ray parenchyma horizontal walls sparsely or distinctly pitted; Ray heights up to 18 cells.</paragraph>
<paragraph id="8BBD6F74CC00B51E4B369DA7FE0CFBA9" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
11A. Horizontal walls of ray parenchyma sparsely pitted; rays uniseriate and occasionally biseriate; axial parenchyma abundant; crassulae present
<emphasis id="B976B366CC00B51E49EB9A46FA33FBCB" bold="true" box="[938,1426,1054,1078]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E49859A46FA33FBCB" authority="Dolezych 2016" authorityName="Dolezych" authorityYear="2016" box="[964,1426,1054,1078]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="schneiderianum">
<emphasis id="B976B366CC00B51E49859A46FABEFBCB" bold="true" box="[964,1311,1054,1078]" italics="true" pageId="4" pageNumber="19">Juniperoxylon schneiderianum</emphasis>
<bibRefCitation id="EF931285CC00B51E4F6B9A46FA33FBCB" author="Dolezych, M." box="[1322,1426,1054,1078]" pageId="4" pageNumber="19" pagination="5 - 31" refId="ref5819" refString="Dolezych, M., 2016. A remarkable extinct wood from Lusatia (central Europe) - Juniperoxylon schneiderianum sp. nov. with affinity to Cupressospermum saxonicum Mai. Palaeontographica B, 295: 5 - 31." type="journal article" year="2016">Dolezych</bibRefCitation>
</taxonomicName>
</emphasis>
<emphasis id="B976B366CC00B51E4B369A64FE0CFBA9" bold="true" box="[375,429,1084,1108]" pageId="4" pageNumber="19">2016</emphasis>
</paragraph>
<paragraph id="8BBD6F74CC00B51E4B369A03FB4BFB69" blockId="4.[224,1426,164,1172]" pageId="4" pageNumber="19">
11B. Horizontal walls of ray parenchyma distinctly pitted; rays uniseriate; axial parenchyma present; crassulae absent
<emphasis id="B976B366CC00B51E494E9A24FB4BFB69" bold="true" box="[783,1258,1148,1172]" pageId="4" pageNumber="19">
&gt; 
<taxonomicName id="4C0214F7CC00B51E49649A24FB4BFB69" authority="Houlbert, 1910" authorityName="Houlbert" authorityYear="1910" box="[805,1258,1148,1172]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="turonense">
<emphasis id="B976B366CC00B51E49649A24FB99FB69" bold="true" box="[805,1080,1148,1172]" italics="true" pageId="4" pageNumber="19">Juniperoxylon turonense</emphasis>
<bibRefCitation id="EF931285CC00B51E4E7E9A24FB4BFB69" author="Houlbert, C." box="[1087,1258,1148,1172]" pageId="4" pageNumber="19" pagination="70 - 76" refId="ref5919" refString="Houlbert, C., 1910. Les bois des faluns de Touraine. Feuille des jeunes Naturalistes 40: 70 - 76." type="journal article" year="1910">Houlbert, 1910</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3183CFFCC00B5104AD69A98FEAFFC2D" lastPageId="10" lastPageNumber="25" pageId="4" pageNumber="19" type="discussion">
<paragraph id="8BBD6F74CC00B51E4AD69A98FDDFFAE7" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
According to the identification key and the given characteristics in 
<tableCitation id="C6805ACFCC00B51E4B519AB9FEC2FB04" box="[272,355,1249,1273]" captionStart="Table 1" captionStartId="8.[521,586,418,442]" captionTargetBox="[112,1970,497,1772]" captionTargetId="graphics-77@8.[97,1987,1049,1152]" captionTargetPageId="8" captionText="Table 1 Comparison of the new species with the former fossil species (Ruiz and Bodnar, 2019)" pageId="4" pageNumber="19">Table 1</tableCitation>
, the new fossil 
<taxonomicName id="4C0214F7CC00B51E48559AB9FCA2FB04" box="[532,771,1249,1273]" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E48559AB9FD0CFB04" box="[532,685,1249,1273]" italics="true" pageId="4" pageNumber="19">Juniperoxylon</emphasis>
species
</taxonomicName>
has differences from all former fossil species.
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD69B7BFE90FA62" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
<taxonomicName id="4C0214F7CC00B51E4AD69B7BFE8EFAA1" authority="(Bodnar et al., 2015) Ruiz &amp; Bodnar, 2019" authorityName="Ruiz &amp; Bodnar" authorityYear="2019" baseAuthorityName="Bodnar" baseAuthorityYear="2015" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="zamunerae">
<emphasis id="B976B366CC00B51E4AD69B7BFE11FAC6" box="[151,432,1315,1339]" italics="true" pageId="4" pageNumber="19">Juniperoxylon zamunerae</emphasis>
(Bodnar et al., 2015) 
<bibRefCitation id="EF931285CC00B51E48F09B7BFE8EFAA1" author="Ruiz, D. P. &amp; Bodnar, J." pageId="4" pageNumber="19" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz &amp; Bodnar, 2019</bibRefCitation>
</taxonomicName>
differs from the new species in having scarce and diffuse axial parenchyma, and 2-34 cells height of rays.
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD69BF0FDFFF9B9" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
<taxonomicName id="4C0214F7CC00B51E4AD69BF0FEAFFA1C" authority="(Stopes, 1915) Eckhold, 1923" authorityName="Eckhold" authorityYear="1923" baseAuthorityName="Stopes" baseAuthorityYear="1915" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="maidstonense">
<emphasis id="B976B366CC00B51E4AD69BF0FDABFA3D" box="[151,522,1448,1472]" italics="true" pageId="4" pageNumber="19">Protojuniperoxylon maidstonense</emphasis>
(
<bibRefCitation id="EF931285CC00B51E48639BF0FD18FA3D" author="Stopes, M. C." box="[546,697,1448,1472]" pageId="4" pageNumber="19" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) 
<bibRefCitation id="EF931285CC00B51E48909BF0FEAFFA1C" author="Eckhold, W." pageId="4" pageNumber="19" pagination="472 - 505" refId="ref5859" refString="Eckhold, W., 1923. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch der preussischen geologischen Landesanstalt zu Berlin, 42: 472 - 505." type="journal article" year="1923">Eckhold, 1923</bibRefCitation>
</taxonomicName>
differ from the present specimen in having scarce axial parenchyma and predominantly uniseriate tracheidal pitting on radial walls, and 4-6 cupressoid pits per cross-field (
<bibRefCitation id="EF931285CC00B51E4B7F9874FDEEF9B9" author="Bodnar, J. &amp; Artabe, A. E." box="[318,591,1580,1604]" pageId="4" pageNumber="19" pagination="303 - 319" refId="ref5743" refString="Bodnar, J. &amp; Artabe, A. E., 2007. Estudio sistematico y paleodendrologico del leno de una Cupressaceae triasica de la Formacion Ischigualasto provincia de San Juan, Argentina. Ameghiniana, 44: 303 - 319." type="journal article" year="2007">Bodnar and Artabe, 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD69815FE80F914" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
<taxonomicName id="4C0214F7CC00B51E4AD69815FD25F998" authority="Suss &amp; Rather 1998" authorityName="Suss &amp; Rather" authorityYear="1998" box="[151,644,1613,1637]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="wagneri">
<emphasis id="B976B366CC00B51E4AD69815FE32F998" box="[151,403,1613,1637]" italics="true" pageId="4" pageNumber="19">Juniperoxylon wagneri</emphasis>
Süss &amp; Rather 1998
</taxonomicName>
, 
<taxonomicName id="4C0214F7CC00B51E48D59816FEE6F95A" authority="Bonetti, 1966 emend. Bodnar &amp; Artabe, 2007" authorityName="Bonetti, 1966 emend. Bodnar &amp; Artabe" authorityYear="2007" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="ischigualastense">
<emphasis id="B976B366CC00B51E48D59816FE1BF97B" italics="true" pageId="4" pageNumber="19">Protojuniperoxylon ischigualastense</emphasis>
<bibRefCitation id="EF931285CC00B51E4B829836FDF7F97B" author="Bonetti, M. I. R." box="[451,598,1646,1670]" pageId="4" pageNumber="19" pagination="211 - 218" refId="ref5786" refString="Bonetti, M. I. R., 1966. Protojuniperoxylon ischigualastensis sp. nov. Del Triasico de Ischigualasto (San Juan). Ameghiniana 4: 211 - 218." type="journal article" year="1966">Bonetti, 1966</bibRefCitation>
emend. 
<bibRefCitation id="EF931285CC00B51E48F59836FEE6F95A" author="Bodnar, J. &amp; Artabe, A. E." pageId="4" pageNumber="19" pagination="303 - 319" refId="ref5743" refString="Bodnar, J. &amp; Artabe, A. E., 2007. Estudio sistematico y paleodendrologico del leno de una Cupressaceae triasica de la Formacion Ischigualasto provincia de San Juan, Argentina. Ameghiniana, 44: 303 - 319." type="journal article" year="2007">Bodnar &amp; Artabe, 2007</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C0214F7CC00B51E4BC798C8FCA2F95A" authority="Dolezych, 2016" authorityName="Dolezych" authorityYear="2016" box="[390,771,1679,1703]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="schneiderianum">
<emphasis id="B976B366CC00B51E4BC798C8FDECF95A" box="[390,589,1679,1703]" italics="true" pageId="4" pageNumber="19">J. schneiderianum</emphasis>
<bibRefCitation id="EF931285CC00B51E481698D7FCA2F95A" author="Dolezych, M." box="[599,771,1679,1703]" pageId="4" pageNumber="19" pagination="5 - 31" refId="ref5819" refString="Dolezych, M., 2016. A remarkable extinct wood from Lusatia (central Europe) - Juniperoxylon schneiderianum sp. nov. with affinity to Cupressospermum saxonicum Mai. Palaeontographica B, 295: 5 - 31." type="journal article" year="2016">Dolezych, 2016</bibRefCitation>
</taxonomicName>
differ from the new specimen in having only diffuse axial parenchyma.
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD698ABFE3BF8B0" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
<taxonomicName id="4C0214F7CC00B51E4AD698ABFF6CF8D1" authority="Watari &amp; Nishida, 1973" authorityName="Watari &amp; Nishida" authorityYear="1973" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="breviparenchmatosum">
<emphasis id="B976B366CC00B51E4AD698ABFD88F8F6" box="[151,553,1779,1803]" italics="true" pageId="4" pageNumber="19">Juniperoxylon breviparenchmatosum</emphasis>
<bibRefCitation id="EF931285CC00B51E487498ABFF6CF8D1" author="Watari, S. &amp; Nishida, M." pageId="4" pageNumber="19" pagination="154 - 159" refId="ref6746" refString="Watari, S. &amp; Nishida, M., 1973. A Juniperoxylon from the Tertiary of Hokkaido. Journal of Japanese Botany 48: 154 - 159." type="journal article" year="1973">Watari &amp; Nishida, 1973</bibRefCitation>
</taxonomicName>
differs from the new species in having only taxodioid 
<typeStatus id="54B9D1D6CC00B51E4A81996EFF4FF8B3" box="[192,238,1846,1870]" pageId="4" pageNumber="19">type</typeStatus>
cross-field pits.
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD6990EFD48F82C" blockId="4.[151,771,1216,2067]" pageId="4" pageNumber="19">
<taxonomicName id="4C0214F7CC00B51E4AD6990EFD0FF893" authority="van der Burgh, 1973" authorityName="van der Burgh" authorityYear="1973" box="[151,686,1878,1902]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="rhenanum">
<emphasis id="B976B366CC00B51E4AD6990EFE0AF893" box="[151,427,1878,1902]" italics="true" pageId="4" pageNumber="19">Juniperoxylon rhenanum</emphasis>
<bibRefCitation id="EF931285CC00B51E4BFB990FFD0FF893" author="van der Burgh, J." box="[442,686,1878,1902]" pageId="4" pageNumber="19" pagination="73 - 275" refId="ref6680" refString="van der Burgh, J., 1973. Holzer der niederrheinischen braunkohlenformation, 2. Holzer der braunkohlengruben &quot; Maria Theresia &quot; zu herzogenrath, &quot; Zukunft West &quot; zu eschweiler und &quot; Victor &quot; (Zulpich mitte) zu Zulpich. Nebst einer systematisch-anatomischen bearbeitung der gattung Pinus L. Review of Palaeobotany and Palynology, 15 / 2: 73 - 275." type="journal article" year="1973">van der Burgh, 1973</bibRefCitation>
</taxonomicName>
differs from the new species in having 1-3 cupressoid and taxodioid cross-field pits, space of radial tracheidal pits, much longer rays (1–40 cells), and crassulae formation.
