410 lines
55 KiB
XML
410 lines
55 KiB
XML
<document id="7E9DFBAF9F132D4EF864B3183268438B" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A963622879FF852E7719930C89FB3D" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia golovatchi Antić & Reip 2020, sp. nov." docType="treatment" docUuid="AFC4B93A-128A-4FF6-9E97-726F4DDE4CA1" docUuidSource="ZooBank" docVersion="11" lastPageNumber="41" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="37" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ACE2D265CE4AC5F743B5A35D576CC127">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:name id="6809FE5C9DC6CE83DB175587D0DA5F63" type="personal">
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<mods:role id="0099D666CEF7AFB025C9757EBE307C78">
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<mods:roleTerm id="B222F891B91C2D7E18EF503D0124DBAF">Author</mods:roleTerm>
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<mods:namePart id="B4BA9B606EEC5F62E7C966B91DB8A6DB">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="7E73D540A86B4AC9837D528EED53C89C">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="4ECCD6A089CF1EEF55E8EEEFB6DB93EA" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="9D3F5254D6CCBBFA1633E03783F6BF13">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="23905E3FB3C23918D7738AA6BCF41483" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:title id="21BA4AEDE45839E9E1307D6D46A89CCD">European Journal of Taxonomy</mods:title>
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<mods:part id="18BC0BEDAF0DC1E3981B74061571935A">
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<mods:date id="132CFC64E877DEB4EFF8816B448FDF16">2020</mods:date>
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<mods:number id="EE6DD26F2129C2D21B6D929645DE72E7">2020-09-07</mods:number>
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<mods:detail id="DF331AAAA12A08AA1177224F38FFA84E" type="volume">
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<mods:number id="813633C6749EBDE0FD6DA047436F7556">713</mods:number>
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<treatment id="03A963622879FF852E7719930C89FB3D" ID-DOI="http://doi.org/10.5281/zenodo.4335818" ID-GBIF-Taxon="167523038" ID-Zenodo-Dep="4335818" LSID="urn:lsid:zoobank.org:act:AFC4B93A-128A-4FF6-9E97-726F4DDE4CA1" httpUri="http://treatment.plazi.org/id/03A963622879FF852E7719930C89FB3D" lastPageId="40" lastPageNumber="41" pageId="36" pageNumber="37">
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<subSubSection id="C31A81FF2879FF892E7719930CE8FE26" box="[595,992,429,456]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BBFD2742879FF892E7719930CE8FE26" blockId="36.[359,1228,429,526]" box="[595,992,429,456]" pageId="36" pageNumber="37">
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<heading id="D0F765182879FF892E7719930CE8FE26" box="[595,992,429,456]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
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<taxonomicName id="4C00A9F72879FF892E7719930C76FE26" ID-CoL="849PQ" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[595,894,429,456]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
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<emphasis id="B9740E662879FF892E7719930C76FE26" bold="true" box="[595,894,429,456]" italics="true" pageId="36" pageNumber="37">Leucogeorgia golovatchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892FA219900CE8FE26" box="[902,992,430,456]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2879FF892D4319EE0C8FFDE0" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BBFD2742879FF892D4319EE0BC4FE04" blockId="36.[359,1228,429,526]" box="[359,1228,464,490]" pageId="36" pageNumber="37">
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<uri id="FF91DE762879FF892D4319EE0BC4FE04" box="[359,1228,464,490]" pageId="36" pageNumber="37">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA6E8A12879FF892EA319EE0BC4FE04" box="[647,1228,464,490]" pageId="36" pageNumber="37">AFC4B93A-128A-4FF6-9E97-726F4DDE4CA1</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BBFD2742879FF892E8819CD0C8FFDE0" blockId="36.[359,1228,429,526]" box="[684,903,499,526]" pageId="36" pageNumber="37">
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<figureCitation id="133BCEF12879FF892E8819CD0C3AFDE0" box="[684,818,499,526]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="37.[189,232,1895,1921]" captionStartId-1="38.[189,232,1858,1884]" captionStartId-2="39.[189,232,1930,1956]" captionTargetBox-0="[281,1307,265,1868]" captionTargetBox-1="[190,1400,391,1837]" captionTargetBox-2="[260,1319,268,1908]" captionTargetId-0="figure@37.[281,1307,265,1868]" captionTargetId-1="figure@38.[190,1400,391,1837]" captionTargetId-2="figure@39.[247,1353,250,1912]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." captionText-1="Fig. 23. Leucogeorgia golovatchi sp. nov., paratype ♂ from Tsakhi Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Right gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A, C–D = 0.1 mm; B = 0.05 mm." captionText-2="Fig. 24. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM). A. Holotype, right gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, penes, posterior view. D. Paratype, leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020798" figureDoi-1="http://doi.org/10.5281/zenodo.4020800" figureDoi-2="http://doi.org/10.5281/zenodo.4020802" httpUri-0="https://zenodo.org/record/4020798/files/figure.png" httpUri-1="https://zenodo.org/record/4020800/files/figure.png" httpUri-2="https://zenodo.org/record/4020802/files/figure.png" pageId="36" pageNumber="37">Figs 22–24</figureCitation>
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,
