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<document id="50DA8C95BCC1A35DD9A382E9D6B58FCC" ID-DOI="10.3853/j.2201-4349.68.2016.1598" ID-ISSN="2201-4349" ID-Zenodo-Dep="5238225" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629536249708" checkinUser="felipe" docAuthor="Owen, Christopher L. &amp; Moulds, Max S." docDate="2016" docId="039DA814D04BFFBA4AB818D8FE10FB1A" docLanguage="en" docName="RecAustMus.68.4.117-200.pdf" docOrigin="Records of the Australian Museum 68 (4)" docSource="http://dx.doi.org/10.3853/j.2201-4349.68.2016.1598" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Haemopsalta Owen &amp; Moulds 2016, n.gen." docType="treatment" docUuid="B69336DA-794B-4E20-865E-23F691FC2C01" docUuidSource="ZooBank" docVersion="3" lastPageNumber="178" masterDocId="FFA4D06CD077FF874902184FFFD4FF9D" masterDocTitle="Systematics and Phylogeny of the Australian Cicada Genus Pauropsalta Goding and Froggatt, 1904 and Allied Genera (Hemiptera: Cicadidae: Cicadettini)" masterLastPageNumber="200" masterPageNumber="117" pageNumber="177" updateTime="1699077434683" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3744EE5FFF3778F5D065752691FE0BBA">Systematics and Phylogeny of the Australian Cicada Genus Pauropsalta Goding and Froggatt, 1904 and Allied Genera (Hemiptera: Cicadidae: Cicadettini)</mods:title>
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<mods:namePart id="4C7181AD4299C1E6C4558141D851F896">Owen, Christopher L.</mods:namePart>
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<mods:date id="D0F46C58B91267EA856331A63F0E1670">2016</mods:date>
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<paragraph id="8B8B1902D04BFFBB4AB818D8FA9CFF2B" blockId="60.[954,1352,150,182]" box="[954,1352,150,182]" pageId="60" pageNumber="177">
<heading id="D0C3AE6ED04BFFBB4AB818D8FA9CFF2B" box="[954,1352,150,182]" centered="true" fontSize="11" level="2" pageId="60" pageNumber="177" reason="2">
<emphasis id="B940C510D04BFFBB4AB818D8FA9CFF2B" bold="true" box="[954,1352,150,182]" pageId="60" pageNumber="177">
Genus
<taxonomicName id="4C346281D04BFFBB4D2118D9FB37FF28" ID-CoL="8HK89" authority="Owen &amp; Moulds, 2016" authorityName="Owen &amp; Moulds" authorityYear="2016" box="[1059,1251,150,181]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B940C510D04BFFBB4D2118D9FB37FF28" bold="true" box="[1059,1251,150,181]" italics="true" pageId="60" pageNumber="177">Haemopsalta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273786BD04BFFBB4DEF18D8FA9CFF2B" box="[1261,1352,151,182]" pageId="60" pageNumber="177" rank="genus">n.gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32E4A89D04BFFBB4A671886FB95FF64" pageId="60" pageNumber="177" type="reference_group">
<paragraph id="8B8B1902D04BFFBB4A671886FB95FF64" blockId="60.[869,1436,201,249]" pageId="60" pageNumber="177">
<treatmentCitationGroup id="AB243E2CD04BFFBB4A671886FB95FF64" pageId="60" pageNumber="177">
<treatmentCitation id="0A953F13D04BFFBB4A671886FAD7FF42" author="Owen, C &amp; Marshall, K" box="[869,1283,201,223]" page="261" pageId="60" pageNumber="177" year="2015">