</paragraph>
<paragraph id="8BBD6F74CC00B51E4AD69982FBF8FA63" blockId="4.[151,771,1216,2067]" lastBlockId="4.[816,1437,1216,2067]" pageId="4" pageNumber="19">
The closest fossil species are 
<taxonomicName id="4C0214F7CC00B51E4B9C9982FE43F7EE" authority="(Goppert, 1850) Krausel, 1949" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Goppert" baseAuthorityYear="1850" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="pachyderma">
<emphasis id="B976B366CC00B51E4B9C9982FCA2F80F" box="[477,771,2010,2034]" italics="true" pageId="4" pageNumber="19">Juniperoxylon pachyderma</emphasis>
(Göppert, 1850) 
<bibRefCitation id="EF931285CC00B51E4B0B99A3FE43F7EE" author="Krausel, R." box="[330,482,2043,2067]" pageId="4" pageNumber="19" pagination="83 - 203" refId="ref6132" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungen zur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica B, 89: 83 - 203." type="journal article" year="1949">Kräusel, 1949</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0214F7CC00B51E4BAC99A4FBAAFB25" authority="(Kownas, 1951) Huard, 1966" authorityName="Huard" authorityYear="1966" baseAuthorityName="Kownas" baseAuthorityYear="1951" box="[493,1035,1216,2067]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="juniperoides">
<emphasis id="B976B366CC00B51E4BAC99A4FD2AF7EE" box="[493,651,2043,2067]" italics="true" pageId="4" pageNumber="19">J. juniperoides</emphasis>
(Kownas, 1951) 
<bibRefCitation id="EF931285CC00B51E493D9A98FBAAFB25" author="Huard, J." box="[892,1035,1216,1240]" pageId="4" pageNumber="19" pagination="1 - 85" refId="ref5943" refString="Huard, J., 1966. Etude anatomique des bois de coniferes des couches a lignite neogenes des Landes. Memoires de la Societe geologique de France 105: 1 - 85." type="journal article" year="1966">Huard, 1966</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C0214F7CC00B51E4E109A99FC75FB04" authority="(Stopes, 1915) Krausel, 1949" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Stopes" baseAuthorityYear="1915" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="pottoniense">
<emphasis id="B976B366CC00B51E4E109A99FB4DFB25" box="[1105,1260,1217,1240]" italics="true" pageId="4" pageNumber="19">J. pottoniense</emphasis>
(
<bibRefCitation id="EF931285CC00B51E4F439A98FA35FB25" author="Stopes, M. C." box="[1282,1428,1216,1240]" pageId="4" pageNumber="19" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) 
<bibRefCitation id="EF931285CC00B51E49719AB9FC75FB04" author="Krausel, R." box="[816,980,1249,1273]" pageId="4" pageNumber="19" pagination="83 - 203" refId="ref6132" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungen zur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica B, 89: 83 - 203." type="journal article" year="1949">Kräusel, 1949</bibRefCitation>
</taxonomicName>
. These three fossil species have only spaced radial tracheidal pits. The new fossil species has both spaced and contiguous radial tracheidal pits. 
<taxonomicName id="4C0214F7CC00B51E4FCB9B7CFC15FAA1" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Goppert" baseAuthorityYear="1850" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="pachyderma">
<emphasis id="B976B366CC00B51E4FCB9B7CFC15FAA1" italics="true" pageId="4" pageNumber="19">J. pachyderma</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0214F7CC00B51E49AF9B1DFB2EFAA1" authorityName="Huard" authorityYear="1966" baseAuthorityName="Kownas" baseAuthorityYear="1951" box="[1006,1167,1348,1372]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="juniperoides">
<emphasis id="B976B366CC00B51E49AF9B1DFB2EFAA1" box="[1006,1167,1348,1372]" italics="true" pageId="4" pageNumber="19">J. juniperoides</emphasis>
</taxonomicName>
have also crassulae formation, and 
<taxonomicName id="4C0214F7CC00B51E49F89B3EFBEEFA80" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Stopes" baseAuthorityYear="1915" box="[953,1103,1382,1405]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="pottoniense">
<emphasis id="B976B366CC00B51E49F89B3EFBEEFA80" box="[953,1103,1382,1405]" italics="true" pageId="4" pageNumber="19">J. pottoniense</emphasis>
</taxonomicName>
has exclusively uniseriate pits on radial walls of tracheids.
</paragraph>
<paragraph id="8BBD6F74CC00B51E49719BF0FC26F8F6" blockId="4.[816,1437,1216,2067]" pageId="4" pageNumber="19">
Based on these differences, the specimen was described as a new 
<taxonomicName id="4C0214F7CC00B51E49D59B91FB20FA1C" box="[916,1153,1481,1505]" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E49D59B91FB8CFA1C" box="[916,1069,1481,1505]" italics="true" pageId="4" pageNumber="19">Juniperoxylon</emphasis>
species
</taxonomicName>
and named 
<taxonomicName id="4C0214F7CC00B51E4F459B91FC53F9FF" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC00B51E4F459B91FC27F9FF" italics="true" pageId="4" pageNumber="19">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC00B51E49BA9BB3FBF0F9FF" box="[1019,1105,1515,1538]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
The descriptions of 
<taxonomicName id="4C0214F7CC00B51E4F729BB3FA3DF9FF" baseAuthorityName="Ruiz and Bodnar" baseAuthorityYear="2019" box="[1331,1436,1515,1538]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC00B51E4F729BB3FA3DF9FF" box="[1331,1436,1515,1538]" italics="true" pageId="4" pageNumber="19">Juniperus</emphasis>
</taxonomicName>
from Hoçaş Fossil site (
<bibRefCitation id="EF931285CC00B51E4E7B9853FA86F9DE" author="Akkemik, U. &amp; Arslan M. &amp; Poole I. &amp; Tosun, S. &amp; Kose, N. &amp; Karlioglu Kilic, N. &amp; Aydin, A." box="[1082,1319,1547,1571]" pageId="4" pageNumber="19" pagination="31 - 50" refId="ref5392" refString="Akkemik, U., Arslan M., Poole I., Tosun, S., Kose, N., Karlioglu Kilic, N. &amp; Aydin, A., 2016. Silicified woods from two previously undescribed early Miocene forest sites near Seben, northwest Turkey. Review of Palaeobotany and Palynology 235: 31 - 50, https: // doi. org / 10.1016 / j. revpalbo. 2016.09.012" type="journal article" year="2016">Akkemik et al., 2016</bibRefCitation>
) and from Aşağıgüney Fossil site (
<bibRefCitation id="EF931285CC00B51E4E779874FAF8F9B9" author="Acarca Bayam, N. N. &amp; Akkemik, U. &amp; Poole, I." box="[1078,1369,1580,1605]" pageId="4" pageNumber="19" pagination="1 - 42" refId="ref5199" refString="Acarca Bayam, N. N., Akkemik, U., Poole, I., &amp; Akarsu. F., 2018. Further Contributions to the early Miocene forest vegetation of the Galatean Volcanic Province, Turkey. Palaeobotanica Electronica, 21.3.40 A 1 - 42. https: // doi. org / 10.26879 / 816 palaeoelectronica. org / content / 2018 / 2353 - trees-of-gvp-inmiocene" type="book chapter" year="2018">Acarca Bayam et al., 2018</bibRefCitation>
) were compared with the new species and results showed that they both had very similar features. It can be concluded that the new species, 
<taxonomicName id="4C0214F7CC00B51E4E7198D7FA3CF95A" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[1072,1437,1679,1703]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC00B51E4E7198D7FA8DF95A" box="[1072,1324,1679,1703]" italics="true" pageId="4" pageNumber="19">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC00B51E497198E9FCDFF935" box="[816,894,1713,1736]" pageId="4" pageNumber="19" rank="species">sp.nov.</taxonomicNameLabel>
has an area through the valley between Beypazarı- Aşağıgüney Village and Seben-Kozyaka Village (
<figureCitation id="133973F1CC00B51E497998ABFCD6F8F6" box="[824,887,1779,1803]" captionStart="Fig" captionStartId="1.[230,263,1449,1470]" captionTargetBox="[265,1326,226,1402]" captionTargetId="figure-343@1.[230,1358,187,1440]" captionTargetPageId="1" captionText="Fig. 1 The location map of the studies samples in Turkey, and the generalized stratigraphic section of the region covering the western parts of the Galatia Massif (Akbaş et al., 2002) and zone of fossils (Hançili Formation). The detailed geological structure of the area was given in Akkemik et al. (2016) and Acarca Bayam et al. (2018)." figureDoi="http://doi.org/10.5281/zenodo.10520762" httpUri="https://zenodo.org/record/10520762/files/figure.png" pageId="4" pageNumber="19">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BBD6F74CC00B51E4971994CFAB6F7EE" blockId="4.[816,1437,1216,2067]" pageId="4" pageNumber="19">
<emphasis id="B976B366CC00B51E4971994CFC02F8D1" bold="true" box="[816,931,1812,1836]" pageId="4" pageNumber="19">Affinities:</emphasis>
The fossil 
<taxonomicName id="4C0214F7CC00B51E4E25994CFB5CF8D1" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" box="[1124,1277,1812,1836]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC00B51E4E25994CFB5CF8D1" box="[1124,1277,1812,1836]" italics="true" pageId="4" pageNumber="19">Juniperoxylon</emphasis>
</taxonomicName>
descriptions (e.g.Vaudois and Privé, 1971; 
<bibRefCitation id="EF931285CC00B51E4EFA996DFC2EF893" author="Bodnar, J. &amp; Artabe, A. E." pageId="4" pageNumber="19" pagination="303 - 319" refId="ref5743" refString="Bodnar, J. &amp; Artabe, A. E., 2007. Estudio sistematico y paleodendrologico del leno de una Cupressaceae triasica de la Formacion Ischigualasto provincia de San Juan, Argentina. Ameghiniana, 44: 303 - 319." type="journal article" year="2007">Bodnar and Artabe, 2007</bibRefCitation>
; 
<bibRefCitation id="EF931285CC00B51E49DD990EFB8EF893" author="Klusek, M." box="[924,1071,1878,1902]" pageId="4" pageNumber="19" pagination="697 - 708" refId="ref6076" refString="Klusek, M., 2014. Miocene coniferous woods of the Polish Carpathian Foredeep. Acta Palaeontologica Polonica, 59 (3): 697 - 708." type="journal article" year="2014">Kłusek, 2014</bibRefCitation>
; 
<bibRefCitation id="EF931285CC00B51E4E7A990EFA82F893" author="Akkemik, U. &amp; Arslan M. &amp; Poole I. &amp; Tosun, S. &amp; Kose, N. &amp; Karlioglu Kilic, N. &amp; Aydin, A." box="[1083,1315,1878,1902]" pageId="4" pageNumber="19" pagination="31 - 50" refId="ref5392" refString="Akkemik, U., Arslan M., Poole I., Tosun, S., Kose, N., Karlioglu Kilic, N. &amp; Aydin, A., 2016. Silicified woods from two previously undescribed early Miocene forest sites near Seben, northwest Turkey. Review of Palaeobotany and Palynology 235: 31 - 50, https: // doi. org / 10.1016 / j. revpalbo. 2016.09.012" type="journal article" year="2016">Akkemik et al., 2016</bibRefCitation>
; 
<bibRefCitation id="EF931285CC00B51E4F6E990EFC2FF872" author="Dolezych, M." pageId="4" pageNumber="19" pagination="5 - 31" refId="ref5819" refString="Dolezych, M., 2016. A remarkable extinct wood from Lusatia (central Europe) - Juniperoxylon schneiderianum sp. nov. with affinity to Cupressospermum saxonicum Mai. Palaeontographica B, 295: 5 - 31." type="journal article" year="2016">Dolezych, 2016</bibRefCitation>
, 
<bibRefCitation id="EF931285CC00B51E49DF992FFB6AF872" author="Acarca Bayam, N. N. &amp; Akkemik, U. &amp; Poole, I." box="[926,1227,1911,1936]" pageId="4" pageNumber="19" pagination="1 - 42" refId="ref5199" refString="Acarca Bayam, N. N., Akkemik, U., Poole, I., &amp; Akarsu. F., 2018. Further Contributions to the early Miocene forest vegetation of the Galatean Volcanic Province, Turkey. Palaeobotanica Electronica, 21.3.40 A 1 - 42. https: // doi. org / 10.26879 / 816 palaeoelectronica. org / content / 2018 / 2353 - trees-of-gvp-inmiocene" type="book chapter" year="2018">Acarca Bayam et al., 2018</bibRefCitation>
) revealed that this 
<typeStatus id="54B9D1D6CC00B51E491799C1FC25F84C" box="[854,900,1945,1969]" pageId="4" pageNumber="19">type</typeStatus>
of wood is closely related to modern 
<taxonomicName id="4C0214F7CC00B51E491799E1FC46F82C" authorityName="Gray" authorityYear="1822" box="[854,999,1977,2001]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
due to the the presence of “ 
<emphasis id="B976B366CC00B51E4F6D99E1FAA4F7EE" italics="true" pageId="4" pageNumber="19">abietinean tracheid pitting, distinctly pitted ray cell walls, axial parenchyma and cupressoid cross-fields</emphasis>
”.
