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<document id="9FF3DBC35A19DA2887E95E7C5704DA55" ID-DOI="10.5281/zenodo.214402" ID-GBIF-Dataset="fad9cc6c-e4d1-4f03-9c3e-6fd0dc5f5c4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="214402" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460526876090" checkinUser="plazi" docAuthor="Grootaert, Patrick &amp; Meuffels, Henk" docDate="2012" docId="03A987850A1AFFD946A15EEEFD493295" docLanguage="en" docName="zt03478p580.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Griphophanes gravicaudatus Grootaert &amp; Meuffels" docType="treatment" docVersion="9" lastPageNumber="575" masterDocId="FF90FFFD0A1EFFDC46365D3AFFEB3361" masterDocTitle="New Griphophanes Grootaert &amp; Meuffels (Diptera, Dolichopodidae, Peloropeodinae) from Southeast Asia" masterLastPageNumber="580" masterPageNumber="570" pageNumber="574" updateTime="1698315185162" updateUser="plazi">
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<mods:title id="3CFE42F42DC4895BD2B7B2C158A9A625">New Griphophanes Grootaert &amp; Meuffels (Diptera, Dolichopodidae, Peloropeodinae) from Southeast Asia</mods:title>
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<mods:namePart id="EB56BE4D089A0717169F34EFE5858DB9">Grootaert, Patrick</mods:namePart>
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<mods:namePart id="A2DE7FDAF04867DB12B96AC37DCC56D3">Meuffels, Henk</mods:namePart>
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<mods:date id="D912680249D85ECB66F396F3C2241A3A">2012</mods:date>
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<mods:identifier id="B768FF2D5A40AC6E646EAD80376105E3" type="DOI">10.5281/zenodo.214402</mods:identifier>
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<treatment id="03A987850A1AFFD946A15EEEFD493295" ID-DOI="http://doi.org/10.5281/zenodo.6174866" ID-GBIF-Taxon="119526295" ID-Zenodo-Dep="6174866" LSID="urn:lsid:plazi:treatment:03A987850A1AFFD946A15EEEFD493295" httpUri="http://treatment.plazi.org/id/03A987850A1AFFD946A15EEEFD493295" lastPageId="5" lastPageNumber="575" pageId="4" pageNumber="574">
<subSubSection id="C31A65180A1AFFD846A15EEEFED0376F" pageId="4" pageNumber="574" type="nomenclature">
<paragraph id="8BBF36930A1AFFD846A15EEEFCCA308C" blockId="4.[151,801,979,1038]" box="[151,801,979,1006]" pageId="4" pageNumber="574">
<heading id="D0F781FF0A1AFFD846A15EEEFCCA308C" bold="true" box="[151,801,979,1006]" fontSize="11" level="1" pageId="4" pageNumber="574" reason="1">
<taxonomicName id="4C004D100A1AFFD846A15EEEFCCA308C" authority="Grootaert &amp; Meuffels" authorityName="Grootaert &amp; Meuffels" box="[151,801,979,1006]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="gravicaudatus">
<emphasis id="B974EA810A1AFFD846A15EEEFCCA308C" bold="true" box="[151,801,979,1006]" pageId="4" pageNumber="574">
<emphasis id="B974EA810A1AFFD846A15EEEFE1D308F" bold="true" box="[151,502,980,1006]" italics="true" pageId="4" pageNumber="574">Griphophanes gravicaudatus</emphasis>
(Grootaert &amp; Meuffels)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBF36930A1AFFD846A15ECCFED0376F" blockId="4.[151,801,979,1038]" box="[151,315,1014,1038]" pageId="4" pageNumber="574">
(
<figureCitation id="133B2A160A1AFFD846A95ECCFEE1376F" box="[159,266,1014,1038]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[186,1408,547,1943]" captionTargetId="figure@5.[151,1436,515,1958]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Griphophanes gravicaudatus (Grootaert &amp; Meuffels, 1997) male: 5. antenna; 6. tip of hind tibia anteriorly; 7. postabdomen. Scale 0.1 mm." httpUri="https://zenodo.org/record/214405/files/figure.png" pageId="4" pageNumber="574">Figs. 5–7</figureCitation>
,
<figureCitation id="133B2A160A1AFFD847205ECCFEDA376F" box="[278,305,1014,1038]" captionStart="FIGURES 11 – 14" captionStartId="9.[151,262,1950,1973]" captionTargetBox="[226,1369,742,1929]" captionTargetId="figure@9.[217,1371,735,1929]" captionTargetPageId="9" captionText="FIGURES 11 – 14. Griphophanes males: 11. gravicaudatus (Grootaert &amp; Meuffels) habitus (scale 0.5 mm); 12. obscurus sp. nov. abdomen (scale 0.1 mm); 13. furcatulus sp. nov. habitus (scale 0.5 mm); 14. furcatus sp. nov. postabdomen (scale 0.1 mm)." httpUri="https://zenodo.org/record/214408/files/figure.png" pageId="4" pageNumber="574">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A65180A1AFFD846A15905FBF83712" pageId="4" pageNumber="574" type="reference_group">