<taxonomicName id="4C346281D04BFFBB4A671886FAD7FF42" ID-CoL="8HK89" authority="Owen et al., 2015: 259 - 261" authorityName="Owen" authorityPageNumber="259 - 261" authorityYear="2015" box="[869,1283,201,223]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B940C510D04BFFBB4A671886FC31FF43" box="[869,997,201,222]" italics="true" pageId="60" pageNumber="177">Haemopsalta</emphasis>
<bibRefCitation id="EFA564F3D04BFFBB4AEF1886FAD7FF42" author="Owen, C &amp; Marshall, K" box="[1005,1283,201,223]" pageId="60" pageNumber="177" pagination="258 - 277" refId="ref83466" refString="Owen, C. L., D. C. Marshall, K. B. R. Hill, and C. Simon. 2015. The phylogenetic utility of acetyltransferase (ARD 1) and glutaminyl tRNA synthetase (QtRNA) for reconstructing Cenozoic relationships as exemplified by the large Australian cicada Pauropsalta complex. Molecular phylogenetics and evolution 83: 258 - 277. http: // dx. doi. org / 10.1016 / j. ympev. 2014.07.008" type="journal article" year="2015">
Owen
<emphasis id="B940C510D04BFFBB4D2C1885FBB5FF43" box="[1070,1121,201,223]" italics="true" pageId="60" pageNumber="177">et al.</emphasis>
, 2015: 259–261
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 263, 271, 273, 274,
<emphasis id="B940C510D04BFFBB4AB618ABFBEEFF64" box="[948,1082,228,249]" italics="true" pageId="60" pageNumber="177">nomen nudum</emphasis>
.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C32E4A89D04BFFBB4A451940FCAEFE12" pageId="60" pageNumber="177" type="type_taxon">
<paragraph id="8B8B1902D04BFFBB4A451940FA95FEBA" blockId="60.[839,1345,271,295]" box="[839,1345,271,295]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A451940FC00FEBA" bold="true" box="[839,980,271,295]" pageId="60" pageNumber="177">
<typeStatus id="548FA7A0D04BFFBB4A451940FCABFEBA" box="[839,895,271,295]" pageId="60" pageNumber="177">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C346281D04BFFBB4AE31940FAE9FEBA" ID-CoL="75V7N" authority="Ewart, 1989" authorityName="Ewart" authorityYear="1989" box="[993,1341,271,295]" class="Insecta" family="Cicadidae" genus="Pauropsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="species" species="aktites">
<emphasis id="B940C510D04BFFBB4AE31940FB65FEBA" box="[993,1201,271,295]" italics="true" pageId="60" pageNumber="177">Pauropsalta aktites</emphasis>
<bibRefCitation id="EFA564F3D04BFFBB4DB5195FFAE9FEBA" author="Ewart, A" box="[1207,1341,271,295]" pageId="60" pageNumber="177" pagination="289 - 375" refId="ref82284" refString="Ewart, A. 1989. Revisionary notes on the genus Pauropsalta Goding and Froggatt (Homoptera: Cicadidae) with special reference to Queensland. Memoirs of the Queensland Museum 27 (2): 289 - 375." type="journal article" year="1989">Ewart, 1989</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A451972FCAEFE12" blockId="60.[839,1466,316,399]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A451972FC28FEC9" bold="true" box="[839,1020,316,341]" pageId="60" pageNumber="177">Included species</emphasis>
.