</paragraph>
<caption id="DF7D3FFCCC01B51F4AAB9958FB68F892" ID-DOI="http://doi.org/10.5281/zenodo.10520766" ID-Zenodo-Dep="10520766" httpUri="https://zenodo.org/record/10520766/files/figure.png" pageId="5" pageNumber="20" startId="5.[234,267,1792,1813]" targetBox="[251,1342,151,1750]" targetPageId="5" targetType="figure">
<paragraph id="8BBD6F74CC01B51F4AAB9958FB68F892" blockId="5.[234,1352,1791,1903]" pageId="5" pageNumber="20">
<emphasis id="B976B366CC01B51F4AAB9958FE82F8E8" bold="true" box="[234,291,1792,1813]" pageId="5" pageNumber="20">Fig. 3</emphasis>
The thin sections of 
<taxonomicName id="4C0214F7CC01B51F4BAC98A7FC88F8E8" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[493,809,1791,1813]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC01B51F4BAC98A7FD6BF8E8" box="[493,714,1791,1813]" italics="true" pageId="5" pageNumber="20">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC01B51F49719958FCDAF8E8" box="[816,891,1792,1813]" pageId="5" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
a) Transversal section with false rings and without resin canals, b) Marginal (zonate) axial parenchyma (arrows) and spaces between tracheids, c) Spaces between tracheids, and diffuse axial parenchyma cells, d) Tangential section with uni-to partly biseriate rays, e) Uniseriate pits on tangential walls of tracheids, f) Biseriate and alternate pits on tangential walls of tracheids.
</paragraph>
</caption>
<caption id="DF7D3FFCCC02B51C4A969965FC36F873" ID-DOI="http://doi.org/10.5281/zenodo.10520768" ID-Zenodo-Dep="10520768" httpUri="https://zenodo.org/record/10520768/files/figure.png" pageId="6" pageNumber="21" startId="6.[215,248,1853,1874]" targetBox="[230,1363,188,1812]" targetPageId="6" targetType="figure">
<paragraph id="8BBD6F74CC02B51C4A969965FC36F873" blockId="6.[215,1365,1853,1935]" pageId="6" pageNumber="21">
<emphasis id="B976B366CC02B51C4A969965FEB0F8AF" bold="true" box="[215,273,1853,1874]" pageId="6" pageNumber="21">Fig. 4</emphasis>
The thin sections of 
<taxonomicName id="4C0214F7CC02B51C4B9A9965FCB7F8AE" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[475,790,1853,1875]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC02B51C4B9A9965FD16F8AE" box="[475,695,1853,1875]" italics="true" pageId="6" pageNumber="21">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC02B51C495C9966FCC9F8AE" box="[797,872,1854,1875]" pageId="6" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
a) Tangential section with short rays, b-d) Nodular and smooth end walls of axial parenchyma, e) Mostly uniseriate and sometimes biseriate pits on radial walls of tracheids (arrows), f) Frequent uniseriate pits and crassulae formation (arrows).
</paragraph>
</caption>
<caption id="DF7D3FFCCC03B51D4AD699C3FBBBF811" ID-DOI="http://doi.org/10.5281/zenodo.10520770" ID-Zenodo-Dep="10520770" httpUri="https://zenodo.org/record/10520770/files/figure.png" pageId="7" pageNumber="22" startId="7.[151,184,1947,1968]" targetBox="[156,1365,168,1905]" targetPageId="7" targetType="figure">
<paragraph id="8BBD6F74CC03B51D4AD699C3FBBBF811" blockId="7.[151,1366,1946,2028]" pageId="7" pageNumber="22">
<emphasis id="B976B366CC03B51D4AD699C3FF73F84D" bold="true" box="[151,210,1947,1968]" pageId="7" pageNumber="22">Fig. 5</emphasis>
The thin sections of 
<taxonomicName id="4C0214F7CC03B51D4BE399C2FD41F84C" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[418,736,1946,1969]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="22" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC03B51D4BE399C2FDDEF84D" box="[418,639,1946,1968]" italics="true" pageId="7" pageNumber="22">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC03B51D48A699C3FC95F84D" box="[743,820,1947,1968]" pageId="7" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
a) Smooth end walls of rays (arrow), b-c) Slightly dentate end walls of ray, and sparsely pitted horizontal walls of rays, d) Indenture (vertical arrow) and sparsely nodular horizontal walls of rays (horizontal arrow), e-g) Cupressoid type cross-field pits, 2-5 pits per cross-field.
</paragraph>
</caption>
<caption id="DF7D3FFCCC0CB51248489FFAF9A9FE47" box="[521,1544,418,442]" pageId="8" pageNumber="23" startId="8.[521,586,418,442]" targetBox="[112,1970,497,1772]" targetIsTable="true" targetPageId="8" targetType="table">
<paragraph id="8BBD6F74CC0CB51248489FFAF9A9FE47" blockId="8.[521,1544,418,442]" box="[521,1544,418,442]" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB51248489FFAFDFCFE47" bold="true" box="[521,605,418,442]" pageId="8" pageNumber="23">Table 1</emphasis>
Comparison of the new species with the former fossil species (
<bibRefCitation id="EF931285CC0CB5124F469FFAF9A0FE47" author="Ruiz, D. P. &amp; Bodnar, J." box="[1287,1537,418,442]" pageId="8" pageNumber="23" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar, 2019</bibRefCitation>
)
</paragraph>
</caption>
<paragraph id="8BBD6F74CC0CB5124A319C42F82AF92C" pageId="8" pageNumber="23">
<table id="F9029DD4CC0C4AE54A319FA9F813F911" box="[112,1970,497,1772]" gridcols="13" gridrows="7" pageId="8" pageNumber="23">
<tr id="35326D36CC0C4AE54A319FA9F813FD48" box="[112,1970,497,693]" gridrow="0" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319FA9FEFAFD48" box="[112,347,497,693]" gridcol="0" gridrow="0" pageId="8" pageNumber="23">
Fossil species 
<taxonomicName id="4C0214F7CC0CB5124A319C20FF61FD5A" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319C20FE94FD71" box="[112,309,632,652]" italics="true" pageId="8" pageNumber="23">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC0CB5124A879CCBFEABFD5A" box="[198,266,659,679]" pageId="8" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="76E3044ACC0C4AE54B399FA9FE5DFD48" box="[376,508,497,693]" gridcol="1" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124B399C5AFE03FDE8" bold="true" box="[376,418,514,533]" pageId="8" pageNumber="23">Age;</emphasis>
<emphasis id="B976B366CC0CB5124B399C6AFE64FDB8" bold="true" box="[376,453,562,581]" pageId="8" pageNumber="23">Country</emphasis>
Early Miocene; Turkey
</th>
<th id="76E3044ACC0C4AE548529FA9FD50FD48" box="[531,753,497,693]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB51248559C55FD51FDDD" bold="true" box="[532,752,525,544]" pageId="8" pageNumber="23">Pits on radial tracheidal</emphasis>
<emphasis id="B976B366CC0CB512482A9C70FD38FDC6" bold="true" box="[619,665,552,571]" pageId="8" pageNumber="23">walls</emphasis>
Uniseriate, Frequent, sometimes contiguous biseriate
</th>
<th id="76E3044ACC0C4AE549529FA9FC2FFD48" box="[787,910,497,693]" gridcol="4" gridrow="0" pageId="8" pageNumber="23">Arrangement of multiseriate pits Opposite</th>
<th id="76E3044ACC0C4AE549F09FA9FB3EFD48" box="[945,1183,497,693]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB51249999FAEFBDEFDF4" bold="true" box="[984,1151,502,521]" pageId="8" pageNumber="23">Axial parenchyma</emphasis>
<emphasis id="B976B366CC0CB51249899C6EFB39FDB4" bold="true" box="[968,1176,566,585]" pageId="8" pageNumber="23">
<typeStatus id="54B9D1D6CC0CB51249899C6EFC57FDB4" box="[968,1014,566,585]" pageId="8" pageNumber="23">Type</typeStatus>
End walls
</emphasis>
Diffuse, Smooth and and zonate nodular
</th>
<th id="76E3044ACC0C4AE54E8B9FA9FAA2FD48" box="[1226,1283,497,693]" gridcol="7" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124E959C55FB58FDDD" bold="true" box="[1236,1273,525,544]" pageId="8" pageNumber="23">Ray</emphasis>
height 1-16
</th>
<th id="76E3044ACC0C4AE54F669FA9FA32FD48" box="[1319,1427,497,693]" gridcol="8" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124F6F9C42FA2DFDD0" bold="true" box="[1326,1420,538,557]" pageId="8" pageNumber="23">Ray width</emphasis>
Uniseriate, rarely partly biseriate
</th>
<th id="76E3044ACC0C4AE54F819FA9F9AFFD48" box="[1472,1550,497,693]" gridcol="9" gridrow="0" pageId="8" pageNumber="23">Ray tracheids -</th>
<th id="76E3044ACC0C4AE54C7B9FA9F906FD48" box="[1594,1703,497,693]" gridcol="10" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124C7F9FA7F922FDB5" bold="true" pageId="8" pageNumber="23">Horizontal walls of rays</emphasis>
Smooth or pitted
</th>
<th id="76E3044ACC0C4AE54C8C9FA9F892FD48" box="[1741,1843,497,693]" gridcol="11" gridrow="0" pageId="8" pageNumber="23">Number and type of cross-field pits 2-3 (5), cupressoid</th>
<th id="76E3044ACC0C4AE54D199FA9F813FD48" box="[1880,1970,497,693]" gridcol="12" gridrow="0" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124D199C42F813FDD0" bold="true" box="[1880,1970,538,557]" pageId="8" pageNumber="23">Crassulae</emphasis>
±
</th>
</tr>
<tr id="35326D36CC0C4AE54A319C92F813FCE8" box="[112,1970,714,789]" gridrow="1" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319C92FEFAFCE8" box="[112,347,714,789]" gridcol="0" gridrow="1" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319C92FF48FCE8" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="23" phylum="Tracheophyta" rank="genus">Juniperoxylon breviparenchymatosum Watari &amp; Nishida, 1973</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399C92FE5DFCE8" box="[376,508,714,789]" gridcol="1" gridrow="1" pageId="8" pageNumber="23">Eocene; Japan</td>
<td id="76E3044ACC0C4AE548529C92FDD0FCE8" box="[531,625,714,789]" gridcol="2" gridrow="1" pageId="8" pageNumber="23">Uniseriate, biseriate</td>
<td id="76E3044ACC0C4AE548D29C92FD50FCE8" box="[659,753,714,789]" gridcol="3" gridrow="1" pageId="8" pageNumber="23">Spaced</td>
<td id="76E3044ACC0C4AE549529C92FC2FFCE8" box="[787,910,714,789]" gridcol="4" gridrow="1" pageId="8" pageNumber="23">Opposite</td>
<td id="76E3044ACC0C4AE549F09C92FBACFCE8" box="[945,1037,714,789]" gridcol="5" gridrow="1" pageId="8" pageNumber="23">Abundant, diffuse, and zonate</td>
<td id="76E3044ACC0C4AE54E7F9C92FB3EFCE8" box="[1086,1183,714,789]" gridcol="6" gridrow="1" pageId="8" pageNumber="23">Smooth or irregularly thickened</td>
<td id="76E3044ACC0C4AE54E8B9C92FAA2FCE8" box="[1226,1283,714,789]" gridcol="7" gridrow="1" pageId="8" pageNumber="23">1-14</td>
<td id="76E3044ACC0C4AE54F669C92FA32FCE8" box="[1319,1427,714,789]" gridcol="8" gridrow="1" pageId="8" pageNumber="23">Uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0C4AE54F819C92F9AFFCE8" box="[1472,1550,714,789]" gridcol="9" gridrow="1" pageId="8" pageNumber="23">-</td>
<td id="76E3044ACC0C4AE54C7B9C92F906FCE8" box="[1594,1703,714,789]" gridcol="10" gridrow="1" pageId="8" pageNumber="23">Smooth or rarely pitted</td>
<td id="76E3044ACC0C4AE54C8C9C92F892FCE8" box="[1741,1843,714,789]" gridcol="11" gridrow="1" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124C8F9C80F893FD11" box="[1742,1842,728,748]" italics="true" pageId="8" pageNumber="23">2-6, taxodioid</emphasis>
</td>
<td id="76E3044ACC0C4AE54D199C92F813FCE8" box="[1880,1970,714,789]" gridcol="12" gridrow="1" pageId="8" pageNumber="23">-</td>
</tr>