<paragraph id="8BBF36930A1AFFD846A15905FC873735" blockId="4.[151,1043,1086,1139]" box="[151,876,1086,1109]" pageId="4" pageNumber="574">
<treatmentCitationGroup id="AB1011BD0A1AFFD846A15905FC873735" box="[151,876,1086,1109]" pageId="4" pageNumber="574">
<taxonomicName id="4C004D100A1AFFD846A15905FD1D3735" authority="Grootaert &amp; Meuffels, 1997: 108" authorityName="Grootaert &amp; Meuffels" authorityPageNumber="108" authorityYear="1997" box="[151,758,1086,1109]" class="Insecta" family="Tephritidae" genus="Griphomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="gravicaudata">
<emphasis id="B974EA810A1AFFD846A15905FE753734" box="[151,414,1087,1109]" italics="true" pageId="4" pageNumber="574">Griphomyia gravicaudata</emphasis>
<treatmentCitation id="0AA110820A1AFFD847935905FD1D3735" author="Grootaert" box="[421,758,1086,1108]" page="108" pageId="4" pageNumber="574" year="1997">
<bibRefCitation id="EF914B620A1AFFD847935905FD2F3735" author="Grootaert" box="[421,708,1086,1108]" pageId="4" pageNumber="574" refString="Grootaert, P. &amp; Meuffels, H. (1997) Griphomyia (Diptera, Dolichopodidae, Peloropeodinae) a new genus from southern Thailand. Belgian Journal of Zoology, 127, 107 - 114." type="journal article" year="1997">Grootaert &amp; Meuffels, 1997</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
, figs. 1–15.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBF36930A1AFFD846A15967FBF83712" blockId="4.[151,1043,1086,1139]" box="[151,1043,1117,1139]" pageId="4" pageNumber="574">
<treatmentCitationGroup id="AB1011BD0A1AFFD846A15967FBF83712" box="[151,1043,1117,1139]" pageId="4" pageNumber="574">
<taxonomicName id="4C004D100A1AFFD846A15967FC353712" authority="Grootaert &amp; Meuffels" authorityName="Grootaert &amp; Meuffels" authorityYear="1998" box="[151,990,1117,1139]" class="Insecta" family="Dolichopodidae" genus="Griphophanes" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="gravicaudatus">
<emphasis id="B974EA810A1AFFD846A15967FE553712" box="[151,446,1117,1139]" italics="true" pageId="4" pageNumber="574">Griphophanes gravicaudatus</emphasis>
(
<treatmentCitation id="0AA110820A1AFFD847FA5964FC353712" author="Grootaert" box="[460,990,1117,1139]" page="111" pageId="4" pageNumber="574" year="1998">
<bibRefCitation id="EF914B620A1AFFD847FA5964FC353712" author="Grootaert" box="[460,990,1117,1139]" pageId="4" pageNumber="574" refString="Grootaert, P. &amp; Meuffels, H. (1998) New data on Griphophanes nom. nov. from South Thailand (Diptera, Dolichopodidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, 68, 179." type="journal article" year="1998">Grootaert &amp; Meuffels): Grootaert &amp; Meuffels, 1998</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: 111.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31A65180A1AFFD846A159A5FBF9360D" pageId="4" pageNumber="574" type="materials_examined">
<paragraph id="8BBF36930A1AFFD846A159A5FBF9360D" blockId="4.[151,1436,1183,2036]" pageId="4" pageNumber="574">
<emphasis id="B974EA810A1AFFD846A159A5FE6A37D6" bold="true" box="[151,385,1183,1208]" pageId="4" pageNumber="574">Material examined.</emphasis>
<collectingCountry id="F31776030A1AFFD847BF59A5FDF437D6" box="[393,543,1183,1207]" name="Malaysia" pageId="4" pageNumber="574">MALAYSIA</collectingCountry>
: 2 3,
<emphasis id="B974EA810A1AFFD8445559A5FCBE37D9" bold="true" box="[611,853,1183,1208]" pageId="4" pageNumber="574">Endau-Rompin N.P.</emphasis>
(
<geoCoordinate id="EE3450540A1AFFD8455259A5FC5D37D6" box="[868,950,1183,1207]" direction="north" orientation="latitude" pageId="4" pageNumber="574" precision="925" value="2.5333333">2°32'N</geoCoordinate>
,
<geoCoordinate id="EE3450540A1AFFD845F459A5FBC637D6" box="[962,1069,1183,1207]" direction="east" orientation="longitude" pageId="4" pageNumber="574" precision="925" value="103.4">103°24'E</geoCoordinate>
; alt.