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<figureCitation id="133BCEF12879FF892F1A19CD0C56FDE0" box="[830,862,499,526]" captionStart="Fig" captionStartId="96.[189,232,1921,1947]" captionTargetBox="[189,1399,979,1893]" captionTargetId="figure@96.[189,1399,979,1893]" captionTargetPageId="96" captionText="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020873" httpUri="https://zenodo.org/record/4020873/files/figure.png" pageId="36" pageNumber="37">56</figureCitation>
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,
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<figureCitation id="133BCEF12879FF892F4E19CD0C8FFDE0" box="[874,903,499,526]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="36" pageNumber="37">58</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2879FF892C991A020E81FC2C" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="8BBFD2742879FF892C991A020E3EFDB8" blockId="36.[189,310,572,598]" box="[189,310,572,598]" pageId="36" pageNumber="37">
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<heading id="D0F765182879FF892C991A020E3EFDB8" bold="true" box="[189,310,572,598]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
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<emphasis id="B9740E662879FF892C991A020E3EFDB8" bold="true" box="[189,310,572,598]" pageId="36" pageNumber="37">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742879FF892C991A550E81FC2C" blockId="36.[189,1399,619,962]" pageId="36" pageNumber="37">
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This species belongs to the group of
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<taxonomicName id="4C00A9F72879FF892E571A520C1DFD6B" authorityName="Verhoeff" authorityYear="1930" box="[627,789,620,645]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662879FF892E571A520C1DFD6B" box="[627,789,620,645]" italics="true" pageId="36" pageNumber="37">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. with neither modified mouthparts nor teeth on the mesomeral claw (vs teeth present in
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<taxonomicName id="4C00A9F72879FF892EBB1AB10C3FFD46" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[671,823,654,680]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662879FF892EBB1AB10C3FFD46" box="[671,823,654,680]" italics="true" pageId="36" pageNumber="37">L. abchasica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C00A9F72879FF892F631AB10CCEFD46" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[839,966,654,680]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
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<emphasis id="B9740E662879FF892F631AB10CCEFD46" box="[839,966,654,680]" italics="true" pageId="36" pageNumber="37">L. borealis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892FEB1AB10B26FD47" box="[975,1070,655,681]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C00A9F72879FF89281F1AB10B82FD46" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1083,1162,655,680]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
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<emphasis id="B9740E662879FF89281F1AB10B82FD46" box="[1083,1162,655,680]" italics="true" pageId="36" pageNumber="37">L. gioi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF8928B61AB10BF9FD47" box="[1170,1265,655,681]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C00A9F72879FF8928DA1AB10A7EFD46" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1278,1398,654,680]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
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<emphasis id="B9740E662879FF8928DA1AB10A7EFD46" box="[1278,1398,654,680]" italics="true" pageId="36" pageNumber="37">L. oculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892C991A8C0E1FFD22" box="[189,279,690,716]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C00A9F72879FF892D691A8C0EC2FD25" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[333,458,690,715]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="satunini">
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<emphasis id="B9740E662879FF892D691A8C0EC2FD25" box="[333,458,690,715]" italics="true" pageId="36" pageNumber="37">L. satunini</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C00A9F72879FF892DC51A8C0C0DFD25" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[481,773,689,715]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
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<emphasis id="B9740E662879FF892DC51A8C0C0DFD25" box="[481,773,689,715]" italics="true" pageId="36" pageNumber="37">Leucogeorgia golovatchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892F281A8C0C6DFD22" box="[780,869,690,716]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C00A9F72879FF892FD91A8C0B6EFD25" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1021,1126,689,715]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
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<emphasis id="B9740E662879FF892FD91A8C0B6EFD25" box="[1021,1126,689,715]" italics="true" pageId="36" pageNumber="37">L. lobata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF8928481A8C0BCEFD22" box="[1132,1222,690,716]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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by the absence of a strongly developed lobe on the mesomeral lamella (vs presence of a strongly developed lobe in
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<taxonomicName id="4C00A9F72879FF892C991AC70E22FCFC" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,298,760,786]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
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<emphasis id="B9740E662879FF892C991AC70E22FCFC" box="[189,298,760,786]" italics="true" pageId="36" pageNumber="37">L. lobata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892D111AC60E9FFCFC" box="[309,407,760,786]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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).