<tr id="35326D36CC0C4AE54A319D69F813FC81" box="[112,1970,817,892]" gridrow="2" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319D69FEFAFC81" box="[112,347,817,892]" gridcol="0" gridrow="2" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319D69FF3AFC81" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319D69FEFAFCB8" box="[112,347,817,837]" italics="true" pageId="8" pageNumber="23">Juniperoxylon juniperoides</emphasis>
(Kownas, 1951) Huard, 1966
</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399D69FE5DFC81" box="[376,508,817,892]" gridcol="1" gridrow="2" pageId="8" pageNumber="23">Miocene; Germany</td>
<td id="76E3044ACC0C4AE548529D69FDD0FC81" box="[531,625,817,892]" gridcol="2" gridrow="2" pageId="8" pageNumber="23">Uniseriate, biseriate</td>
<td id="76E3044ACC0C4AE548D29D69FD50FC81" box="[659,753,817,892]" gridcol="3" gridrow="2" pageId="8" pageNumber="23">Spaced, contiguous</td>
<td id="76E3044ACC0C4AE549529D69FC2FFC81" box="[787,910,817,892]" gridcol="4" gridrow="2" pageId="8" pageNumber="23">Opposite</td>
<td id="76E3044ACC0C4AE549F09D69FBACFC81" box="[945,1037,817,892]" gridcol="5" gridrow="2" pageId="8" pageNumber="23">Abundant, diffuse, and zonate</td>
<td id="76E3044ACC0C4AE54E7F9D69FB3EFC81" box="[1086,1183,817,892]" gridcol="6" gridrow="2" pageId="8" pageNumber="23">Nodular</td>
<td id="76E3044ACC0C4AE54E8B9D69FAA2FC81" box="[1226,1283,817,892]" gridcol="7" gridrow="2" pageId="8" pageNumber="23">1-12</td>
<td id="76E3044ACC0C4AE54F669D69FA32FC81" box="[1319,1427,817,892]" gridcol="8" gridrow="2" pageId="8" pageNumber="23">Uniseriate</td>
<td id="76E3044ACC0C4AE54F819D69F9AFFC81" box="[1472,1550,817,892]" gridcol="9" gridrow="2" pageId="8" pageNumber="23">-</td>
<td id="76E3044ACC0C4AE54C7B9D69F906FC81" box="[1594,1703,817,892]" gridcol="10" gridrow="2" pageId="8" pageNumber="23">Distinctly pitted</td>
<td id="76E3044ACC0C4AE54C8C9D69F892FC81" box="[1741,1843,817,892]" gridcol="11" gridrow="2" pageId="8" pageNumber="23">1-4 (6), cupressoid</td>
<td id="76E3044ACC0C4AE54D199D69F813FC81" box="[1880,1970,817,892]" gridcol="12" gridrow="2" pageId="8" pageNumber="23">+</td>
</tr>
<tr id="35326D36CC0C4AE54A319DC2F813FB97" box="[112,1970,922,1130]" gridrow="3" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319DC2FEFAFB97" box="[112,347,922,1130]" gridcol="0" gridrow="3" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319DF0FF3AFC0E" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319DF0FEFBFC41" box="[112,346,936,956]" italics="true" pageId="8" pageNumber="23">Juniperoxylon pachyderma</emphasis>
(Göppert, 1850) Krausel, 1949
</taxonomicName>
<taxonomicName id="4C0214F7CC0CB5124A319A47FF3AFB97" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319A47FEF3FBCE" box="[112,338,1055,1075]" italics="true" pageId="8" pageNumber="23">Juniperoxylon pottoniense</emphasis>
(
<bibRefCitation id="EF931285CC0CB5124A369A62FF46FBB3" author="Stopes, M. C." box="[119,231,1082,1102]" pageId="8" pageNumber="23" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) Krausel, 1949
</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399DC2FE5DFB97" box="[376,508,922,1130]" gridcol="1" gridrow="3" pageId="8" pageNumber="23">Miocene; Germany, Belgium, Netherlands, Poland, Siberia Cretaceous-Eocene; England, Denmark</td>
<td id="76E3044ACC0C4AE548529DC2FDD0FB97" box="[531,625,922,1130]" gridcol="2" gridrow="3" pageId="8" pageNumber="23">Uniseriate, rarely biseriate, triseriate Uniseriate</td>
<td id="76E3044ACC0C4AE548D29DC2FD50FB97" box="[659,753,922,1130]" gridcol="3" gridrow="3" pageId="8" pageNumber="23">Spaced Spaced</td>
<td id="76E3044ACC0C4AE549529DC2FC2FFB97" box="[787,910,922,1130]" gridcol="4" gridrow="3" pageId="8" pageNumber="23">Opposite -</td>
<td id="76E3044ACC0C4AE549F09DC2FBACFB97" box="[945,1037,922,1130]" gridcol="5" gridrow="3" pageId="8" pageNumber="23">Abundant, diffuse, and zonate Abundant, zonate</td>
<td id="76E3044ACC0C4AE54E7F9DC2FB3EFB97" box="[1086,1183,922,1130]" gridcol="6" gridrow="3" pageId="8" pageNumber="23">Smooth or nodular Smooth?</td>
<td id="76E3044ACC0C4AE54E8B9DC2FAA2FB97" box="[1226,1283,922,1130]" gridcol="7" gridrow="3" pageId="8" pageNumber="23">1-15 1-12</td>
<td id="76E3044ACC0C4AE54F669DC2FA32FB97" box="[1319,1427,922,1130]" gridcol="8" gridrow="3" pageId="8" pageNumber="23">Uniseriate, rarely partly biseriate Uniseriate</td>
<td id="76E3044ACC0C4AE54F819DC2F9AFFB97" box="[1472,1550,922,1130]" gridcol="9" gridrow="3" pageId="8" pageNumber="23">- -</td>
<td id="76E3044ACC0C4AE54C7B9DC2F906FB97" box="[1594,1703,922,1130]" gridcol="10" gridrow="3" pageId="8" pageNumber="23">Smooth Distinctly pitted</td>
<td id="76E3044ACC0C4AE54C8C9DC2F892FB97" box="[1741,1843,922,1130]" gridcol="11" gridrow="3" pageId="8" pageNumber="23">1-6, cupressoid Cupressoid</td>
<td id="76E3044ACC0C4AE54D199DC2F813FB97" box="[1880,1970,922,1130]" gridcol="12" gridrow="3" pageId="8" pageNumber="23">+ -</td>
</tr>
<tr id="35326D36CC0C4AE54A319ADEF813FB2C" box="[112,1970,1158,1233]" gridrow="4" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319ADEFEFAFB2C" box="[112,347,1158,1233]" gridcol="0" gridrow="4" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319ACCFE81FB3E" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319ACCFEEBFB55" box="[112,330,1172,1192]" italics="true" pageId="8" pageNumber="23">Juniperoxylon rhenanurn</emphasis>
<bibRefCitation id="EF931285CC0CB5124A319AF7FE81FB3E" author="van der Burgh, J." box="[112,288,1199,1219]" pageId="8" pageNumber="23" pagination="73 - 275" refId="ref6680" refString="van der Burgh, J., 1973. Holzer der niederrheinischen braunkohlenformation, 2. Holzer der braunkohlengruben &quot; Maria Theresia &quot; zu herzogenrath, &quot; Zukunft West &quot; zu eschweiler und &quot; Victor &quot; (Zulpich mitte) zu Zulpich. Nebst einer systematisch-anatomischen bearbeitung der gattung Pinus L. Review of Palaeobotany and Palynology, 15 / 2: 73 - 275." type="journal article" year="1973">van der Burgh, 1973</bibRefCitation>
</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399ADEFE5DFB2C" box="[376,508,1158,1233]" gridcol="1" gridrow="4" pageId="8" pageNumber="23">Miocene; Germany</td>
<td id="76E3044ACC0C4AE548529ADEFDD0FB2C" box="[531,625,1158,1233]" gridcol="2" gridrow="4" pageId="8" pageNumber="23">Uniseriate, biseriate</td>
<td id="76E3044ACC0C4AE548D29ADEFD50FB2C" box="[659,753,1158,1233]" gridcol="3" gridrow="4" pageId="8" pageNumber="23">Spaced</td>
<td id="76E3044ACC0C4AE549529ADEFC2FFB2C" box="[787,910,1158,1233]" gridcol="4" gridrow="4" pageId="8" pageNumber="23">Opposite</td>
<td id="76E3044ACC0C4AE549F09ADEFBACFB2C" box="[945,1037,1158,1233]" gridcol="5" gridrow="4" pageId="8" pageNumber="23">Present, diffuse, and zonate</td>
<td id="76E3044ACC0C4AE54E7F9ADEFB3EFB2C" box="[1086,1183,1158,1233]" gridcol="6" gridrow="4" pageId="8" pageNumber="23">Nodular</td>
<td id="76E3044ACC0C4AE54E8B9ADEFAA2FB2C" box="[1226,1283,1158,1233]" gridcol="7" gridrow="4" pageId="8" pageNumber="23">1-40</td>
<td id="76E3044ACC0C4AE54F669ADEFA32FB2C" box="[1319,1427,1158,1233]" gridcol="8" gridrow="4" pageId="8" pageNumber="23">Uniseriate, occasionally biseriate</td>
<td id="76E3044ACC0C4AE54F819ADEF9AFFB2C" box="[1472,1550,1158,1233]" gridcol="9" gridrow="4" pageId="8" pageNumber="23">±</td>
<td id="76E3044ACC0C4AE54C7B9ADEF906FB2C" box="[1594,1703,1158,1233]" gridcol="10" gridrow="4" pageId="8" pageNumber="23">Distinctly pitted</td>
<td id="76E3044ACC0C4AE54C8C9ADEF892FB2C" box="[1741,1843,1158,1233]" gridcol="11" gridrow="4" pageId="8" pageNumber="23">1-3, cupressoid, taxodioid</td>
<td id="76E3044ACC0C4AE54D199ADEF813FB2C" box="[1880,1970,1158,1233]" gridcol="12" gridrow="4" pageId="8" pageNumber="23">+</td>
</tr>
<tr id="35326D36CC0C4AE54A319AB5F813FAC5" box="[112,1970,1261,1336]" gridrow="5" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319AB5FEFAFAC5" box="[112,347,1261,1336]" gridcol="0" gridrow="5" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319AA3FE93FAD7" authorityName="Krausel" authorityYear="1949" baseAuthorityName="Houlbert" baseAuthorityYear="1910" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="23" phylum="Tracheophyta" rank="genus">
Juniperoxylon schneiderianum 
<bibRefCitation id="EF931285CC0CB5124AEA9B4EFE93FAD7" author="Dolezych, M." box="[171,306,1302,1322]" pageId="8" pageNumber="23" pagination="5 - 31" refId="ref5819" refString="Dolezych, M., 2016. A remarkable extinct wood from Lusatia (central Europe) - Juniperoxylon schneiderianum sp. nov. with affinity to Cupressospermum saxonicum Mai. Palaeontographica B, 295: 5 - 31." type="journal article" year="2016">Dolezych, 2016</bibRefCitation>
</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399AB5FE5DFAC5" box="[376,508,1261,1336]" gridcol="1" gridrow="5" pageId="8" pageNumber="23">Miocene; Germany</td>
<td id="76E3044ACC0C4AE548529AB5FDD0FAC5" box="[531,625,1261,1336]" gridcol="2" gridrow="5" pageId="8" pageNumber="23">Uniseriate, biseriate</td>
<td id="76E3044ACC0C4AE548D29AB5FD50FAC5" box="[659,753,1261,1336]" gridcol="3" gridrow="5" pageId="8" pageNumber="23">Spaced</td>
<td id="76E3044ACC0C4AE549529AB5FC2FFAC5" box="[787,910,1261,1336]" gridcol="4" gridrow="5" pageId="8" pageNumber="23">Opposite</td>
<td id="76E3044ACC0C4AE549F09AB5FBACFAC5" box="[945,1037,1261,1336]" gridcol="5" gridrow="5" pageId="8" pageNumber="23">Abundant, diffuse</td>
<td id="76E3044ACC0C4AE54E7F9AB5FB3EFAC5" box="[1086,1183,1261,1336]" gridcol="6" gridrow="5" pageId="8" pageNumber="23">Nodular</td>
<td id="76E3044ACC0C4AE54E8B9AB5FAA2FAC5" box="[1226,1283,1261,1336]" gridcol="7" gridrow="5" pageId="8" pageNumber="23">2-18</td>
<td id="76E3044ACC0C4AE54F669AB5FA32FAC5" box="[1319,1427,1261,1336]" gridcol="8" gridrow="5" pageId="8" pageNumber="23">Uniseriate, occasionally biseriate</td>
<td id="76E3044ACC0C4AE54F819AB5F9AFFAC5" box="[1472,1550,1261,1336]" gridcol="9" gridrow="5" pageId="8" pageNumber="23">-</td>
<td id="76E3044ACC0C4AE54C7B9AB5F906FAC5" box="[1594,1703,1261,1336]" gridcol="10" gridrow="5" pageId="8" pageNumber="23">Sparsely pitted</td>
<td id="76E3044ACC0C4AE54C8C9AB5F892FAC5" box="[1741,1843,1261,1336]" gridcol="11" gridrow="5" pageId="8" pageNumber="23">2-4, cupressoid, taxodioid</td>
<td id="76E3044ACC0C4AE54D199AB5F813FAC5" box="[1880,1970,1261,1336]" gridcol="12" gridrow="5" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124D3E9B51F82AFAE0" box="[1919,1931,1289,1309]" italics="true" pageId="8" pageNumber="23">+</emphasis>
</td>
</tr>
<tr id="35326D36CC0C4AE54A319B0CF813F911" box="[112,1970,1364,1772]" gridrow="6" pageId="8" pageNumber="23">
<th id="76E3044ACC0C4AE54A319B0CFEFAF911" box="[112,347,1364,1772]" gridcol="0" gridrow="6" pageId="8" pageNumber="23">
<taxonomicName id="4C0214F7CC0CB5124A319B3AFE80FA6C" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319B3AFE97FA8B" box="[112,310,1378,1398]" italics="true" pageId="8" pageNumber="23">Juniperoxylon wagneri</emphasis>
Süss &amp; Rather, 1998
</taxonomicName>
<taxonomicName id="4C0214F7CC0CB5124A319BEEFE88F9E6" pageId="8" pageNumber="23">