<quantity id="4CF89B760A1AFFD8425259A5FB7437D6" box="[1124,1183,1183,1207]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="574" unit="m" value="34.0">34 m</quantity>
) river banks, tributary Endau river,
<date id="FFBE10530A1AFFD8471059FEFE4537BD" box="[294,430,1220,1244]" pageId="4" pageNumber="574" value="2001-07-11">11.VII.2001</date>
(reg. 21062; leg. P. Grootaert); 1 Ƥ, Endau-Rompin, banks Endau river,
<date id="FFBE10530A1AFFD842EE59FEFA8A37BD" box="[1240,1377,1220,1244]" pageId="4" pageNumber="574" value="2001-07-12">12.VII.2001</date>
(reg. 21065); 1 3, Endau-Rompin, secondary rain forest along Endau river,
<date id="FFBE10530A1AFFD845F559DDFBA7379E" box="[963,1100,1255,1279]" pageId="4" pageNumber="574" value="2001-07-12">12.VII.2001</date>
(reg. 21066); 4 3,
<collectingCountry id="F31776030A1AFFD8431859DDFA7C379E" box="[1326,1431,1255,1279]" name="Malaysia" pageId="4" pageNumber="574">Malaysia</collectingCountry>
, Endau-Rompin, Sungei 1, tributary Endau river,
<date id="FFBE10530A1AFFD8448A5836FCAE3645" box="[700,837,1292,1316]" pageId="4" pageNumber="574" value="2001-07-13">13.VII.2001</date>
(reg. 21070); 15 3, Endau-Rompin, banks Sungei 2, tributary Endau river,
<date id="FFBE10530A1AFFD847955815FDC73626" box="[419,556,1327,1351]" pageId="4" pageNumber="574" value="2001-07-13">13.VII.2001</date>
(reg. 21072). 4 3, 2 Ƥ,
<emphasis id="B974EA810A1AFFD8455B5815FC0E3629" bold="true" box="[877,997,1327,1352]" pageId="4" pageNumber="574">Langkawi</emphasis>
(
<geoCoordinate id="EE3450540A1AFFD845CD5815FB983626" box="[1019,1139,1327,1351]" direction="north" orientation="latitude" pageId="4" pageNumber="574" precision="15" value="6.3722224">6°22'20''N</geoCoordinate>
,
<geoCoordinate id="EE3450540A1AFFD842B35815FAE33626" box="[1157,1288,1327,1351]" direction="east" orientation="longitude" pageId="4" pageNumber="574" precision="15" value="99.69416">99°40'99''E</geoCoordinate>
; alt.
<quantity id="4CF89B760A1AFFD8437A5815FA653626" box="[1356,1422,1327,1351]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="574" unit="m" value="54.0">54 m</quantity>
),
<date id="FFBE10530A1AFFD846A1586EFEE0360D" box="[151,267,1364,1388]" pageId="4" pageNumber="574" value="2005-09-03">3.IX.2005</date>
, rain forest (reg. 25326, Tio170, leg. P. Grootaert &amp; I. Van de Velde).
</paragraph>
</subSubSection>
<subSubSection id="C31A65180A1AFFD846F1584DFDAB36D5" pageId="4" pageNumber="574" type="diagnosis">
<paragraph id="8BBF36930A1AFFD846F1584DFDAB36D5" blockId="4.[151,1436,1183,2036]" pageId="4" pageNumber="574">
<emphasis id="B974EA810A1AFFD846F1584DFED136F1" bold="true" box="[199,314,1399,1424]" pageId="4" pageNumber="574">Diagnosis</emphasis>
: Medium-sized species with yellow legs. Acr uniseriate. No bristling on fore and hind femora. Cerci not-bifurcate but bent in a right angle.