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<taxonomicName id="4C00A9F72879FF892D8B1AC70DDFFCFC" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[431,727,760,786]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
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<emphasis id="B9740E662879FF892D8B1AC70DDFFCFC" box="[431,727,760,786]" italics="true" pageId="36" pageNumber="37">Leucogeorgia golovatchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2879FF892EC61AC60C48FCFC" box="[738,832,760,786]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
|
||
<taxonomicName id="4C00A9F72879FF892FC11AC70B87FCFC" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[997,1167,760,786]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
|
||
<emphasis id="B9740E662879FF892FC11AC70B87FCFC" box="[997,1167,760,786]" italics="true" pageId="36" pageNumber="37">L. prometheus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2879FF8928BE1AC60BF0FCFC" box="[1178,1272,760,786]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by both a shorter mesomeral claw and a long solenomere clearly exceeding the height of the mesomeral lamella (vs a longer mesomeral claw and a short solenomere not exceeding the height of the mesomeral lamella in
|
||
<taxonomicName id="4C00A9F72879FF892CF81B5C0E8CFC95" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[220,388,865,891]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
|
||
<emphasis id="B9740E662879FF892CF81B5C0E8CFC95" box="[220,388,865,891]" italics="true" pageId="36" pageNumber="37">L. prometheus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2879FF892DA81B5C0EE3FC92" box="[396,491,866,892]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Additionally,
|
||
<taxonomicName id="4C00A9F72879FF892E851B5C0C35FC95" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[673,829,865,891]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
|
||
<emphasis id="B9740E662879FF892E851B5C0C35FC95" box="[673,829,865,891]" italics="true" pageId="36" pageNumber="37">L. golovatchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2879FF892F611B5C0CA8FC92" box="[837,928,866,892]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
clearly differs from all other congeners by having strongly developed ventral lobes on body ring
|
||
<quantity id="4CF87F912879FF892F731BBB0C8CFC71" box="[855,900,901,927]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="36" pageNumber="37" unit="in" value="2.0">2 in</quantity>
|
||
both sexes (vs absence of such lobes in all other congeners).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2879FF892C991BCF0B3AFB6E" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="8BBFD2742879FF892C991BCF0E4CFBE5" blockId="36.[189,324,1009,1035]" box="[189,324,1009,1035]" pageId="36" pageNumber="37">
|
||
<heading id="D0F765182879FF892C991BCF0E4CFBE5" bold="true" box="[189,324,1009,1035]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
|
||
<emphasis id="B9740E662879FF892C991BCF0E4CFBE5" bold="true" box="[189,324,1009,1035]" pageId="36" pageNumber="37">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991C210B3AFB6E" blockId="36.[189,1399,1055,1152]" pageId="36" pageNumber="37">This new species is dedicated to Sergei Golovatch, one of the collectors, a well-known diplopodologist worldwide, and our good friend and colleague. His collection of millipedes from the Caucasus, now in ZMUM, is one of the largest and most complete. Noun in the genitive case.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2879FF892C991C910DA4F9FA" pageId="36" pageNumber="37" type="materials_examined">
|
||
<paragraph id="8BBFD2742879FF892C991C910EA2FB27" blockId="36.[189,426,1199,1225]" box="[189,426,1199,1225]" pageId="36" pageNumber="37">
|
||
<heading id="D0F765182879FF892C991C910EA2FB27" bold="true" box="[189,426,1199,1225]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
|
||
<emphasis id="B9740E662879FF892C991C910EA2FB27" bold="true" box="[189,426,1199,1225]" pageId="36" pageNumber="37">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B68D8292879FF892CCE1CE00B45FAD0" ID-GBIF-Occurrence="2858644349" collectingDate="1939-07-30" collectionCode="ZMUM" collectorName="J. Birstein" country="Georgia" county="Cave" latitude="42.54" location="Racha-Lechkhumi" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BBFD2742879FF892CCE1CE00E54FB16" blockId="36.[189,1399,1246,1343]" box="[234,348,1246,1272]" pageId="36" pageNumber="37">
|
||
<emphasis id="B9740E662879FF892CCE1CE00E54FB16" bold="true" box="[234,348,1246,1272]" pageId="36" pageNumber="37">
|
||
<typeStatus id="54BB6CD62879FF892CCE1CE00E54FB16" box="[234,348,1246,1272]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991D3F0B45FAD0" blockId="36.[189,1399,1246,1343]" pageId="36" pageNumber="37">
|
||
CENTRAL-WEST
|
||
<collectingCountry id="F31792E42879FF892D831D3F0D21FAF5" box="[423,553,1281,1307]" name="Georgia" pageId="36" pageNumber="37">GEORGIA</collectingCountry>
|
||
–
|
||
<emphasis id="B9740E662879FF892E681D3E0C3DFAF4" bold="true" box="[588,821,1280,1306]" pageId="36" pageNumber="37">
|
||