<emphasis id="B976B366CC0CB5124A319BEEFEEFFA37" box="[112,334,1462,1482]" italics="true" pageId="8" pageNumber="23">Juniperoxylon zamunerae</emphasis>
(Bodnar, Ruiz, Artabe, Morel,&amp; Ganuza, 2015) 
<bibRefCitation id="EF931285CC0CB5124A31985FFE88F9E6" author="Ruiz, D. P. &amp; Bodnar, J." box="[112,297,1543,1563]" pageId="8" pageNumber="23" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz &amp; Bodnar, 2019</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C0214F7CC0CB5124A319875FF3AF96F" authorityName="W.Eckhold" authorityYear="1921" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="23" phylum="Tracheophyta" rank="genus">
Protojuniperoxylon ischigualastense 
<bibRefCitation id="EF931285CC0CB5124A9A9810FEEEF9A1" author="Bonetti, M. I. R." box="[219,335,1608,1628]" pageId="8" pageNumber="23" pagination="211 - 218" refId="ref5786" refString="Bonetti, M. I. R., 1966. Protojuniperoxylon ischigualastensis sp. nov. Del Triasico de Ischigualasto (San Juan). Ameghiniana 4: 211 - 218." type="journal article" year="1966">Bonetti, 1966</bibRefCitation>
emend.Bodnar &amp; Artabe, 2007
</taxonomicName>
<taxonomicName id="4C0214F7CC0CB5124A3198FCFF4CF911" authorityName="W.Eckhold" authorityYear="1921" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="23" phylum="Tracheophyta" rank="genus">
Protojuniperoxylon maidstonense (
<bibRefCitation id="EF931285CC0CB5124A8798E5FE97F92C" author="Stopes, M. C." box="[198,310,1725,1745]" pageId="8" pageNumber="23" refId="ref6455" refString="Stopes, M. C., 1915. Catalogue of the Mesozoic Plants in the British Museum (Natural History). Part II: Lower Greensand (Aptian) Plants of Britain. xxxvi + 360 pp. Trustees of British Museum, London." type="book" year="1915">Stopes, 1915</bibRefCitation>
) 
<bibRefCitation id="EF931285CC0CB5124A319880FF4CF911" author="Eckhold, W." box="[112,237,1752,1772]" pageId="8" pageNumber="23" pagination="472 - 505" refId="ref5859" refString="Eckhold, W., 1923. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch der preussischen geologischen Landesanstalt zu Berlin, 42: 472 - 505." type="journal article" year="1923">Eckhold, 1923</bibRefCitation>
</taxonomicName>
</th>
<td id="76E3044ACC0C4AE54B399B0CFE5DF911" box="[376,508,1364,1772]" gridcol="1" gridrow="6" pageId="8" pageNumber="23">Miocene; Germany Middle Triassic; Argentina Late Triassic; Argentina Lower Cretaceous; England</td>
<td id="76E3044ACC0C4AE548529B0CFDD0F911" box="[531,625,1364,1772]" gridcol="2" gridrow="6" pageId="8" pageNumber="23">Uniseriate, rarely biseriate Uniseriate, rarely biseriate Uniseriate, biseriate Uniseriate</td>
<td id="76E3044ACC0C4AE548D29B0CFD50F911" box="[659,753,1364,1772]" gridcol="3" gridrow="6" pageId="8" pageNumber="23">Spaced Spaced Spaced, contiguous Spaced, contiguous</td>
<td id="76E3044ACC0C4AE549529B0CFC2FF911" box="[787,910,1364,1772]" gridcol="4" gridrow="6" pageId="8" pageNumber="23">Opposite Opposite Alternate, opposite?</td>
<td id="76E3044ACC0C4AE549F09B0CFBACF911" box="[945,1037,1364,1772]" gridcol="5" gridrow="6" pageId="8" pageNumber="23">Abundant, diffuse Scarce, diffuse Present, diffuse?</td>
<td id="76E3044ACC0C4AE54E7F9B0CFB3EF911" box="[1086,1183,1364,1772]" gridcol="6" gridrow="6" pageId="8" pageNumber="23">Nodular Smooth or irregularly thickened Smooth?</td>
<td id="76E3044ACC0C4AE54E8B9B0CFAA2F911" box="[1226,1283,1364,1772]" gridcol="7" gridrow="6" pageId="8" pageNumber="23">1-5 2-34 1-45 1-20</td>
<td id="76E3044ACC0C4AE54F669B0CFA32F911" box="[1319,1427,1364,1772]" gridcol="8" gridrow="6" pageId="8" pageNumber="23">
Uniseriate Uniseriate, rarely partly biseriate 
<emphasis id="B976B366CC0CB5124F6F9875FA2DF9BC" box="[1326,1420,1581,1601]" italics="true" pageId="8" pageNumber="23">Uniseriate,</emphasis>
less frequent partly bi- or triseriate Uniseriate, rarely partly biseriate
</td>
<td id="76E3044ACC0C4AE54F819B0CF9AFF911" box="[1472,1550,1364,1772]" gridcol="9" gridrow="6" pageId="8" pageNumber="23">- - + -</td>
<td id="76E3044ACC0C4AE54C7B9B0CF906F911" box="[1594,1703,1364,1772]" gridcol="10" gridrow="6" pageId="8" pageNumber="23">Smooth Distinctly pitted Distinctly pitted Smooth?</td>
<td id="76E3044ACC0C4AE54C8C9B0CF892F911" box="[1741,1843,1364,1772]" gridcol="11" gridrow="6" pageId="8" pageNumber="23">
2-8, cupressoid 1-2(3-4), cupressoid 2-4, cupressoid 
<emphasis id="B976B366CC0CB5124C9F98E8F880F939" box="[1758,1825,1712,1732]" italics="true" pageId="8" pageNumber="23">4-6, cupressoid</emphasis>
</td>
<td id="76E3044ACC0C4AE54D199B0CF813F911" box="[1880,1970,1364,1772]" gridcol="12" gridrow="6" pageId="8" pageNumber="23">- - - +</td>
</tr>
</table>
</paragraph>
<caption id="DF7D3FFCCC0DB5134B5C9E84FC13FEF2" pageId="9" pageNumber="24" startId="9.[285,344,220,241]" targetBox="[155,1432,311,1610]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="8BBD6F74CC0DB5134B5C9E84FC13FEF2" blockId="9.[285,1303,220,271]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134B5C9E84FED1FF0C" bold="true" box="[285,368,220,241]" pageId="9" pageNumber="24">Table 2.</emphasis>
Comparison of modern 
<taxonomicName id="4C0214F7CC0DB513481A9E85FCA4FF0F" box="[603,773,220,242]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB513481A9E85FD1BFF0F" box="[603,698,221,242]" italics="true" pageId="9" pageNumber="24">Juniperus</emphasis>
species
</taxonomicName>
(Esteban et al., 2004; 
<bibRefCitation id="EF931285CC0DB513499C9E84FB49FF0F" author="Akkemik, U. &amp; Yaman, B." box="[989,1256,220,242]" pageId="9" pageNumber="24" refId="ref5315" refString="Akkemik, U. &amp; Yaman, B., 2012. Wood Anatomy of Eastern Mediterranean Species. Kessel Publishing House, Germany." type="book" year="2012">Akkemik and Yaman, 2012</bibRefCitation>
) and 
<taxonomicName id="4C0214F7CC0DB51348C29EA1FCFEFEF2" authority="Akkemik, 2021" authorityName="Akkemik" authorityYear="2021" box="[643,863,249,271]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC0DB51348C29EA1FCFEFEF2" box="[643,863,249,271]" italics="true" pageId="9" pageNumber="24">Juniperoxylon acarcae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC0DB51349279EA2FC13FEF2" box="[870,946,250,271]" pageId="9" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBD6F74CC0DB5134ADA9F6FFA97F9B7" pageId="9" pageNumber="24">
<table id="F9029DD4CC0D4AE54ADA9F6FFA39F9B7" box="[155,1432,311,1610]" gridcols="9" gridrows="14" pageId="9" pageNumber="24">
<tr id="35326D36CC0D4AE54ADA9F6FFA39FE50" box="[155,1432,311,429]" gridrow="0" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9F6FFEB7FE50" box="[155,278,311,429]" gridcol="0" gridrow="0" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9F6FFF60FE50" bold="true" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9F6FFF48FE86" bold="true" italics="true" pageId="9" pageNumber="24">Fossil species / Features</emphasis>
Growth ring
</emphasis>
</th>
<th id="76E3044ACC0D4AE54B6B9F6FFE09FE50" box="[298,424,311,429]" gridcol="1" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB5134B6B9F6FFEC7FE86" authority="Labill. Distinct" authorityName="Labill. Distinct" box="[298,387,311,379]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="drupacea">
<emphasis id="B976B366CC0DB5134B6B9F6FFEC7FE86" bold="true" box="[298,387,311,379]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134B6B9F6FFEDDFE9F" bold="true" box="[298,387,311,354]" italics="true" pageId="9" pageNumber="24">Juniperus drupacea</emphasis>
Labill.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE54BF79F6FFD87FE50" box="[438,550,311,429]" gridcol="2" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB5134BF79F6FFE5FFE86" authority="M. Bieb. Distinct" authorityName="M. Bieb. Distinct" box="[438,527,311,379]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B976B366CC0DB5134BF79F6FFE5FFE86" bold="true" box="[438,527,311,379]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134BF79F6FFE57FE9F" bold="true" box="[438,527,311,354]" italics="true" pageId="9" pageNumber="24">Juniperus excelsa</emphasis>
M.Bieb.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE5487F9F6FFD12FE50" box="[574,691,311,429]" gridcol="3" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB513487F9F6FFDD6FE86" authority="Willd. Distinct" authorityName="Willd. Distinct" box="[574,680,311,379]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="foetidissima">
<emphasis id="B976B366CC0DB513487F9F6FFDD6FE86" bold="true" box="[574,680,311,379]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB513487F9F6FFD09FE9F" bold="true" box="[574,680,311,354]" italics="true" pageId="9" pageNumber="24">Juniperus foetidissima</emphasis>
Willd.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE548829F6FFC94FE50" box="[707,821,311,429]" gridcol="4" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB51348829F1BFC94FE93" authority="L. Distinct" authorityName="L. Distinct" box="[707,821,323,366]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="oxycedrus">
<emphasis id="B976B366CC0DB51348829F1BFC94FE93" bold="true" box="[707,821,323,366]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB51348829F1BFCBDFE93" bold="true" box="[707,796,323,366]" italics="true" pageId="9" pageNumber="24">Juniperus oxycedrus</emphasis>
L.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE549089F6FFC1AFE50" box="[841,955,311,429]" gridcol="5" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB51349089F1BFC1AFE93" authority="L. Distinct" authorityName="L. Distinct" box="[841,955,323,366]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="phoenicea">
<emphasis id="B976B366CC0DB51349089F1BFC1AFE93" bold="true" box="[841,955,323,366]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB51349089F1BFC02FE93" bold="true" box="[841,931,323,366]" italics="true" pageId="9" pageNumber="24">Juniperus phoenicea</emphasis>
L.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE5498F9F6FFBEEFE50" box="[974,1103,311,429]" gridcol="6" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB513498F9F6FFBEEFE86" baseAuthorityName="Ruiz and Bodnar" baseAuthorityYear="2019" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB513498F9F6FFBEEFE86" bold="true" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB513498F9F6FFBA5FE9F" bold="true" italics="true" pageId="9" pageNumber="24">Juniperus saltuaria</emphasis>
Rehder &amp; E.H.Wilson
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE54E249F6FFB4BFE50" box="[1125,1258,311,429]" gridcol="7" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB5134E249F1BFB6CFE93" authority="L. Distinct" authorityName="L. Distinct" box="[1125,1229,323,366]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="thurifera">
<emphasis id="B976B366CC0DB5134E249F1BFB6CFE93" bold="true" box="[1125,1229,323,366]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134E249F1BFB15FE93" bold="true" box="[1125,1214,323,366]" italics="true" pageId="9" pageNumber="24">Juniperus thurifera</emphasis>
L.