<taxonomicName id="4C346281D04BFFBB4D0A1973FB37FEC8" ID-CoL="75V7N" authority="(Ewart, 1989)" authorityName="Owen &amp; Moulds" authorityYear="2016" baseAuthorityName="Ewart" baseAuthorityYear="1989" box="[1032,1251,316,341]" class="Insecta" family="Cicadidae" genus="Pauropsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="species" species="aktites" status="comb. nov.">
<emphasis id="B940C510D04BFFBB4D0A1973FB99FEC9" box="[1032,1101,316,340]" italics="true" pageId="60" pageNumber="177">aktites</emphasis>
(
<bibRefCitation id="EFA564F3D04BFFBB4D5A1972FB0FFEC8" author="Ewart, A" box="[1112,1243,317,341]" pageId="60" pageNumber="177" pagination="289 - 375" refId="ref82284" refString="Ewart, A. 1989. Revisionary notes on the genus Pauropsalta Goding and Froggatt (Homoptera: Cicadidae) with special reference to Queensland. Memoirs of the Queensland Museum 27 (2): 289 - 375." type="journal article" year="1989">Ewart, 1989</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A273786BD04BFFBB4DEE1973FA90FEC9" box="[1260,1348,316,340]" pageId="60" pageNumber="177" rank="species">n.comb.</taxonomicNameLabel>
;
<taxonomicName id="4C346281D04BFFBB4C491973FA6DFEC9" ID-CoL="3J968" authority="Owen &amp; Moulds, 2016" authorityName="Owen &amp; Moulds" authorityYear="2016" box="[1355,1465,316,340]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="species" species="flammeata" status="sp. nov.">
<emphasis id="B940C510D04BFFBB4C491973FA6DFEC9" box="[1355,1465,316,340]" italics="true" pageId="60" pageNumber="177">flammeata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273786BD04BFFBB4A451915FCA8FEEC" box="[839,892,346,369]" pageId="60" pageNumber="177" rank="species">n.sp.</taxonomicNameLabel>
;
<taxonomicName id="4C346281D04BFFBB4A851915FB39FEEF" ID-CoL="3J96C" authority="(Goding &amp; Froggatt, 1904)" authorityName="Owen &amp; Moulds" authorityYear="2016" baseAuthorityName="Goding &amp; Froggatt" baseAuthorityYear="1904" box="[903,1261,346,370]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="species" species="rubea" status="comb. nov.">
<emphasis id="B940C510D04BFFBB4A851915FC11FEEF" box="[903,965,346,370]" italics="true" pageId="60" pageNumber="177">rubea</emphasis>
(Goding &amp; Froggatt, 1904)
</taxonomicName>
,
<taxonomicNameLabel id="A273786BD04BFFBB4DFA1915FA85FEEF" box="[1272,1361,346,370]" pageId="60" pageNumber="177" rank="species">n.comb.</taxonomicNameLabel>
;
<taxonomicName id="4C346281D04BFFBB4C591914FA6EFEEF" ID-CoL="6KWDR" authority="Owen &amp; Moulds, 2016" authorityName="Owen &amp; Moulds" authorityYear="2016" box="[1371,1466,347,370]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="177" phylum="Arthropoda" rank="species" species="georgina" status="sp. nov.">
<emphasis id="B940C510D04BFFBB4C591914FA6EFEEF" box="[1371,1466,347,370]" italics="true" pageId="60" pageNumber="177">georgina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273786BD04BFFBB4A451937FCAEFE12" box="[839,890,376,399]" pageId="60" pageNumber="177" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C32E4A89D04BFFBB4A4519EDFC66FD8F" pageId="60" pageNumber="177" type="etymology">
<paragraph id="8B8B1902D04BFFBB4A4519EDFC66FD8F" blockId="60.[837,1467,418,567]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A4519EDFC17FE27" bold="true" box="[839,963,418,442]" pageId="60" pageNumber="177">Etymology</emphasis>
. From the Greek root
<emphasis id="B940C510D04BFFBB4DC119EDFAD2FE27" box="[1219,1286,418,442]" italics="true" pageId="60" pageNumber="177">haem-</emphasis>
meaning blood, referring to the dominant red colour of most species, and from psalta, derived from
<emphasis id="B940C510D04BFFBB4D531993FB7EFE69" box="[1105,1194,476,500]" italics="true" pageId="60" pageNumber="177">psaltria,</emphasis>
meaning a female harpist. Feminine.