<location id="8EDF84AF2879FF892E681D3E0C3DFAF4" LSID="urn:lsid:plazi:treatment:03A963622879FF852E7719930C89FB3D:8EDF84AF2879FF892E681D3E0C3DFAF4" box="[588,821,1280,1306]" country="Georgia" county="Cave" latitude="42.54" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" name="Racha-Lechkhumi" pageId="36" pageNumber="37">Racha-Lechkhumi</location>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D0619FD2879FF892F771CC10C62FAF5" box="[851,874,1279,1307]" pageId="36" pageNumber="37" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8EDF84AF2879FF892F5D1D3E0B6FFAF4" LSID="urn:lsid:plazi:treatment:03A963622879FF852E7719930C89FB3D:8EDF84AF2879FF892F5D1D3E0B6FFAF4" box="[889,1127,1280,1306]" country="Georgia" county="Cave" latitude="42.54" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" name="Ambrolauri District" pageId="36" pageNumber="37">Ambrolauri District</location>
|
||
,
|
||
<locationDeviation id="9A0DCB562879FF8928511D3F0A13FAF4" box="[1141,1307,1280,1307]" pageId="36" pageNumber="37">near Gogoleti</locationDeviation>
|
||
,
|
||
<location id="8EDF84AF2879FF89290D1D3E0E9AFAD0" LSID="urn:lsid:plazi:treatment:03A963622879FF852E7719930C89FB3D:8EDF84AF2879FF89290D1D3E0E9AFAD0" country="Georgia" county="Cave" latitude="42.54" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" name="Tsakhi (= Gogoleti) Cave" pageId="36" pageNumber="37">Tsakhi (= Gogoleti) Cave</location>
|
||
;
|
||
<geoCoordinate id="EE34B4B32879FF892D861D1A0D07FAD0" box="[418,527,1316,1343]" degrees="42.54" direction="north" orientation="latitude" pageId="36" pageNumber="37" precision="555" value="42.54">42.54° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32879FF892E3F1D1A0D8DFAD0" box="[539,645,1316,1342]" degrees="42.90" direction="east" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="42.9">42.90° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42879FF892EB61D1A0C2DFAD0" box="[658,805,1316,1342]" pageId="36" pageNumber="37" value="1939-07-30">
|
||
<collectingDate id="EFFA0D5C2879FF892EB61D1A0C2DFAD0" box="[658,805,1316,1342]" pageId="36" pageNumber="37" value="1939-07-30">30 Jul. 1939</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22879FF892F151D1A0CA0FAD0" box="[817,936,1316,1342]" pageId="36" pageNumber="37">J. Birstein</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12879FF892FCE1D1A0B41FAD0" box="[1002,1097,1316,1342]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="36" pageNumber="37">ZMUM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BBFD2742879FF892CCE1D530E6FFA69" blockId="36.[189,1398,1389,1556]" box="[234,359,1389,1415]" pageId="36" pageNumber="37">
|
||
<emphasis id="B9740E662879FF892CCE1D530E6FFA69" bold="true" box="[234,359,1389,1415]" pageId="36" pageNumber="37">
|
||
<typeStatus id="54BB6CD62879FF892CCE1D530E6FFA69" box="[234,359,1389,1415]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991DAE0DA4F9FA" blockId="36.[189,1398,1389,1556]" pageId="36" pageNumber="37">
|
||
<materialsCitation id="3B68D8292879FF892C991DAE0C3AFA44" ID-GBIF-Occurrence="2858644341" box="[189,818,1424,1450]" collectorName="Racha-Lechkhumi" country="Georgia" location="Racha-Lechkhumi" pageId="36" pageNumber="37" specimenCount="1" typeStatus="paratype">
|
||
CENTRAL-WEST
|
||
<collectingCountry id="F31792E42879FF892D821DAE0D20FA44" box="[422,552,1424,1450]" name="Georgia" pageId="36" pageNumber="37">GEORGIA</collectingCountry>
|
||
–
|
||
<emphasis id="B9740E662879FF892E6D1DAE0C3AFA44" bold="true" box="[585,818,1424,1450]" pageId="36" pageNumber="37">
|
||
<location id="8EDF84AF2879FF892E6D1DAE0C3AFA44" LSID="urn:lsid:plazi:treatment:03A963622879FF852E7719930C89FB3D:8EDF84AF2879FF892E6D1DAE0C3AFA44" box="[585,818,1424,1450]" country="Georgia" name="Racha-Lechkhumi" pageId="36" pageNumber="37">Racha-Lechkhumi</location>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292879FF892F6A1DAE0E98FA20" ID-GBIF-Occurrence="2858644400" collectingDate="1939-07-30" collectionCode="ZMUM" collectorName="J. Birstein" country="Georgia" county="Cave" latitude="42.54" location="Racha-Lechkhumi" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" pageId="36" pageNumber="37" specimenCount="15" specimenCount-female="1" specimenCount-male="8" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2879FF892F6A1DAE0C9AFA44" box="[846,914,1422,1450]" pageId="36" pageNumber="37" type="male">8 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2879FF892F851DAE0CC7FA44" box="[929,975,1424,1450]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2879FF892FFB1DAE0B23FA44" box="[991,1067,1424,1450]" pageId="36" pageNumber="37">6 juvs</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12879FF892D111D8A0E98FA20" box="[309,400,1460,1486]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="36" pageNumber="37">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292879FF892D891D8D0CD3FA20" ID-GBIF-Occurrence="2858644323" box="[429,987,1457,1486]" collectingDate="1939-07-30" collectionCode="IZB" collectorName="J. Birstein" country="Georgia" county="Cave" latitude="42.54" location="Racha-Lechkhumi" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2879FF892D891D8D0ED3FA23" box="[429,475,1457,1485]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12879FF892F881D8A0CD3FA20" box="[940,987,1460,1486]" pageId="36" pageNumber="37">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292879FF892FDC1D8D0E85FA1E" ID-GBIF-Occurrence="2858644371" collectingDate="1939-07-30" collectionCode="SMNG" collectorName="J. Birstein" country="Georgia" county="Cave" latitude="42.54" location="Racha-Lechkhumi" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2879FF892FDC1D8D0B2FFA23" box="[1016,1063,1457,1485]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED114AB12879FF892D101DE80E85FA1E" box="[308,397,1494,1520]" collectionName="Germany, Goerlitz, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="36" pageNumber="37">SMNG</collectionCode>