</paragraph>
</subSubSection>
<subSubSection id="C31A65180A1AFFD946F15885FB3F32AE" lastPageId="5" lastPageNumber="575" pageId="4" pageNumber="574" type="description">
<paragraph id="8BBF36930A1AFFD946F15885FB3F32AE" blockId="4.[151,1436,1183,2036]" lastBlockId="5.[151,1436,151,500]" lastPageId="5" lastPageNumber="575" pageId="4" pageNumber="574">
<emphasis id="B974EA810A1AFFD846F15885FE5536B9" bold="true" box="[199,446,1471,1496]" pageId="4" pageNumber="574">Redescription. Male.</emphasis>
Body length
<quantity id="4CF89B760A1AFFD8446E5885FD5936B6" box="[600,690,1471,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="574" unit="mm" value="1.7">1.7 mm</quantity>
; wing length
<quantity id="4CF89B760A1AFFD8457A5885FC4D36B6" box="[844,934,1471,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="574" unit="mm" value="1.3">1.3 mm</quantity>
.
<emphasis id="B974EA810A1AFFD8458458FAFC1A36B9" bold="true" box="[946,1009,1472,1496]" pageId="4" pageNumber="574">Head</emphasis>
. Frons and face with blackish ground colour. Eyes nearly touching each other below middle of face. Palpus brown, with small, black apical bristle. Rostrum dark brown. Occiput black. Postoculars all black. Antenna (
<figureCitation id="133B2A160A1AFFD8459F5B3DFC06357E" box="[937,1005,1543,1567]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[186,1408,547,1943]" captionTargetId="figure@5.[151,1436,515,1958]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Griphophanes gravicaudatus (Grootaert &amp; Meuffels, 1997) male: 5. antenna; 6. tip of hind tibia anteriorly; 7. postabdomen. Scale 0.1 mm." httpUri="https://zenodo.org/record/214405/files/figure.png" pageId="4" pageNumber="574">Fig. 5</figureCitation>
) dark brown. Third segment about as long as deep, triangular, with rather blunt apex. Arista dorsal, more than
<date id="FFBE10530A1AFFD845F85B16FC1A3525" box="[974,1009,1580,1604]" pageId="4" pageNumber="574">3X</date>
as long as antenna, bare; basal aristal segment half as long as third antennal segment.
<emphasis id="B974EA810A1AFFD8449A5B6AFCEF3509" bold="true" box="[684,772,1616,1640]" pageId="4" pageNumber="574">Thorax</emphasis>
. Mesoscutum and scutellum greenish black, feebly shining. Pleurae brownish black to dark brown, partly with green metallic gloss. Acr uniseriate. Legs. Fore coxa yellow, mid and hind coxae brown. Legs, including trochanters, yellow. Tarsi indistinctly darkened towards tips.
<emphasis id="B974EA810A1AFFD842C55BA2FAB835D1" bold="true" box="[1267,1363,1688,1712]" pageId="4" pageNumber="574">Fore leg</emphasis>
. Coxa anteriorly with few, very short, dark hairs; at apex a few short and weak bristles. Trochanter bare. Femur and tibia without bristles; tibia about as long as femur. Length of tibia and tarsal segments (in mm): 0.35: 0.21: 0.08: 0.07: 0.04: 0.05.
<emphasis id="B974EA810A1AFFD8472F5A39FE98347D" bold="true" box="[281,371,1795,1820]" pageId="4" pageNumber="574">Mid leg</emphasis>
. Coxa anteriorly with a few, very short hairs; exteriorly with short, weak bristle. Trochanter with very small bristle. Femur without bristles, with only 1 or 2 tiny preapical pv. Tibia slightly longer than femur; 1 ad and 1 pd at basal fourth; a few short and thin apicals. Length of tibia and tarsal segments (in mm): 0.55: 0.24: 0.15: 0.11: 0.08: 0.07.