</emphasis>
</taxonomicName>
Distinct
</th>
<th id="76E3044ACC0D4AE54EB69F6FFA39FE50" box="[1271,1432,311,429]" gridcol="8" gridrow="0" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134EB69F6FFA9CFE86" bold="true" pageId="9" pageNumber="24">
<taxonomicName id="4C0214F7CC0DB5134EB69F6FFA37FE9F" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" box="[1271,1430,311,354]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC0DB5134EB69F6FFA9DFE9F" bold="true" box="[1271,1399,311,354]" italics="true" pageId="9" pageNumber="24">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC0DB5134EB69F30FA9CFE86" box="[1271,1341,360,379]" pageId="9" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Distinct
</th>
</tr>
<tr id="35326D36CC0D4AE54ADA9FE0FA39FE19" box="[155,1432,440,484]" gridrow="1" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9FE0FEB7FE19" box="[155,278,440,484]" gridcol="0" gridrow="1" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9FE0FF51FE19" bold="true" pageId="9" pageNumber="24">Partial and false ring</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9FE0FE09FE19" box="[298,424,440,484]" gridcol="1" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE54BF79FE0FD87FE19" box="[438,550,440,484]" gridcol="2" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE5487F9FE0FD12FE19" box="[574,691,440,484]" gridcol="3" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE548829FE0FC94FE19" box="[707,821,440,484]" gridcol="4" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE549089FE0FC1AFE19" box="[841,955,440,484]" gridcol="5" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE5498F9FE0FBEEFE19" box="[974,1103,440,484]" gridcol="6" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE54E249FE0FB4BFE19" box="[1125,1258,440,484]" gridcol="7" gridrow="1" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE54EB69FE0FA39FE19" box="[1271,1432,440,484]" gridcol="8" gridrow="1" pageId="9" pageNumber="24">Present</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9FB7FA39FD91" box="[155,1432,495,620]" gridrow="2" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9FB7FEB7FD91" box="[155,278,495,620]" gridcol="0" gridrow="2" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9C52FF4CFD9A" bold="true" pageId="9" pageNumber="24">Transition from earlywood to latewood</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9FB7FE09FD91" box="[298,424,495,620]" gridcol="1" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE54BF79FB7FD87FD91" box="[438,550,495,620]" gridcol="2" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE5487F9FB7FD12FD91" box="[574,691,495,620]" gridcol="3" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE548829FB7FC94FD91" box="[707,821,495,620]" gridcol="4" gridrow="2" pageId="9" pageNumber="24">Gradual; 2-3 seriate of flattened latewood tracheids</td>
<td id="76E3044ACC0D4AE549089FB7FC1AFD91" box="[841,955,495,620]" gridcol="5" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE5498F9FB7FBEEFD91" box="[974,1103,495,620]" gridcol="6" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE54E249FB7FB4BFD91" box="[1125,1258,495,620]" gridcol="7" gridrow="2" pageId="9" pageNumber="24">Gradual</td>
<td id="76E3044ACC0D4AE54EB69FB7FA39FD91" box="[1271,1432,495,620]" gridcol="8" gridrow="2" pageId="9" pageNumber="24">Gradual; 1-2 seriate of flattened latewood tracheids</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9CD2FA39FCC3" box="[155,1432,650,830]" gridrow="3" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9CD2FEB7FCC3" box="[155,278,650,830]" gridcol="0" gridrow="3" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9CF7FF6CFCC3" bold="true" box="[155,270,687,830]" pageId="9" pageNumber="24">Radial pitting Intercellular space</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9CD2FE09FCC3" box="[298,424,650,830]" gridcol="1" gridrow="3" pageId="9" pageNumber="24">Predominantly uniseriate Commonly present</td>
<td id="76E3044ACC0D4AE54BF79CD2FBEEFCC3" box="[438,1103,650,830]" colspan="5" colspanRight="4" gridcol="2" gridrow="3" pageId="9" pageNumber="24">Predominantly Predominantly Predominantly Predominantly Predominantly uniseriate uniseriate uniseriate uniseriate uniseriate Commonly Commonly Commonly Commonly present present present Present present</td>
<td id="76E3044ACC0D4AE54E249CD2FB4BFCC3" box="[1125,1258,650,830]" gridcol="7" gridrow="3" pageId="9" pageNumber="24">Predominantly uniseriate Present</td>
<td id="76E3044ACC0D4AE54EB69CD2FA39FCC3" box="[1271,1432,650,830]" gridcol="8" gridrow="3" pageId="9" pageNumber="24">Uniseriate, Sometimes biseriate, opposite, spaced and contiguous Commonly present</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9D0CFA39FC82" box="[155,1432,852,895]" gridrow="4" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9D0CFEB7FC82" box="[155,278,852,895]" gridcol="0" gridrow="4" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9D0CFF6CFC82" bold="true" pageId="9" pageNumber="24">Helical thickening</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9D0CFE09FC82" box="[298,424,852,895]" gridcol="1" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54BF79D0CFD87FC82" box="[438,550,852,895]" gridcol="2" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE5487F9D0CFD12FC82" box="[574,691,852,895]" gridcol="3" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE548829D0CFC94FC82" box="[707,821,852,895]" gridcol="4" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE549089D0CFC1AFC82" box="[841,955,852,895]" gridcol="5" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE5498F9D0CFBEEFC82" box="[974,1103,852,895]" gridcol="6" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54E249D0CFB4BFC82" box="[1125,1258,852,895]" gridcol="7" gridrow="4" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54EB69D0CFA39FC82" box="[1271,1432,852,895]" gridcol="8" gridrow="4" pageId="9" pageNumber="24">Absent</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9DD5FA39FC45" box="[155,1432,909,952]" gridrow="5" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9DD5FEB7FC45" box="[155,278,909,952]" gridcol="0" gridrow="5" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9DD5FF4EFC45" bold="true" box="[155,245,909,952]" pageId="9" pageNumber="24">Latewood tracheids</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9DD5FE09FC45" box="[298,424,909,952]" gridcol="1" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE54BF79DD5FD87FC45" box="[438,550,909,952]" gridcol="2" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE5487F9DD5FD12FC45" box="[574,691,909,952]" gridcol="3" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE548829DD5FC94FC45" box="[707,821,909,952]" gridcol="4" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE549089DD5FC1AFC45" box="[841,955,909,952]" gridcol="5" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE5498F9DD5FBEEFC45" box="[974,1103,909,952]" gridcol="6" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE54E249DD5FB4BFC45" box="[1125,1258,909,952]" gridcol="7" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
<td id="76E3044ACC0D4AE54EB69DD5FA39FC45" box="[1271,1432,909,952]" gridcol="8" gridrow="5" pageId="9" pageNumber="24">Thick walled</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9D9EFA39FC0C" box="[155,1432,966,1009]" gridrow="6" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9D9EFEB7FC0C" box="[155,278,966,1009]" gridcol="0" gridrow="6" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9D9EFF79FC0C" bold="true" pageId="9" pageNumber="24">Axial parenchyma</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9D9EFE09FC0C" box="[298,424,966,1009]" gridcol="1" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE54BF79D9EFD87FC0C" box="[438,550,966,1009]" gridcol="2" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE5487F9D9EFD12FC0C" box="[574,691,966,1009]" gridcol="3" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE548829D9EFC94FC0C" box="[707,821,966,1009]" gridcol="4" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE549089D9EFC1AFC0C" box="[841,955,966,1009]" gridcol="5" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE5498F9D9EFBEEFC0C" box="[974,1103,966,1009]" gridcol="6" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE54E249D9EFB4BFC0C" box="[1125,1258,966,1009]" gridcol="7" gridrow="6" pageId="9" pageNumber="24">Common</td>
<td id="76E3044ACC0D4AE54EB69D9EFA39FC0C" box="[1271,1432,966,1009]" gridcol="8" gridrow="6" pageId="9" pageNumber="24">Common</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9DA3FA39FBC2" box="[155,1432,1019,1087]" gridrow="7" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9DA3FEB7FBC2" box="[155,278,1019,1087]" gridcol="0" gridrow="7" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9DA3FF79FBC2" bold="true" pageId="9" pageNumber="24">Marginal axial parenchyma</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9DA3FE09FBC2" box="[298,424,1019,1087]" gridcol="1" gridrow="7" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54BF79DA3FD87FBC2" box="[438,550,1019,1087]" gridcol="2" gridrow="7" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE5487F9DA3FD12FBC2" box="[574,691,1019,1087]" gridcol="3" gridrow="7" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE548829DA3FC94FBC2" box="[707,821,1019,1087]" gridcol="4" gridrow="7" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE549089DA3FC1AFBC2" box="[841,955,1019,1087]" gridcol="5" gridrow="7" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE5498F9DA3FBEEFBC2" box="[974,1103,1019,1087]" gridcol="6" gridrow="7" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54E249DA3FB4BFBC2" box="[1125,1258,1019,1087]" gridcol="7" gridrow="7" pageId="9" pageNumber="24">Present and 1-3 seriate</td>
<td id="76E3044ACC0D4AE54EB69DA3FA39FBC2" box="[1271,1432,1019,1087]" gridcol="8" gridrow="7" pageId="9" pageNumber="24">Present, uniseriate</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9A1DFA39FB83" box="[155,1432,1093,1150]" gridrow="8" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9A1DFEB7FB83" box="[155,278,1093,1150]" gridcol="0" gridrow="8" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9A06FF55FB8C" bold="true" box="[155,244,1118,1137]" pageId="9" pageNumber="24">End walls</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9A1DFE09FB83" box="[298,424,1093,1150]" gridcol="1" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
<td id="76E3044ACC0D4AE54BF79A1DFD87FB83" box="[438,550,1093,1150]" gridcol="2" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
<td id="76E3044ACC0D4AE5487F9A1DFD12FB83" box="[574,691,1093,1150]" gridcol="3" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
<td id="76E3044ACC0D4AE548829A1DFC94FB83" box="[707,821,1093,1150]" gridcol="4" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
<td id="76E3044ACC0D4AE549089A1DFC1AFB83" box="[841,955,1093,1150]" gridcol="5" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
<td id="76E3044ACC0D4AE5498F9A1DFBEEFB83" box="[974,1103,1093,1150]" gridcol="6" gridrow="8" pageId="9" pageNumber="24">Smooth Pitted</td>
<td id="76E3044ACC0D4AE54E249A1DFB4BFB83" box="[1125,1258,1093,1150]" gridcol="7" gridrow="8" pageId="9" pageNumber="24">Smooth Pitted</td>