|
||
</materialsCitation>
|
||
; •
|
||
<materialsCitation id="3B68D8292879FF892D8F1DE90DA0F9FA" ID-GBIF-Occurrence="2858644384" collectingDate="1974-08-09" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Georgia" county="Cave" latitude="42.54" location="Racha-Lechkhumi" longLatPrecision="689" longitude="42.9" municipality="Tsakhi" pageId="36" pageNumber="37" specimenCount="17" specimenCount-female="3" specimenCount-male="7" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2879FF892D8F1DE90EE6FA1F" box="[427,494,1493,1521]" pageId="36" pageNumber="37" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2879FF892DD81DE80D48FA1F" box="[508,576,1494,1521]" pageId="36" pageNumber="37" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2879FF892E6A1DE90D9EFA1E" box="[590,662,1494,1521]" pageId="36" pageNumber="37">7 juvs</specimenCount>
|
||
(mainly fragmented); same collection data as for holotype but
|
||
<date id="FFBEF4B42879FF892C991DC40E59F9FA" box="[189,337,1530,1556]" pageId="36" pageNumber="37" value="1974-08-09">
|
||
<collectingDate id="EFFA0D5C2879FF892C991DC40E59F9FA" box="[189,337,1530,1556]" pageId="36" pageNumber="37" value="1974-08-09">9 Aug. 1974</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22879FF892D791DC40D0FF9FD" box="[349,519,1529,1556]" pageId="36" pageNumber="37">S.I. Golovatch</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12879FF892E6D1DC40DA0F9FA" box="[585,680,1530,1556]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="36" pageNumber="37">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2879FF852C991E7C0C38FB80" lastPageId="40" lastPageNumber="41" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BBFD2742879FF892C991E7C0E46F9B2" blockId="36.[189,334,1602,1628]" box="[189,334,1602,1628]" pageId="36" pageNumber="37">
|
||
<heading id="D0F765182879FF892C991E7C0E46F9B2" bold="true" box="[189,334,1602,1628]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
|
||
<emphasis id="B9740E662879FF892C991E7C0E46F9B2" bold="true" box="[189,334,1602,1628]" pageId="36" pageNumber="37">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991E4F0D86F8F6" blockId="36.[189,1399,1649,1816]" pageId="36" pageNumber="37">
|
||
SIZE AND NUMBER OF BODY RINGS.
|
||
<typeStatus id="54BB6CD62879FF892E181E4F0DA1F965" box="[572,681,1649,1675]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912879FF892ED71E4F0C4EF962" box="[755,838,1649,1676]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="37" unit="mm" value="25.0">25 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912879FF8929761E4F0FE6F941" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="36" pageNumber="37" unit="mm" value="1.6">1.6 mm</quantity>
|
||
, body with 50 podous rings + 1 apodous ring + telson.
|
||
<typeStatus id="54BB6CD62879FF892FB41EAB0CFFF941" box="[912,1015,1685,1711]" pageId="36" pageNumber="37" type="paratype">Paratype</typeStatus>
|
||
males
|
||
<quantity id="4CF87F912879FF89286B1EAA0BD8F941" box="[1103,1232,1684,1711]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="36" pageNumber="37" unit="mm" value="25.5" valueMax="29.0" valueMin="22.0">22–29 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912879FF892E331E860DAFF93C" box="[535,679,1720,1746]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="36" pageNumber="37" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 mm</quantity>
|
||
, body with 43–52 podous rings + 0–2 apodous rings + telson.
|
||
<typeStatus id="54BB6CD62879FF892C991EE50E2CF91B" box="[189,292,1755,1781]" pageId="36" pageNumber="37" type="paratype">Paratype</typeStatus>
|
||
females
|
||
<quantity id="4CF87F912879FF892DAA1EE50D05F91B" box="[398,525,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="36" pageNumber="37" unit="mm" value="25.0" valueMax="29.0" valueMin="21.0">21–29 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912879FF8928371EE50BACF91B" box="[1043,1188,1755,1781]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="36" pageNumber="37" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 mm</quantity>
|
||
, body with 46–50 podous rings + 1 apodous ring + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991F010DDDF8B7" blockId="36.[189,725,1855,1882]" box="[189,725,1855,1882]" pageId="36" pageNumber="37">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF12879FF892D0C1F010E77F8B7" box="[296,383,1855,1882]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="36" pageNumber="37">Fig. 22</figureCitation>
|
||
). Yellowish white in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742879FF892C991FBF0A7FF80F" blockId="36.[189,1399,1921,2018]" pageId="36" pageNumber="37">
|
||
HEAD (
|
||