<emphasis id="B974EA810A1AFFD847485A55FE0D34E9" bold="true" box="[382,486,1903,1928]" pageId="4" pageNumber="574">
<collectingCountry id="F31776030A1AFFD847485A55FE5134E9" box="[382,442,1903,1928]" name="India" pageId="4" pageNumber="574">Hind</collectingCountry>
leg
</emphasis>
. Coxa with thin and weak exterior bristle. Trochanter bare. Femur without bristles. Tibia slightly longer than femur; tiny ad at basal fourth; 1 dorsal bristle near apex (
<figureCitation id="133B2A160A1AFFD8425C5AAEFB5934CD" box="[1130,1202,1940,1964]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[186,1408,547,1943]" captionTargetId="figure@5.[151,1436,515,1958]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Griphophanes gravicaudatus (Grootaert &amp; Meuffels, 1997) male: 5. antenna; 6. tip of hind tibia anteriorly; 7. postabdomen. Scale 0.1 mm." httpUri="https://zenodo.org/record/214405/files/figure.png" pageId="4" pageNumber="574">Fig. 6</figureCitation>
), preceded by some lengthened hairs. Tip of hind tibia with scale-like appendage ventrally. Length of tibia and tarsal segments (in mm): 0.53: 0.16: 0.2: 0.15: 0.1: 0.07.
<emphasis id="B974EA810A1AFFD8442C5AE1FDB13495" bold="true" box="[538,602,2011,2036]" pageId="4" pageNumber="574">Wing</emphasis>
. Brownish tinged with brown veins. Tp about half as long as apical part of Cu. Halter darkened. Squama brown, with black border, and 5 long, black cilia.
<emphasis id="B974EA810A1BFFD942145DADFB7D33D1" bold="true" box="[1058,1174,151,176]" pageId="5" pageNumber="575">Abdomen</emphasis>
above blackish brown, feebly shining; sterna brownish. Hairs on terga small, black; hind marginal bristles long on first tergum, short on following terga. Hypopygium (
<figureCitation id="133B2A160A1BFFD947C15DE5FDD03396" box="[503,571,223,247]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[186,1408,547,1943]" captionTargetId="figure@5.[151,1436,515,1958]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Griphophanes gravicaudatus (Grootaert &amp; Meuffels, 1997) male: 5. antenna; 6. tip of hind tibia anteriorly; 7. postabdomen. Scale 0.1 mm." httpUri="https://zenodo.org/record/214405/files/figure.png" pageId="5" pageNumber="575">Fig. 7</figureCitation>
) brownish black; cerci small, yellowish-brown, with tip bent less than right angle. Ventral surstylus digitiform with strong apical bristle. Dorsal surstylus twice as broad as ventral surstylus also bearing apical spine-like bristle. Epandrial lobe very large without epandrial bristles. Detailed description of complex aedeagus and asymmetrical hypandrium found in
<bibRefCitation id="EF914B620A1BFFD945115C76FB873205" author="Grootaert" box="[807,1132,332,356]" pageId="5" pageNumber="575" refString="Grootaert, P. &amp; Meuffels, H. (1997) Griphomyia (Diptera, Dolichopodidae, Peloropeodinae) a new genus from southern Thailand. Belgian Journal of Zoology, 127, 107 - 114." type="journal article" year="1997">Grootaert &amp; Meuffels (1997)</bibRefCitation>
.
<emphasis id="B974EA810A1BFFD9424E5C76FB253205" bold="true" box="[1144,1230,332,356]" pageId="5" pageNumber="575">Female</emphasis>
. Body length
<quantity id="4CF89B760A1BFFD9435D5C76FF2D32E9" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="575" unit="mm" value="1.55">1.55 mm</quantity>
; wing length
<quantity id="4CF89B760A1BFFD947555C55FE5432E9" box="[355,447,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="575" unit="mm" value="1.3">1.3 mm</quantity>
. Resembles male except as follows: Face at narrowest point about as wide as distance between ocellar bristles. Palpus and rostrum slightly larger than in male and more yellowish brown. Oviscapt yellow, with row of 6 small acanthae (2 black acanthae on each side, and 2 yellowish acanthae in middle).
</paragraph>
</subSubSection>
<subSubSection id="C31A65180A1BFFD946F15CE1FD493295" box="[199,674,475,500]" pageId="5" pageNumber="575" type="distribution">
<paragraph id="8BBF36930A1BFFD946F15CE1FD493295" blockId="5.[151,1436,151,500]" box="[199,674,475,500]" pageId="5" pageNumber="575">
<emphasis id="B974EA810A1BFFD946F15CE1FEB23295" bold="true" box="[199,345,475,500]" pageId="5" pageNumber="575">Distribution</emphasis>
:
<collectingCountry id="F31776030A1BFFD9475E5CE6FE203295" box="[360,459,476,500]" name="Thailand" pageId="5" pageNumber="575">Thailand</collectingCountry>
(South),
<collectingCountry id="F31776030A1BFFD944025CE6FD753295" box="[564,670,476,500]" name="Malaysia" pageId="5" pageNumber="575">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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