<td id="76E3044ACC0D4AE54EB69A1DFA39FB83" box="[1271,1432,1093,1150]" gridcol="8" gridrow="8" pageId="9" pageNumber="24">Distinctly pitted</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9AC9FA39FB09" box="[155,1432,1169,1268]" gridrow="9" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9AC9FEB7FB09" box="[155,278,1169,1268]" gridcol="0" gridrow="9" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9A9CFF69FB2A" bold="true" box="[155,200,1220,1239]" pageId="9" pageNumber="24">Rays</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9AC9FE09FB09" box="[298,424,1169,1268]" gridcol="1" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0D4AE54BF79AC9FD87FB09" box="[438,550,1169,1268]" gridcol="2" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0D4AE5487F9AC9FD12FB09" box="[574,691,1169,1268]" gridcol="3" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0D4AE548829AC9FC94FB09" box="[707,821,1169,1268]" gridcol="4" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0D4AE549089AC9FC1AFB09" box="[841,955,1169,1268]" gridcol="5" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, rarely partly biseriate</td>
<td id="76E3044ACC0D4AE5498F9AC9FBEEFB09" box="[974,1103,1169,1268]" gridcol="6" gridrow="9" pageId="9" pageNumber="24">Exclusively uniseriate, partly biseriate</td>
<td id="76E3044ACC0D4AE54E249AC9FB4BFB09" box="[1125,1258,1169,1268]" gridcol="7" gridrow="9" pageId="9" pageNumber="24">Uniseriate</td>
<td id="76E3044ACC0D4AE54EB69AC9FA39FB09" box="[1271,1432,1169,1268]" gridcol="8" gridrow="9" pageId="9" pageNumber="24">Rays exclusively uniseriate, rarely partly biseriate.</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9B4FFA39FABE" box="[155,1432,1303,1347]" gridrow="10" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9B4FFEB7FABE" box="[155,278,1303,1347]" gridcol="0" gridrow="10" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9B4FFF78FABE" bold="true" pageId="9" pageNumber="24">End walls of rays</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9B4FFE09FABE" box="[298,424,1303,1347]" gridcol="1" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE54BF79B4FFD87FABE" box="[438,550,1303,1347]" gridcol="2" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE5487F9B4FFD12FABE" box="[574,691,1303,1347]" gridcol="3" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE548829B4FFC94FABE" box="[707,821,1303,1347]" gridcol="4" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE549089B4FFC1AFABE" box="[841,955,1303,1347]" gridcol="5" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE5498F9B4FFBEEFABE" box="[974,1103,1303,1347]" gridcol="6" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE54E249B4FFB4BFABE" box="[1125,1258,1303,1347]" gridcol="7" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
<td id="76E3044ACC0D4AE54EB69B4FFA39FABE" box="[1271,1432,1303,1347]" gridcol="8" gridrow="10" pageId="9" pageNumber="24">Nodded</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9B17FA39FA75" box="[155,1432,1359,1416]" gridrow="11" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9B17FEB7FA75" box="[155,278,1359,1416]" gridcol="0" gridrow="11" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9B05FF79FA75" bold="true" box="[155,255,1373,1416]" pageId="9" pageNumber="24">Cross-field pitting</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9B17FE09FA75" box="[298,424,1359,1416]" gridcol="1" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE54BF79B17FD87FA75" box="[438,550,1359,1416]" gridcol="2" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE5487F9B17FD12FA75" box="[574,691,1359,1416]" gridcol="3" gridrow="11" pageId="9" pageNumber="24">Cupressoid and taxodioid</td>
<td id="76E3044ACC0D4AE548829B17FC94FA75" box="[707,821,1359,1416]" gridcol="4" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE549089B17FC1AFA75" box="[841,955,1359,1416]" gridcol="5" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE5498F9B17FBEEFA75" box="[974,1103,1359,1416]" gridcol="6" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE54E249B17FB4BFA75" box="[1125,1258,1359,1416]" gridcol="7" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
<td id="76E3044ACC0D4AE54EB69B17FA39FA75" box="[1271,1432,1359,1416]" gridcol="8" gridrow="11" pageId="9" pageNumber="24">Cupressoid</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9BC3FA39F9F1" box="[155,1432,1435,1548]" gridrow="12" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9BC3FEB7F9F1" box="[155,278,1435,1548]" gridcol="0" gridrow="12" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA9BC3FEA7F9F6" bold="true" pageId="9" pageNumber="24">Pit number per cross-field Ray heights</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9BC3FE09F9F1" box="[298,424,1435,1548]" gridcol="1" gridrow="12" pageId="9" pageNumber="24">1-2 (1-4) 3 (1-6)</td>
<td id="76E3044ACC0D4AE54BF79BC3FD87F9F1" box="[438,550,1435,1548]" gridcol="2" gridrow="12" pageId="9" pageNumber="24">2 (2-4) 1-4 (-13)</td>
<td id="76E3044ACC0D4AE5487F9BC3FD12F9F1" box="[574,691,1435,1548]" gridcol="3" gridrow="12" pageId="9" pageNumber="24">2 (1-4) 1-11 (-19)</td>
<td id="76E3044ACC0D4AE548829BC3FC94F9F1" box="[707,821,1435,1548]" gridcol="4" gridrow="12" pageId="9" pageNumber="24">1-2 (1-5) 1-8 (-19)</td>
<td id="76E3044ACC0D4AE549089BC3FC1AF9F1" box="[841,955,1435,1548]" gridcol="5" gridrow="12" pageId="9" pageNumber="24">1-2 (1-5) 1-7 (-14)</td>
<td id="76E3044ACC0D4AE5498F9BC3FBEEF9F1" box="[974,1103,1435,1548]" gridcol="6" gridrow="12" pageId="9" pageNumber="24">1-4 1-15</td>
<td id="76E3044ACC0D4AE54E249BC3FB4BF9F1" box="[1125,1258,1435,1548]" gridcol="7" gridrow="12" pageId="9" pageNumber="24">1-4 1-15</td>
<td id="76E3044ACC0D4AE54EB69BC3FA39F9F1" box="[1271,1432,1435,1548]" gridcol="8" gridrow="12" pageId="9" pageNumber="24">2-3 (-5) 2-6 (-16)</td>
</tr>
<tr id="35326D36CC0D4AE54ADA9872FA39F9B7" box="[155,1432,1578,1610]" gridrow="13" pageId="9" pageNumber="24">
<th id="76E3044ACC0D4AE54ADA9872FEB7F9B7" box="[155,278,1578,1610]" gridcol="0" gridrow="13" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB5134ADA986EFF57F9B4" bold="true" box="[155,246,1590,1609]" pageId="9" pageNumber="24">Indenture</emphasis>
</th>
<td id="76E3044ACC0D4AE54B6B9872FE09F9B7" box="[298,424,1578,1610]" gridcol="1" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE54BF79872FD87F9B7" box="[438,550,1578,1610]" gridcol="2" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE5487F9872FD12F9B7" box="[574,691,1578,1610]" gridcol="3" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE548829872FC94F9B7" box="[707,821,1578,1610]" gridcol="4" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE549089872FC1AF9B7" box="[841,955,1578,1610]" gridcol="5" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE5498F9872FBEEF9B7" box="[974,1103,1578,1610]" gridcol="6" gridrow="13" pageId="9" pageNumber="24">Present</td>
<td id="76E3044ACC0D4AE54E249872FB4BF9B7" box="[1125,1258,1578,1610]" gridcol="7" gridrow="13" pageId="9" pageNumber="24">Absent</td>
<td id="76E3044ACC0D4AE54EB69872FA39F9B7" box="[1271,1432,1578,1610]" gridcol="8" gridrow="13" pageId="9" pageNumber="24">Present</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBD6F74CC0DB5134AFC98FEFA89F7F5" blockId="9.[151,771,1701,2056]" lastBlockId="9.[816,1437,1701,2056]" pageId="9" pageNumber="24">
In 
<taxonomicName id="4C0214F7CC0DB5134AA498FDFED9F940" authorityName="Gray" authorityYear="1822" box="[229,376,1701,1725]" class="Pinopsida" family="Cupressaceae" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
, normal resin canals and helical thickenings are also absent. When some fossil species such as 
<taxonomicName id="4C0214F7CC0DB5134ABD98B0FE2EF902" box="[252,399,1768,1791]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="zamurense">
<emphasis id="B976B366CC0DB5134ABD98B0FE2EF902" box="[252,399,1768,1791]" italics="true" pageId="9" pageNumber="24">J. zamurense</emphasis>
</taxonomicName>
are related to different modern genera, 
<taxonomicName id="4C0214F7CC0DB5134AB49950FE47F8DD" authority="Kurz." authorityName="Kurz." box="[245,486,1800,1824]" class="Pinopsida" family="Cupressaceae" genus="Chamaecyparis" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB5134AB49950FE3DF8DD" box="[245,412,1800,1824]" italics="true" pageId="9" pageNumber="24">Chamaecyparis</emphasis>
Kurz.
</taxonomicName>
, 
<taxonomicName id="4C0214F7CC0DB5134BBC9951FDCDF8DD" box="[509,620,1801,1824]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB5134BBC9951FDCDF8DD" box="[509,620,1801,1824]" italics="true" pageId="9" pageNumber="24">Cupressus</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C0214F7CC0DB51348E99951FF79F8BC" authority="Lindl." authorityName="Lindl." class="Pinopsida" family="Cupressaceae" genus="Fitzroya" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB51348E99951FCA3F8DD" box="[680,770,1801,1824]" italics="true" pageId="9" pageNumber="24">Fitzroya</emphasis>
Lindl.
</taxonomicName>
and 
<taxonomicName id="4C0214F7CC0DB5134B529972FD3FF8BC" authority="(Ruiz and Bodnar, 2019)" baseAuthorityName="Ruiz and Bodnar" baseAuthorityYear="2019" box="[275,670,1833,1857]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB5134B529972FEDDF8BC" box="[275,380,1834,1857]" italics="true" pageId="9" pageNumber="24">Juniperus</emphasis>
(
<bibRefCitation id="EF931285CC0DB5134BCF9971FD37F8BC" author="Ruiz, D. P. &amp; Bodnar, J." box="[398,662,1833,1857]" pageId="9" pageNumber="24" pagination="481 - 488" refId="ref6337" refString="Ruiz, D. P. &amp; Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar, 2019</bibRefCitation>
)
</taxonomicName>
, the new species, 
<taxonomicName id="4C0214F7CC0DB5134AB49913FE4FF89E" authorityName="Akkemik" authorityYear="2021" box="[245,494,1867,1891]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="acarcae">
<emphasis id="B976B366CC0DB5134AB49913FE4FF89E" box="[245,494,1867,1891]" italics="true" pageId="9" pageNumber="24">Juniperoxylon acarcae</emphasis>
</taxonomicName>
, is closely related to the modern 
<taxonomicName id="4C0214F7CC0DB5134AB59935FEFCF879" baseAuthorityName="Ruiz and Bodnar" baseAuthorityYear="2019" box="[244,349,1901,1924]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0DB5134AB59935FEFCF879" box="[244,349,1901,1924]" italics="true" pageId="9" pageNumber="24">Juniperus</emphasis>
</taxonomicName>
. A comparison of the features of the new fossil species with the five modern 
<taxonomicName id="4C0214F7CC0DB513480799D6FCA2F858" box="[582,771,1933,1957]" pageId="9" pageNumber="24">
<emphasis id="B976B366CC0DB513480799D6FD0EF858" box="[582,687,1934,1957]" italics="true" pageId="9" pageNumber="24">Juniperus</emphasis>
species
</taxonomicName>
in 
<collectingCountry id="F3152FE4CC0DB5134AF299F6FEA1F83B" box="[179,256,1966,1990]" name="Turkey" pageId="9" pageNumber="24">Turkey</collectingCountry>
, which are native, and two similar species given in Esteban et al. (2004), which are 
<taxonomicName id="4C0214F7CC0DB51348739988FE2BF7F5" authority="Rehder et E. H. Wilson" authorityName="Rehder et E. H. Wilson" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="saltuaria">
<emphasis id="B976B366CC0DB51348739988FCA2F81A" box="[562,771,1999,2023]" italics="true" pageId="9" pageNumber="24">Juniperus saltuaria</emphasis>
Rehder et E.H. Wilson
</taxonomicName>
from south of 
<collectingCountry id="F3152FE4CC0DB513486999A8FDC9F7F5" box="[552,616,2032,2056]" name="China" pageId="9" pageNumber="24">China</collectingCountry>
and 
<collectingRegion id="49C6A196CC0DB51348DA99A8FD72F7F5" box="[667,723,2032,2056]" country="China" name="Xizang" pageId="9" pageNumber="24">Tibet</collectingRegion>
and 
<taxonomicName id="4C0214F7CC0DB513497198FEFBA4F940" box="[816,1029,1701,1725]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="thurifera">
<emphasis id="B976B366CC0DB513497198FEFBA4F940" box="[816,1029,1701,1725]" italics="true" pageId="9" pageNumber="24">Juniperus thurifera</emphasis>
</taxonomicName>
L. from the western Mediterranean Basin, showed the highest similarities. However, 
<taxonomicName id="4C0214F7CC0DB5134F27989FFBE1F902" authority="Willd." authorityName="Willd." class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="foetidissima">
<emphasis id="B976B366CC0DB5134F27989FFC55F902" italics="true" pageId="9" pageNumber="24">Juniperus foetidissima</emphasis>
Willd.
</taxonomicName>
has both cupressoid and taxodioid cross-field pits, and 
<taxonomicName id="4C0214F7CC0DB5134E789951FAABF8DD" box="[1081,1290,1800,1824]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="24" phylum="Tracheophyta" rank="species" species="saltuaria">
<emphasis id="B976B366CC0DB5134E789951FAABF8DD" box="[1081,1290,1800,1824]" italics="true" pageId="9" pageNumber="24">Juniperus saltuaria</emphasis>
</taxonomicName>
has not marginal and metatraheal axial parenchyma, therefore, these two species slightly differ from the fossil species and the other five modern Mediterranean species (
<tableCitation id="C6805ACFCC0DB5134F6E9934FA31F879" box="[1327,1424,1900,1924]" captionStart="Table 2" captionStartId="9.[285,344,220,241]" captionTargetPageId="9" captionText="Table 2. Comparison of modern Juniperus species (Esteban et al., 2004; Akkemik and Yaman, 2012) and Juniperoxylon acarcae sp. nov." pageId="9" pageNumber="24">Table 2</tableCitation>
). Wood features demonstrated that both fossil and modern species are rather close to each other, and therefore, the fossil species may be evaluated as a possible ancestor of the modern species in the Mediterranean basin.
</paragraph>
<paragraph id="8BBD6F74CC0EB5104AD69EC6FE8CFDBE" blockId="10.[151,772,158,976]" pageId="10" pageNumber="25">
<emphasis id="B976B366CC0EB5104AD69EC6FDB8FF4B" bold="true" box="[151,537,158,182]" pageId="10" pageNumber="25">Possible growing site conditions:</emphasis>
The described 
<taxonomicName id="4C0214F7CC0EB510488C9EC7FE70FF2A" authority="Akkemik" authorityName="Akkemik" authorityYear="2021" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="species" species="acarcae" status="sp. nov.">
<emphasis id="B976B366CC0EB510488C9EC7FEC2FF2A" italics="true" pageId="10" pageNumber="25">Juniperoxylon acarcae</emphasis>
Akkemik
</taxonomicName>
<taxonomicNameLabel id="A2450E1DCC0EB5104B9D9E98FD95FF2A" box="[476,564,192,215]" pageId="10" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
is a forest tree of the Galatian Volcanic Province (GVP). The new species was one of the elements within a rich woody flora of GVP (
<bibRefCitation id="EF931285CC0EB5104A9F9F7AFE63FEC7" author="Akkemik, U. &amp; Arslan M. &amp; Poole I. &amp; Tosun, S. &amp; Kose, N. &amp; Karlioglu Kilic, N. &amp; Aydin, A." box="[222,450,290,314]" pageId="10" pageNumber="25" pagination="31 - 50" refId="ref5392" refString="Akkemik, U., Arslan M., Poole I., Tosun, S., Kose, N., Karlioglu Kilic, N. &amp; Aydin, A., 2016. Silicified woods from two previously undescribed early Miocene forest sites near Seben, northwest Turkey. Review of Palaeobotany and Palynology 235: 31 - 50, https: // doi. org / 10.1016 / j. revpalbo. 2016.09.012" type="journal article" year="2016">Akkemik et al., 2016</bibRefCitation>
; 
<bibRefCitation id="EF931285CC0EB5104B939F7AFD54FEC7" author="Acarca Bayam, N. N. &amp; Akkemik, U. &amp; Poole, I." box="[466,757,290,315]" pageId="10" pageNumber="25" pagination="1 - 42" refId="ref5199" refString="Acarca Bayam, N. N., Akkemik, U., Poole, I., &amp; Akarsu. F., 2018. Further Contributions to the early Miocene forest vegetation of the Galatean Volcanic Province, Turkey. Palaeobotanica Electronica, 21.3.40 A 1 - 42. https: // doi. org / 10.26879 / 816 palaeoelectronica. org / content / 2018 / 2353 - trees-of-gvp-inmiocene" type="book chapter" year="2018">Acarca Bayam et al., 2018</bibRefCitation>
). The early Miocene trees of 
<taxonomicName id="4C0214F7CC0EB5104B8F9F1CFDAAFEA6" authorityName="Perry" authorityYear="1811" box="[462,523,324,347]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104B8F9F1CFDAAFEA6" box="[462,523,324,347]" italics="true" pageId="10" pageNumber="25">Pinus</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C0214F7CC0EB510487D9F1CFF74FE81" authority=", Cedrus Trew." authorityName="Cedrus Trew." class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB510487D9F1CFCA2FEA6" box="[572,771,323,347]" italics="true" pageId="10" pageNumber="25">Juniperus, Cedrus</emphasis>
Trew.
</taxonomicName>
, 
<taxonomicName id="4C0214F7CC0EB5104AA99F3CFE8CFE81" box="[232,301,356,380]" class="Magnoliopsida" family="Ulmaceae" genus="Ulmus" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104AA99F3CFE8CFE81" box="[232,301,356,380]" italics="true" pageId="10" pageNumber="25">Ulmus</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C0214F7CC0EB5104B229F3CFDABFE81" authority="Spach." authorityName="Spach." box="[355,522,356,380]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104B229F3CFE16FE81" box="[355,439,356,380]" italics="true" pageId="10" pageNumber="25">Zelkova</emphasis>
Spach.
</taxonomicName>
, 
<taxonomicName id="4C0214F7CC0EB510485C9F3CFD09FE81" box="[541,680,356,380]" class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" kingdom="Plantae" order="Saxifragales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB510485C9F3CFD09FE81" box="[541,680,356,380]" italics="true" pageId="10" pageNumber="25">Liquidambar</emphasis>
</taxonomicName>
L., and 
<taxonomicName id="4C0214F7CC0EB5104AD69FDDFF68FE60" box="[151,201,389,413]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104AD69FDDFF68FE60" box="[151,201,389,413]" italics="true" pageId="10" pageNumber="25">Acer</emphasis>
</taxonomicName>
L., grow under the conditions of VU0, and VU5 to VU 
<quantity id="4CFAC291CC0EB5104AFE9FFFFF4AFE42" box="[191,235,423,447]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="25" unit="in" value="7.0">7 in</quantity>
GVP. These vegetation units (VU) classify that VU0 is subtropical, moist or dry light forests, VU5 is well-drained lowland forests, VU6 is well-drained upland forests, and VU7 is well-drained (lowland and) upland conifer forest.
</paragraph>
<paragraph id="8BBD6F74CC0EB5104AD69C14FEAFFC2D" blockId="10.[151,772,158,976]" pageId="10" pageNumber="25">
The 
<taxonomicName id="4C0214F7CC0EB5104A8C9C14FE66FD99" authorityName="Akkemik" authorityYear="2021" box="[205,455,588,612]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="species" species="acarcae">
<emphasis id="B976B366CC0EB5104A8C9C14FE66FD99" box="[205,455,588,612]" italics="true" pageId="10" pageNumber="25">Juniperoxylon acarcae</emphasis>
</taxonomicName>
stems are autochthonous in the field. The other autochthonous early Miocene trees are 
<taxonomicName id="4C0214F7CC0EB5104A839CD7FEFAFD5B" authority="Sect." authorityName="Sect." box="[194,347,654,678]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104A839CD7FEBDFD5B" box="[194,284,655,678]" italics="true" pageId="10" pageNumber="25">Quercus</emphasis>
Sect.
</taxonomicName>
<emphasis id="B976B366CC0EB5104B259CD6FE33FD5B" box="[356,402,654,678]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C0214F7CC0EB5104B259CD6FE2EFD5B" box="[356,399,654,678]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">Ilex</taxonomicName>
,
</emphasis>
palms (later they were identified as 
<taxonomicName id="4C0214F7CC0EB5104AF99CF7FEC0FD3A" box="[184,353,687,711]" genus="Palmoxylon" kingdom="Animalia" pageId="10" pageNumber="25" rank="species" species="undetermined">
<emphasis id="B976B366CC0EB5104AF99CF7FE99FD3A" box="[184,312,687,711]" italics="true" pageId="10" pageNumber="25">Palmoxylon</emphasis>
sp.
</taxonomicName>
cf. 
<taxonomicName id="4C0214F7CC0EB5104BD39CF7FD87FD3A" box="[402,550,687,711]" class="Liliopsida" family="Arecaceae" genus="Trachycarpus" kingdom="Plantae" order="Arecales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104BD39CF7FD87FD3A" box="[402,550,687,711]" italics="true" pageId="10" pageNumber="25">Trachycarpus</emphasis>
</taxonomicName>
by Iamandei et al., 2018), 
<taxonomicName id="4C0214F7CC0EB5104AA89C89FED5FD14" box="[233,372,721,745]" class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" kingdom="Plantae" order="Saxifragales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">
<emphasis id="B976B366CC0EB5104AA89C89FED5FD14" box="[233,372,721,745]" italics="true" pageId="10" pageNumber="25">Liquidambar</emphasis>
</taxonomicName>
and 
<emphasis id="B976B366CC0EB5104BF29C89FDE6FD14" box="[435,583,721,745]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C0214F7CC0EB5104BF29C89FE49FD14" box="[435,488,721,745]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="genus">Salix</taxonomicName>
/Populus
</emphasis>
in Hoçaş Fossil Site (
<bibRefCitation id="EF931285CC0EB5104A909CAAFE1EFCF7" author="Akkemik, U. &amp; Arslan M. &amp; Poole I. &amp; Tosun, S. &amp; Kose, N. &amp; Karlioglu Kilic, N. &amp; Aydin, A." box="[209,447,754,778]" pageId="10" pageNumber="25" pagination="31 - 50" refId="ref5392" refString="Akkemik, U., Arslan M., Poole I., Tosun, S., Kose, N., Karlioglu Kilic, N. &amp; Aydin, A., 2016. Silicified woods from two previously undescribed early Miocene forest sites near Seben, northwest Turkey. Review of Palaeobotany and Palynology 235: 31 - 50, https: // doi. org / 10.1016 / j. revpalbo. 2016.09.012" type="journal article" year="2016">Akkemik et al., 2016</bibRefCitation>
). Because these fossil woods are found in their original positions in the field, to interpret their life conditions is more satisfactory. The presence of these riparian and lowland trees may suggest that the growing area of 
<taxonomicName id="4C0214F7CC0EB5104BC19D2EFDDBFC73" authorityName="Akkemik" authorityYear="2021" box="[384,634,886,910]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="25" phylum="Tracheophyta" rank="species" species="acarcae">
<emphasis id="B976B366CC0EB5104BC19D2EFDDBFC73" box="[384,634,886,910]" italics="true" pageId="10" pageNumber="25">Juniperoxylon acarcae</emphasis>
</taxonomicName>
trees was a forest having an open, well-drained lowland growing conditions.
</paragraph>
</subSubSection>
</treatment>
</document>