<figureCitation id="133BCEF12879FF892D2B1FBF0E74F875" box="[271,380,1921,1947]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="36" pageNumber="37">Figs 22B</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12879FF892DA81FBF0EC9F875" box="[396,449,1921,1947]" captionStart="Fig" captionStartId="38.[189,232,1858,1884]" captionTargetBox="[190,1400,391,1837]" captionTargetId="figure@38.[190,1400,391,1837]" captionTargetPageId="38" captionText="Fig. 23. Leucogeorgia golovatchi sp. nov., paratype ♂ from Tsakhi Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Right gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A, C–D = 0.1 mm; B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4020800" httpUri="https://zenodo.org/record/4020800/files/figure.png" pageId="36" pageNumber="37">23C</figureCitation>
|
||
). Without ommatidia. Frontal setae absent. Labrum with three labral teeth, four supralabral setae and 15 labral setae. Gnathochilarium with rhomboid promentum. Lamellae linguales with 5+5 setae, stipites with 3+3 distolateral and 11+11 medial setae. Antennae
|
||
<quantity id="4CF87F912879FF89287D1FF90BBAF80C" box="[1113,1202,1991,2018]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="36" pageNumber="37" unit="mm" value="2.3">2.3 mm</quantity>
|
||
long in
|
||
<typeStatus id="54BB6CD62879FF8929341FF90A7FF80F" box="[1296,1399,1991,2017]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
</paragraph>
|
||
<caption id="DF7F82FC2878FF882C991F590CF5F805" ID-DOI="http://doi.org/10.5281/zenodo.4020798" ID-Zenodo-Dep="4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="37" pageNumber="38" startId="37.[189,232,1895,1921]" targetBox="[281,1307,265,1868]" targetPageId="37">
|
||
<paragraph id="8BBFD2742878FF882C991F590CF5F805" blockId="37.[189,1399,1893,2027]" pageId="37" pageNumber="38">
|
||
<emphasis id="B9740E662878FF882C991F590E16F86F" bold="true" box="[189,286,1895,1921]" pageId="37" pageNumber="38">Fig. 22.</emphasis>
|
||
<taxonomicName id="4C00A9F72878FF882D0F1F560D5DF86F" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[299,597,1895,1921]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
|
||
<emphasis id="B9740E662878FF882D0F1F560D5DF86F" box="[299,597,1895,1921]" italics="true" pageId="37" pageNumber="38">Leucogeorgia golovatchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2878FF882E461F560DCCF86C" box="[610,708,1896,1922]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype).
|
||
<emphasis id="B9740E662878FF882D591FB40E9AF84A" bold="true" box="[381,402,1930,1956]" pageId="37" pageNumber="38">A</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B9740E662878FF882EB81FB40DB8F84A" bold="true" box="[668,688,1930,1956]" pageId="37" pageNumber="38">B</emphasis>
|
||
. Anterior part of body, lateral view.
|
||
<emphasis id="B9740E662878FF8828751FB40B6EF84A" bold="true" box="[1105,1126,1930,1956]" pageId="37" pageNumber="38">C</emphasis>
|
||
. Posterior part of body, lateral view.
|
||
<emphasis id="B9740E662878FF882D771F900E60F826" bold="true" box="[339,360,1966,1992]" pageId="37" pageNumber="38">D</emphasis>
|
||
. Midbody rings, lateral view.
|
||
<emphasis id="B9740E662878FF882EED1F900DD5F826" bold="true" box="[713,733,1966,1992]" pageId="37" pageNumber="38">E</emphasis>
|
||
. Body ring 2, lateral view.
|
||
<emphasis id="B9740E662878FF88283F1F900B25F826" bold="true" box="[1051,1069,1966,1992]" pageId="37" pageNumber="38">F</emphasis>
|
||
. Body ring 2, anterior view.
|
||
<emphasis id="B9740E662878FF882C991FEE0FDCF804" bold="true" box="[189,212,2000,2026]" pageId="37" pageNumber="38">G</emphasis>
|
||
. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBFD274287BFF8B2C9919350A7EFE82" blockId="38.[189,1399,267,364]" pageId="38" pageNumber="39">male, their length ca 140% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.12 (I), 0.47 (II), 0.41 (III), 0.4 (IV), 0.46 (V), 0.29 (VI), 0.12 (VII) and 0.03 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 3 (II), 3 (III), 2.7 (IV), 2.4 (V), 1.6 (VI) and 0.9 (VII). Antennomeres</paragraph>
|
||
<caption id="DF7F82FC287BFF8B2C991F7C0C58F804" ID-DOI="http://doi.org/10.5281/zenodo.4020800" ID-Zenodo-Dep="4020800" httpUri="https://zenodo.org/record/4020800/files/figure.png" pageId="38" pageNumber="39" startId="38.[189,232,1858,1884]" targetBox="[190,1400,391,1837]" targetPageId="38">
|
||
<paragraph id="8BBFD274287BFF8B2C991F7C0C58F804" blockId="38.[189,1399,1857,2026]" pageId="38" pageNumber="39">
|
||
<emphasis id="B9740E66287BFF8B2C991F7C0E16F8B2" bold="true" box="[189,286,1858,1884]" pageId="38" pageNumber="39">Fig. 23.</emphasis>
|
||
<taxonomicName id="4C00A9F7287BFF8B2D0D1F7D0D5AF8B3" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[297,594,1859,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
|
||
<emphasis id="B9740E66287BFF8B2D0D1F7D0D5AF8B3" box="[297,594,1859,1885]" italics="true" pageId="38" pageNumber="39">Leucogeorgia golovatchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D287BFF8B2E7A1F7D0DB7F8B3" box="[606,703,1859,1885]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype ♂ from Tsakhi Cave (ZMUM).
|
||
<emphasis id="B9740E66287BFF8B28EC1F7C0BD5F8B2" bold="true" box="[1224,1245,1858,1884]" pageId="38" pageNumber="39">A</emphasis>
|
||
. Leg-pair 1, anterior view.
|
||
<emphasis id="B9740E66287BFF8B2D491F580E89F86E" bold="true" box="[365,385,1894,1920]" pageId="38" pageNumber="39">B</emphasis>
|
||
. Right leg 1, lateral view.