</paragraph>
</subSubSection>
<subSubSection id="C32E4A89D04BFFBA4D411A52FE10FB1A" lastPageId="61" lastPageNumber="178" pageId="60" pageNumber="177" type="diagnosis">
<paragraph id="8B8B1902D04BFFBB4D411A52FB6AFDAA" blockId="60.[837,1467,418,567]" box="[1091,1214,541,567]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4D411A52FB6AFDAA" bold="true" box="[1091,1214,541,567]" pageId="60" pageNumber="177">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A451A08FB10FC92" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A451A08FC54FDC2" box="[839,896,583,607]" italics="true" pageId="60" pageNumber="177">Head</emphasis>
including eyes about as wide as mesonotum, sometimes a little narrower, sometimes a little wider; supra-antennal plate meeting or nearly meeting eye; postclypeus broadly rounded transversely across ventral midline, in lateral profile angulate between “top” and “sides”; postclypeus in dorsal view tending confluent with anterior margin of head and tending angular in outline when viewed from above.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671B5BFC16FC3F" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671B5BFC66FCB1" box="[869,946,788,812]" italics="true" pageId="60" pageNumber="177">Thorax</emphasis>
. Pronotal collar width at dorsal midline much less than diameter of eyes; paranota confluent with adjoining pronotal sclerites, no mid lateral tooth; cruciform elevation with its dome wider than long; epimeral lobe not reaching operculum.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671BE7FAE7FC22" blockId="60.[837,1468,583,1986]" box="[869,1331,935,959]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671BE7FC4BFC22" box="[869,927,936,959]" italics="true" pageId="60" pageNumber="177">Legs.</emphasis>
Fore leg femoral primary spine erect.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671B8AFA6FFA09" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671B8AFC7AFC41" box="[869,942,965,988]" italics="true" pageId="60" pageNumber="177">Wings.</emphasis>
Hyaline. Fore wings with 8 apical cells; subapical cells absent; ulnar cell 3 angled to radial cell; basal cell long and narrow; costal vein (C) clearly higher than R+Sc; costa parallel-sided to node; pterostigma present; vein CuA only weakly bowed so that cubital cell no wider than medial cell; veins M and CuA completely fused as one before reaching basal cell; vein RA
<subScript id="17B01B47D04BFFBB4D1A1CCEFBF4FB12" attach="left" box="[1048,1056,1153,1167]" fontSize="6" pageId="60" pageNumber="177">1</subScript>
aligned closely with Sc for its length and not diverging in subapical region; vein CuA
<subScript id="17B01B47D04BFFBB4C5B1CD0FAB5FB30" attach="left" box="[1369,1377,1183,1197]" fontSize="6" pageId="60" pageNumber="177">1</subScript>
divided by crossvein m-cu so that proximal portion shortest; veins CuP and 1A fused in part; infuscation absent; wing outer margin developed for its total length, never reduced to be contiguous with ambient vein. Hind wings with 6 apical cells; infuscation at distal end of vein 2A spread on wing margin; width of 1st cubital cell at distal end at least twice that of 2nd cubital cell; anal lobe broad with vein 3A curved, long, separated from wing margin; veins RP and M fused basally.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671DD5FA6DF994" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671DD5FC2BFA2F" box="[869,1023,1434,1458]" italics="true" pageId="60" pageNumber="177">Male opercula</emphasis>
more or less reaching margin of tympanal cavity, directed towards distomedial margin of tympanal cavity, apically broadly rounded, clearly not meeting, clearly raised above level of tympanal cavity on its outer half or so.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671E40FC3AF901" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671E40FBDEF9BA" box="[869,1034,1551,1575]" italics="true" pageId="60" pageNumber="177">Male abdomen</emphasis>
in cross-section with sides of tergites straight or weakly convex, epipleurites reflexed ventrally from junction with tergites; tergites 2–7 all similar in size (2 and 3 not considerably larger); sternites III–VII in crosssection convex.
</paragraph>
<paragraph id="8B8B1902D04BFFBB4A671EEDFA7EF969" blockId="60.[837,1468,583,1986]" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671EEDFC15F927" box="[869,961,1698,1722]" italics="true" pageId="60" pageNumber="177">Timbals.</emphasis>
Timbal covers absent; timbal ribs irregular in size and spaced with prominent intermediate short ribs; basal dome very large; timbals not extended below wing bases.
</paragraph>
<paragraph id="8B8B1902D04BFFBA4A671EB5FECEFD67" blockId="60.[837,1468,583,1986]" lastBlockId="61.[119,750,151,821]" lastPageId="61" lastPageNumber="178" pageId="60" pageNumber="177">
<emphasis id="B940C510D04BFFBB4A671EB5FC2BF88F" box="[869,1023,1786,1810]" italics="true" pageId="60" pageNumber="177">Male genitalia.</emphasis>
Pygofer with distal shoulders not developed; upper pygofer lobe shape in lateral view with basal portion of dorsal margin strongly angled to axis of pygofer, very broad basally and tapering to a broad apex; dorsal beak present and a part of chitinized pygofer; pygofer basal lobe peg-like with rounded distal end; pygofer secondary basal lobe with outer face ridged longitudinally, lobe-like and domed, in lateral view not or moderately projecting and broadly rounded, in ventral view well developed with rounded apex that is sometimes twotiered. Uncus in dorsal view broadly rounded, flat or depressed along dorsal midline. Claspers in lateral view extending no more than a little beyond margin of pygofer, claw-like with minimum or no cavity below; claspers extreme basal region projecting forward as a broad rounded lobe.Aedeagus weakly depressed on dorsal midline; with basal plate in lateral view undulated with basal portion of basal plate directed forwards away from thecal shaft; in dorsal view tending to be Y-shaped; ventral rib completely fused with basal plate; junction between theca and basal plate with a functional “hinge” that is small and substantially compressed between theca and basal plate in lateral view; thecal shaft straight or curved in a gentle arc; pseudoparameres arising independently at the base; endotheca exposed, ridged, much of surface weakly sclerotized and in part translucent; endotheca shaft parallel-sided in apical region; endothecal ventral support absent; thecal apex entirely chitinized, thecal subapical cerci absent; flabellum absent; conjunctival claws absent; vesica retractable, vesical opening apical on theca.
</paragraph>
<paragraph id="8B8B1902D04AFFBA49951AB0FEF9FCA8" blockId="61.[119,750,151,821]" pageId="61" pageNumber="178">
The male genitalia of
<taxonomicName id="4C346281D04AFFBA48801AB0FDC4FC8A" box="[386,528,767,791]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B940C510D04AFFBA48801AB0FDC4FC8A" box="[386,528,767,791]" italics="true" pageId="61" pageNumber="178">Haemopsalta</emphasis>
</taxonomicName>
show little variation between species.
</paragraph>
<paragraph id="8B8B1902D04AFFBA497B1B18FE51FCF2" blockId="61.[121,389,855,879]" box="[121,389,855,879]" pageId="61" pageNumber="178">
<emphasis id="B940C510D04AFFBA497B1B18FE51FCF2" bold="true" box="[121,389,855,879]" pageId="61" pageNumber="178">Distinguishing features</emphasis>
</paragraph>
<paragraph id="8B8B1902D04AFFBA497B1BCAFE10FB1A" blockId="61.[119,749,901,1159]" pageId="61" pageNumber="178">
<taxonomicName id="4C346281D04AFFBA497B1BCAFED3FC00" box="[121,263,901,925]" class="Insecta" family="Cicadidae" genus="Haemopsalta" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B940C510D04AFFBA497B1BCAFED3FC00" box="[121,263,901,925]" italics="true" pageId="61" pageNumber="178">Haemopsalta</emphasis>
</taxonomicName>
can be separated from all other genera by the following combination of characters: fore wing veins M and CuA with their stems meeting the basal cell completely fused as one; claspers in lateral view extending no more than a little beyond the margin of the pygofer, claw-like with minimum or no cavity below; uncus in dorsal view triangular and depressed along the dorsal midline; endotheca trumpetshaped in apical region, the apical margin continuously serrated laterally and ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>
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