|
||
<emphasis id="B9740E66287BFF8B2EF41F580DEDF86E" bold="true" box="[720,741,1894,1920]" pageId="38" pageNumber="39">C</emphasis>
|
||
. Head, ventral view.
|
||
<emphasis id="B9740E66287BFF8B2FCB1F580B0CF86E" bold="true" box="[1007,1028,1894,1920]" pageId="38" pageNumber="39">D</emphasis>
|
||
. Right gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A, C–D = 0.1 mm; B = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7F82FC287AFF8A2C991FB40BADF805" ID-DOI="http://doi.org/10.5281/zenodo.4020802" ID-Zenodo-Dep="4020802" httpUri="https://zenodo.org/record/4020802/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1930,1956]" targetBox="[260,1319,268,1908]" targetPageId="39">
|
||
<paragraph id="8BBFD274287AFF8A2C991FB40BADF805" blockId="39.[189,1399,1929,2027]" pageId="39" pageNumber="40">
|
||
<emphasis id="B9740E66287AFF8A2C991FB40E14F84A" bold="true" box="[189,284,1930,1956]" pageId="39" pageNumber="40">Fig. 24.</emphasis>
|
||
<taxonomicName id="4C00A9F7287AFF8A2D021FB50D46F84A" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[294,590,1930,1956]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
|
||
<emphasis id="B9740E66287AFF8A2D021FB50D46F84A" box="[294,590,1930,1956]" italics="true" pageId="39" pageNumber="40">Leucogeorgia golovatchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D287AFF8A2E7C1FB50DB0F84B" box="[600,696,1931,1957]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, type ♂♂ from Tsakhi Cave (ZMUM).
|
||
<emphasis id="B9740E66287AFF8A28BE1FB40BA7F84A" bold="true" box="[1178,1199,1930,1956]" pageId="39" pageNumber="40">A</emphasis>
|
||
. Holotype, right gonopods, mesal view.
|
||
<emphasis id="B9740E66287AFF8A2DFD1F900EE5F826" bold="true" box="[473,493,1966,1992]" pageId="39" pageNumber="40">B</emphasis>
|
||
. Holotype, left gonopods, lateral view.
|
||
<emphasis id="B9740E66287AFF8A2FF51F930CEEF829" bold="true" box="[977,998,1965,1991]" pageId="39" pageNumber="40">C</emphasis>
|
||
. Paratype, penes, posterior view.
|
||
<emphasis id="B9740E66287AFF8A2C991FEF0FDAF805" bold="true" box="[189,210,2001,2027]" pageId="39" pageNumber="40">D</emphasis>
|
||
. Paratype, leg-pair 1, anterior view.
|
||
<emphasis id="B9740E66287AFF8A2E5A1FEF0D9AF805" bold="true" box="[638,658,2001,2027]" pageId="39" pageNumber="40">E</emphasis>
|
||
. Right leg 1, lateral view. Scale bar: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBFD2742875FF852C9919320C60FEA6" blockId="40.[189,1398,267,328]" pageId="40" pageNumber="41">V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C9919570EDAFE27" blockId="40.[189,1399,360,457]" pageId="40" pageNumber="41">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF12875FF852D7119560EA2FE6D" box="[341,426,360,387]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="40" pageNumber="41">Fig. 22</figureCitation>
|
||
D–F). Entire metazonal area with longitudinal striations. Length of midbody setae ca 5% of vertical diameter of rings. Body ring 2 with strongly developed ventral lobes (
|
||
<figureCitation id="133BCEF12875FF85289A19B20A21FE48" box="[1214,1321,396,422]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="40" pageNumber="41">Fig. 22B</figureCitation>
|
||
, E–F), rounded in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C9919D70BBFFDC8" blockId="40.[189,1398,489,550]" pageId="40" pageNumber="41">
|
||
TELSON (
|
||
<figureCitation id="133BCEF12875FF852D0619D70E85FDED" box="[290,397,489,515]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="40" pageNumber="41">Fig. 22C</figureCitation>
|
||
). Epiproct with a very short and blunt preanal process, covered with dorsal and lateral setae. Paraprocts rounded, with numerous setae. Hypoproct without any modifications.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C991A780D6DFD02" blockId="40.[189,1399,582,748]" pageId="40" pageNumber="41">
|
||
LEGS IN MALES. First pair of legs modified, hook-shaped (
|
||
<figureCitation id="133BCEF12875FF852F561A780CC6FD8E" box="[882,974,582,608]" captionStart="Fig" captionStartId="38.[189,232,1858,1884]" captionTargetBox="[190,1400,391,1837]" captionTargetId="figure@38.[190,1400,391,1837]" captionTargetPageId="38" captionText="Fig. 23. Leucogeorgia golovatchi sp. nov., paratype ♂ from Tsakhi Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Right gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A, C–D = 0.1 mm; B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4020800" httpUri="https://zenodo.org/record/4020800/files/figure.png" pageId="40" pageNumber="41">Figs 23</figureCitation>
|
||
A–B, 24D–E), with three complete podomeres; coxa with one seta; prefemur with 8–10 setae; femur, postfemur and tibiotarsus coalesced; femur with 2–3 setae; postfemur with one seta. Tibiotarsus with a short distal lobe (tarsal remnant). Tip slightly tuberculated. Postfemoral and tibial ventral pads well-developed on anterior legs, then gradually disappearing towards posterior legs.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C991B330BEDFCC8" blockId="40.[189,1253,780,807]" box="[189,1253,780,807]" pageId="40" pageNumber="41">
|
||
PENES (
|
||
<figureCitation id="133BCEF12875FF852D361B320E75FCC9" box="[274,381,780,807]" captionStart="Fig" captionStartId="39.[189,232,1930,1956]" captionTargetBox="[260,1319,268,1908]" captionTargetId="figure@39.[247,1353,250,1912]" captionTargetPageId="39" captionText="Fig. 24. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM). A. Holotype, right gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, penes, posterior view. D. Paratype, leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020802" httpUri="https://zenodo.org/record/4020802/files/figure.png" pageId="40" pageNumber="41">Fig. 24C</figureCitation>
|
||
). In form of a short trapezoid, apically with two small subtriangular lobes.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C991B790B64FC8E" blockId="40.[189,1132,838,865]" box="[189,1132,838,865]" pageId="40" pageNumber="41">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF12875FF852EA71B780DF8FC8E" box="[643,752,838,865]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[281,1307,265,1868]" captionTargetId="figure@37.[281,1307,265,1868]" captionTargetPageId="37" captionText="Fig. 22. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM) (A–D: holotype; E–G: paratype). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Posterior part of body, lateral view. D. Midbody rings, lateral view. E. Body ring 2, lateral view. F. Body ring 2, anterior view. G. Body ring 7, lateral view. Scale bars: A–D = 1 mm; E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4020798" httpUri="https://zenodo.org/record/4020798/files/figure.png" pageId="40" pageNumber="41">Fig. 22G</figureCitation>
|
||
). Small, rounded in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C991BBE0C38FB80" blockId="40.[189,1399,896,1134]" pageId="40" pageNumber="41">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF12875FF852D671BBE0EA6FC75" box="[323,430,896,923]" captionStart="Fig" captionStartId="38.[189,232,1858,1884]" captionTargetBox="[190,1400,391,1837]" captionTargetId="figure@38.[190,1400,391,1837]" captionTargetPageId="38" captionText="Fig. 23. Leucogeorgia golovatchi sp. nov., paratype ♂ from Tsakhi Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Right gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A, C–D = 0.1 mm; B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4020800" httpUri="https://zenodo.org/record/4020800/files/figure.png" pageId="40" pageNumber="41">Figs 23D</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12875FF852D9F1BBE0ED0FC74" box="[443,472,896,922]" captionStart="Fig" captionStartId="39.[189,232,1930,1956]" captionTargetBox="[260,1319,268,1908]" captionTargetId="figure@39.[247,1353,250,1912]" captionTargetPageId="39" captionText="Fig. 24. Leucogeorgia golovatchi sp. nov., type ♂♂ from Tsakhi Cave (ZMUM). A. Holotype, right gonopods, mesal view. B. Holotype, left gonopods, lateral view. C. Paratype, penes, posterior view. D. Paratype, leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4020802" httpUri="https://zenodo.org/record/4020802/files/figure.png" pageId="40" pageNumber="41">24</figureCitation>
|
||
A–B). Promere (p) long and slender, with a flagellum (f); apical part with poorly denticulated margins; basal half with two developed ridges. Mesomere (m) with a slender mesomeral claw (mc) lacking teeth; mesomeral lamella (ml) gradually decreasing in height posteriad, distal margin serrate, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a long tip exceeding posterior part of lamella, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2875FF852C991CB30C89FB3D" box="[189,897,1165,1235]" pageId="40" pageNumber="41" type="distribution">
|
||
<paragraph id="8BBFD2742875FF852C991CB30E5EFB49" blockId="40.[189,897,1165,1235]" box="[189,342,1165,1191]" pageId="40" pageNumber="41">
|
||
<heading id="D0F765182875FF852C991CB30E5EFB49" bold="true" box="[189,342,1165,1191]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
|
||
<emphasis id="B9740E662875FF852C991CB30E5EFB49" bold="true" box="[189,342,1165,1191]" pageId="40" pageNumber="41">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742875FF852C991C870C89FB3D" blockId="40.[189,897,1165,1235]" box="[189,897,1208,1235]" pageId="40" pageNumber="41">
|
||
Known only from its
|
||
<typeStatus id="54BB6CD62875FF852D9E1C870EE5FB3D" box="[442,493,1209,1235]" pageId="40" pageNumber="41">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="133BCEF12875FF852E7F1C860DBAFB3D" box="[603,690,1208,1235]" captionStart="Fig" captionStartId="98.[189,232,1964,1990]" captionTargetBox="[189,1399,1026,1935]" captionTargetId="figure@98.[189,1399,1026,1935]" captionTargetPageId="98" captionText="Fig. 58. Distribution of the southeastern group of the genus Leucogeorgia Verhoeff, 1930 and the species Martvilia parva gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4020877" httpUri="https://zenodo.org/record/4020877/files/figure.png" pageId="40" pageNumber="41">Fig. 58</figureCitation>
|
||
, orange triangle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |