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<document id="7667CB446D085A3CF4AEE01B36D536BB" ID-DOI="10.5281/zenodo.186138" ID-GBIF-Dataset="8cf8a401-bc91-4a30-99b8-27492f4f254e" ID-ISSN="1175-5326" ID-Zenodo-Dep="186138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461046376021" checkinUser="plazi" docAuthor="De, Jeferson L., Lindquist, Evert E. &amp; De, Gilberto J." docDate="2009" docId="039687DDFFCE8D5DFF76B22FFEA27926" docLanguage="en" docName="zt02024p032.pdf" docOrigin="Zootaxa 2024" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protogamasellus sigillophorus De, Lindquist &amp; De, 2009, sp. nov." docType="treatment" docVersion="10" lastPageNumber="29" masterDocId="FFAFFFA5FFD78D41FFE1B44BFFF27D22" masterDocTitle="Edaphic ascid mites (Acari: Mesostigmata: Ascidae) from the state of São Paulo, Brazil, with description of five new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="26" updateTime="1698577253990" updateUser="plazi">
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<mods:title id="E032D67929FDDEEE2139E777772D63F5">Edaphic ascid mites (Acari: Mesostigmata: Ascidae) from the state of São Paulo, Brazil, with description of five new species</mods:title>
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<mods:namePart id="F0F436B4416CEECCBAA174A30E52602C">De, Jeferson L.</mods:namePart>
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<mods:namePart id="C4C7161E82F9F611337E21E7206C2016">Lindquist, Evert E.</mods:namePart>
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<mods:namePart id="32CDF5D3767CCA88B20C7D13905CCEF0">De, Gilberto J.</mods:namePart>
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<mods:date id="B889FF22E8DB7D824BA24599F49EC78B">2009</mods:date>
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<treatment id="039687DDFFCE8D5DFF76B22FFEA27926" ID-DOI="http://doi.org/10.5281/zenodo.6222966" ID-GBIF-Taxon="119610861" ID-Zenodo-Dep="6222966" LSID="urn:lsid:plazi:treatment:039687DDFFCE8D5DFF76B22FFEA27926" httpUri="http://treatment.plazi.org/id/039687DDFFCE8D5DFF76B22FFEA27926" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="C3256540FFCE8D58FF76B22FFEDF7B82" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8B8036CBFFCE8D58FF76B22FFD987B5F" blockId="25.[151,618,1635,1696]" box="[151,618,1635,1662]" pageId="25" pageNumber="26">
<heading id="D0C881A7FFCE8D58FF76B22FFD987B5F" bold="true" box="[151,618,1635,1662]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B94BEAD9FFCE8D58FF76B22FFD987B5F" bold="true" box="[151,618,1635,1662]" pageId="25" pageNumber="26">
<taxonomicName id="4C3F4D48FFCE8D58FF76B22FFDF47B5C" ID-CoL="9F2DL" box="[151,518,1636,1662]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sigillophorus" status="sp. nov.">
<emphasis id="B94BEAD9FFCE8D58FF76B22FFDF47B5C" bold="true" box="[151,518,1636,1662]" italics="true" pageId="25" pageNumber="26">Protogamasellus sigillophorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857A2FFCE8D58FDECB228FD987B5F" box="[525,618,1635,1661]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8B8036CBFFCE8D58FF76B2CDFEDF7B82" blockId="25.[151,618,1635,1696]" box="[151,301,1670,1696]" pageId="25" pageNumber="26">
(
<figureCitation id="13042A4EFFCE8D58FF7EB2CDFED77B82" box="[159,293,1670,1696]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="25" pageNumber="26">Figs 6067</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256540FFCE8D5AFF76B298FB607FED" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8B8036CBFFCE8D58FF76B298FDBB7AF4" blockId="25.[151,1437,1747,2006]" pageId="25" pageNumber="26">
<emphasis id="B94BEAD9FFCE8D58FF76B298FEEE7BCF" bold="true" box="[151,284,1747,1773]" pageId="25" pageNumber="26">Diagnosis:</emphasis>
Adult females of this species are similar to those of
<taxonomicName id="4C3F4D48FFCE8D58FC7AB29FFF127A36" authority="Athias-Henriot, 1961" authorityName="Athias-Henriot" authorityYear="1961" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="massula">
<emphasis id="B94BEAD9FFCE8D58FC7AB29FFB3C7BCF" box="[923,1230,1748,1773]" italics="true" pageId="25" pageNumber="26">Protogamasellus massula</emphasis>
(
<bibRefCitation id="EFAE4B3AFFCE8D58FB3EB29FFF2A7A36" author="Athias-Henriot" pageId="25" pageNumber="30" refString="Athias-Henriot, C. (1961) Mesostigmates (Urop. Excl.) edaphiques mediterraneens (Acaromorpha, Anactinotrichida). Acarologia, 3, 381 - 509." type="journal article" year="1961">Athias-Henriot, 1961</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C3F4D48FFCE8D58FEC1B2B0FD587A36" authority="Genis et al., 1967" authorityName="Genis et al." authorityYear="1967" box="[288,682,1786,1812]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="scuticalis">
<emphasis id="B94BEAD9FFCE8D58FEC1B2B0FEC27A36" box="[288,304,1787,1812]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="B94BEAD9FFCE8D58FEA3B2B0FE4E7A36" box="[322,444,1787,1812]" italics="true" pageId="25" pageNumber="26">scuticalis</emphasis>
<bibRefCitation id="EFAE4B3AFFCE8D58FE25B2B1FD587A36" author="Genis" box="[452,682,1786,1812]" pageId="25" pageNumber="31" refString="Genis, N. De L., Loots, G. C. &amp; Ryke, P. A. J. (1967) The genus Protogamasellus Karg (Acari) with descriptions of new species and subspecies from the Ethiopian Region. Journal of Natural History, 1, 337 - 53." type="journal article" year="1967">
Genis
<emphasis id="B94BEAD9FFCE8D58FDF6B2B0FDA37A36" box="[535,593,1787,1812]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
.,
<quantity id="4CC79B2EFFCE8D58FD8AB2B1FD587A36" box="[619,682,1786,1812]" metricMagnitude="1" metricUnit="m" metricValue="4.99618" pageId="25" pageNumber="26" unit="in" value="1967.0">1967</quantity>
</bibRefCitation>
</taxonomicName>
in having opisthogastric setae
<emphasis id="B94BEAD9FFCE8D58FBDBB2B0FB9F7A36" box="[1082,1133,1787,1812]" italics="true" pageId="25" pageNumber="26">ZV1</emphasis>
inserted behind, and in longitudinal alignment with,
<emphasis id="B94BEAD9FFCE8D58FE0AB36AFDEB7A18" box="[491,537,1825,1850]" italics="true" pageId="25" pageNumber="26">JV1</emphasis>
on soft cuticle and in having a pyriform ventri-anal shield that is longer than wide, with 3 pairs of opisthogastric setae (
<emphasis id="B94BEAD9FFCE8D58FD68B303FD457A43" box="[649,695,1864,1889]" italics="true" pageId="25" pageNumber="26">JV2</emphasis>
,
<emphasis id="B94BEAD9FFCE8D58FD24B303FD017A43" box="[709,755,1864,1889]" italics="true" pageId="25" pageNumber="26">JV3</emphasis>
and
<emphasis id="B94BEAD9FFCE8D58FCCDB303FCAB7A43" box="[812,857,1864,1889]" italics="true" pageId="25" pageNumber="26">JV5</emphasis>
) in addition to the circum-anal setae. They differ from the latter two in a variety of aspects, particularly the opisthonotal shield having setae
<emphasis id="B94BEAD9FFCE8D58FB26B324FB107AAA" box="[1223,1250,1903,1928]" italics="true" pageId="25" pageNumber="26">J4</emphasis>
relatively short (i.e., only 1.3 instead of 2.0 longer than
<emphasis id="B94BEAD9FFCE8D58FD95B3DEFD7C7A8C" box="[628,654,1941,1966]" italics="true" pageId="25" pageNumber="26">J5</emphasis>
), and the simply bidentate movable digit, with the proximal tooth much larger than the subapical tooth.
</paragraph>
<caption id="DF406643FFCD8D5BFF76B3D9FF387ACC" httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="26" pageNumber="27" targetBox="[189,1410,186,1907]" targetPageId="26">
<paragraph id="8B8036CBFFCD8D5BFF76B3D9FF387ACC" blockId="26.[151,1436,1938,2030]" pageId="26" pageNumber="27">
<emphasis id="B94BEAD9FFCD8D5BFF76B3D9FE967A88" bold="true" box="[151,356,1938,1962]" pageId="26" pageNumber="27">FIGURES 6067.</emphasis>
<taxonomicName id="4C3F4D48FFCD8D5BFE8DB3D9FD337A8B" box="[364,705,1938,1961]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sigillophorus" status="sp. nov.">
<emphasis id="B94BEAD9FFCD8D5BFE8DB3D9FD337A8B" box="[364,705,1938,1961]" italics="true" pageId="26" pageNumber="27">Protogamasellus sigillophorus</emphasis>
</taxonomicName>
<emphasis id="B94BEAD9FFCD8D5BFD29B3D9FCF57A88" bold="true" box="[712,775,1938,1962]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A27857A2FFCD8D5BFD29B3D9FCF57A88" box="[712,775,1938,1962]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral
<taxonomicName id="4C3F4D48FFCD8D5BFDF9B3FFFD6B7AE9" box="[536,665,1972,1995]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="26" pageNumber="28" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
(64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67).
</paragraph>
</caption>
<paragraph id="8B8036CBFFCC8D5AFF27B4D3FB607FED" blockId="27.[151,1437,152,2034]" pageId="27" pageNumber="28">
<emphasis id="B94BEAD9FFCC8D5AFF27B4D3FEA97D93" box="[198,347,152,177]" italics="true" pageId="27" pageNumber="28">Adult female</emphasis>
.
<emphasis id="B94BEAD9FFCC8D5AFE8BB4D3FDDC7D93" box="[362,558,152,177]" italics="true" pageId="27" pageNumber="28">Dorsal idiosoma</emphasis>
(
<figureCitation id="13042A4EFFCC8D5AFDDFB4D3FD657D90" box="[574,663,152,178]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="27" pageNumber="28">Fig. 60</figureCitation>
): Dorsal shields collectively 270305 µm long and unornamented other than podonotal shield having an uninterrupted transverse line at level of setal bases
<emphasis id="B94BEAD9FFCC8D5AFB2AB4F4FB147DFA" box="[1227,1254,191,216]" italics="true" pageId="27" pageNumber="28">z6</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFAC0B4F4FAC97DFA" box="[1313,1339,191,216]" italics="true" pageId="27" pageNumber="28">s6</emphasis>
, behind which shield desclerotized, and opisthonotal shield having an uninterrupted transverse line at level of setal bases
<emphasis id="B94BEAD9FFCC8D5AFF3EB547FF097C07" box="[223,251,268,293]" italics="true" pageId="27" pageNumber="28">J1</emphasis>
, followed by a couple of short transverse lines laterally near bases of setae
<emphasis id="B94BEAD9FFCC8D5AFB9FB547FB6F7C07" box="[1150,1181,268,293]" italics="true" pageId="27" pageNumber="28">Z1</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFB39B547FB077C07" box="[1240,1269,268,293]" italics="true" pageId="27" pageNumber="28">S1</emphasis>
and a median sigillar scar between bases of setae
<emphasis id="B94BEAD9FFCC8D5AFDD5B579FDA37C69" box="[564,593,306,331]" italics="true" pageId="27" pageNumber="28">J1</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFD69B579FD567C69" box="[648,676,306,331]" italics="true" pageId="27" pageNumber="28">J2</emphasis>
. Podonotal shield 130148 µm long, 98102 µm wide at level of setae
<emphasis id="B94BEAD9FFCC8D5AFF39B512FF1D7C50" box="[216,239,345,370]" italics="true" pageId="27" pageNumber="28">j5</emphasis>
, with 17 pairs of setae, opisthonotal shield 138152 µm long, 6568 µm wide at level of setae
<emphasis id="B94BEAD9FFCC8D5AFAA4B512FA937C50" box="[1349,1377,345,370]" italics="true" pageId="27" pageNumber="28">J2</emphasis>
, (
<emphasis id="B94BEAD9FFCC8D5AFA98B512FA667C50" box="[1401,1428,345,370]" italics="true" pageId="27" pageNumber="28">ca</emphasis>
. 2.12.2 times as long as wide) with 15 pairs of setae; 12 pairs of setae on lateral soft cuticle, including
<emphasis id="B94BEAD9FFCC8D5AFAB1B5CBFA997CBB" box="[1360,1387,384,409]" italics="true" pageId="27" pageNumber="28">s2</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFF76B5ECFF2B7CE2" box="[151,217,422,448]" italics="true" pageId="27" pageNumber="28">r2-r5</emphasis>
anteriorly (
<emphasis id="B94BEAD9FFCC8D5AFE8CB5ECFE7B7CE2" box="[365,393,423,448]" italics="true" pageId="27" pageNumber="28">r6</emphasis>
absent) and
<emphasis id="B94BEAD9FFCC8D5AFDC8B5ECFD857CE2" box="[553,631,422,448]" italics="true" pageId="27" pageNumber="28">R1-R6</emphasis>
and one submarginal
<emphasis id="B94BEAD9FFCC8D5AFC6DB5ECFC467CE2" box="[908,948,423,448]" italics="true" pageId="27" pageNumber="28">UR</emphasis>
seta posteriorly. Podonotal shield with vertical setae
<emphasis id="B94BEAD9FFCC8D5AFEDCB586FEA67CC4" box="[317,340,461,486]" italics="true" pageId="27" pageNumber="28">j1</emphasis>
prominent, stouter and longer (2527 µm) than other setae (1116 µm) which shorter than successive distances between their bases; on opisthonotal shield
<emphasis id="B94BEAD9FFCC8D5AFC79B5BFFC467F2F" box="[920,948,500,525]" italics="true" pageId="27" pageNumber="28">J3</emphasis>
(1214 µm)
<emphasis id="B94BEAD9FFCC8D5AFBB3B5BFFB9C7F2F" box="[1106,1134,500,525]" italics="true" pageId="27" pageNumber="28">ca</emphasis>
. 0.30.4 times as long as alveolar interval
<emphasis id="B94BEAD9FFCC8D5AFE82B651FE547F11" box="[355,422,538,564]" italics="true" pageId="27" pageNumber="28">J3-J4</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFE54B651FE207F11" box="[437,466,538,563]" italics="true" pageId="27" pageNumber="28">J5</emphasis>
(910 µm) shorter than other members of
<emphasis id="B94BEAD9FFCC8D5AFC37B651FC117F11" box="[982,995,538,563]" italics="true" pageId="27" pageNumber="28">J</emphasis>
-series (1216 µm),
<emphasis id="B94BEAD9FFCC8D5AFB39B651FAD37F11" box="[1240,1313,538,564]" italics="true" pageId="27" pageNumber="28">Z1-Z4</emphasis>
similar in length (1518 µm) and 0.60.7 as long as alveolar interval to next seta in series,
<emphasis id="B94BEAD9FFCC8D5AFB93B60AFB607F78" box="[1138,1170,577,602]" italics="true" pageId="27" pageNumber="28">Z5</emphasis>
conspicuously longer (2830 µm);
<emphasis id="B94BEAD9FFCC8D5AFED8B623FE727FA3" box="[313,384,616,642]" italics="true" pageId="27" pageNumber="28">S1-S4</emphasis>
similar in length (1112 µm) and 0.40.5 as long as interval to next seta in series,
<emphasis id="B94BEAD9FFCC8D5AFA9FB623FA6E7FA3" box="[1406,1436,616,641]" italics="true" pageId="27" pageNumber="28">S5</emphasis>
slightly longer (14 µm); all setae smooth except
<emphasis id="B94BEAD9FFCC8D5AFD36B6C4FD017F8A" box="[727,755,655,680]" italics="true" pageId="27" pageNumber="28">J4</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFCE3B6C4FCD37F8A" box="[770,801,655,680]" italics="true" pageId="27" pageNumber="28">Z5</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFCBDB6C4FC8A7F8A" box="[860,888,655,680]" italics="true" pageId="27" pageNumber="28">S5</emphasis>
sparsely barbed. Setae
<emphasis id="B94BEAD9FFCC8D5AFB71B6C4FB557F8A" box="[1168,1191,655,680]" italics="true" pageId="27" pageNumber="28">j2</emphasis>
transversely aligned with
<emphasis id="B94BEAD9FFCC8D5AFF33B6FEFF1B7FEC" box="[210,233,693,718]" italics="true" pageId="27" pageNumber="28">j1</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFEC3B6FEFECE7FEC" box="[290,316,693,718]" italics="true" pageId="27" pageNumber="28">z1</emphasis>
. Setae on lateral soft cuticle short (1013 µm, except
<emphasis id="B94BEAD9FFCC8D5AFC57B6FEFC3D7FEC" box="[950,975,693,718]" italics="true" pageId="27" pageNumber="28">r3</emphasis>
15 µm), simple.
</paragraph>
</subSubSection>
<subSubSection id="C3256540FFCC8D5DFF24B697FE227F79" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8B8036CBFFCC8D5AFF24B697FD4278F6" blockId="27.[151,1437,152,2034]" pageId="27" pageNumber="28">
<emphasis id="B94BEAD9FFCC8D5AFF24B697FE6A7FD7" box="[197,408,732,757]" italics="true" pageId="27" pageNumber="28">Ve n t ra l idiosoma</emphasis>
(
<figureCitation id="13042A4EFFCC8D5AFE4BB697FDFB7FD4" box="[426,521,732,758]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="27" pageNumber="28">Fig. 61</figureCitation>
): Tritosternum normal in shape, with trapezoidal base and slender, pilose laciniae fused along basal one-fifth of their length (6070 µm). Pre-sternal area weakly sclerotized, microtuberculate, without platelets. Sternal shield
<emphasis id="B94BEAD9FFCC8D5AFD15B762FCE27E60" box="[756,784,809,834]" italics="true" pageId="27" pageNumber="28">ca</emphasis>
. 75 µm in median length by 45 µm wide at narrowest width between coxae II, with 3 pairs of setae and 3 pairs of poroids; shield with anterior margin deeply emarginated medially, faintly lineated along lateral margins, smooth over rest of surface, and with posterior margin gently concave. Setae
<emphasis id="B94BEAD9FFCC8D5AFDE6B7D6FDD97E94" box="[519,555,925,950]" italics="true" pageId="27" pageNumber="28">st4</emphasis>
inserted on soft cuticle closely behind postero-lateral corners of sternal shield,
<emphasis id="B94BEAD9FFCC8D5AFF0DB78FFEFC7EFF" box="[236,270,964,989]" italics="true" pageId="27" pageNumber="28">st4</emphasis>
and genital setae
<emphasis id="B94BEAD9FFCC8D5AFE03B78FFDF67EFF" box="[482,516,964,989]" italics="true" pageId="27" pageNumber="28">st5</emphasis>
(1012 µm) slightly shorter than sternal setae
<emphasis id="B94BEAD9FFCC8D5AFBCDB78FFB8B7EFF" box="[1068,1145,964,990]" italics="true" pageId="27" pageNumber="28">st1-st3</emphasis>
(1416 µm). Endopodal plates formed as weak strips alongside coxae IIIIV, not discernible in some specimens. Genital shield smooth, not widened at level of insertions of genital setae, with posterior margin truncate, and hyaline anterior margin broadly rounded, reaching posterior margin of sternal shield. Post-genital groove without discernible sigillar platelets. Metapodal plates hardly discernible as short, narrow strips (length 10 µm) laterad
<emphasis id="B94BEAD9FFCC8D5AFA84B014FA66795A" box="[1381,1428,1119,1144]" italics="true" pageId="27" pageNumber="28">JV1</emphasis>
. Ventri-anal shield pyriform,
<emphasis id="B94BEAD9FFCC8D5AFE0EB0CEFDF979BC" box="[495,523,1157,1182]" italics="true" pageId="27" pageNumber="28">ca</emphasis>
. 1.2 as long (8797 µm) as wide (6575 µm), with anterior margin arched; with 2 or 3 transverse lines arched anteriorly between bases of setae
<emphasis id="B94BEAD9FFCC8D5AFC3EB0E7FBFC79E7" box="[991,1038,1196,1221]" italics="true" pageId="27" pageNumber="28">JV2</emphasis>
and para-anals (lines sometimes interrupted medially), and sparsely punctate on post-anal region; shield with 3 pairs of ventral setae (
<emphasis id="B94BEAD9FFCC8D5AFA87B098FA6679CE" box="[1382,1428,1235,1260]" italics="true" pageId="27" pageNumber="28">JV2</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFF76B0B2FF377830" box="[151,197,1273,1298]" italics="true" pageId="27" pageNumber="28">JV3</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFF35B0B2FEF07830" box="[212,258,1273,1298]" italics="true" pageId="27" pageNumber="28">JV5</emphasis>
) plus 3 circum-anal setae; post-anal seta and
<emphasis id="B94BEAD9FFCC8D5AFCC6B0B2FCA77830" box="[807,853,1273,1298]" italics="true" pageId="27" pageNumber="28">JV5</emphasis>
sparsely barbed and considerably longer (3336 µm and 2123 µm, respectively) than smooth para-anals (1416 µm) and
<emphasis id="B94BEAD9FFCC8D5AFBF3B16BFB8A781B" box="[1042,1144,1312,1338]" italics="true" pageId="27" pageNumber="28">JV2-JV3</emphasis>
(1114 µm); para-anals inserted at level of anterior margin of anus. Three pairs of ventral setae (
<emphasis id="B94BEAD9FFCC8D5AFBC5B10CFBA17842" box="[1060,1107,1351,1376]" italics="true" pageId="27" pageNumber="28">JV1</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFB85B10CFB657842" box="[1124,1175,1351,1376]" italics="true" pageId="27" pageNumber="28">ZV1</emphasis>
,
<emphasis id="B94BEAD9FFCC8D5AFB49B10CFB2A7842" box="[1192,1240,1351,1376]" italics="true" pageId="27" pageNumber="28">JV4</emphasis>
) on soft cuticle anterior and lateral to ventri-anal shield;
<emphasis id="B94BEAD9FFCC8D5AFD9BB126FD5978A4" box="[634,683,1389,1414]" italics="true" pageId="27" pageNumber="28">ZV1</emphasis>
nearly aligned vertically behind
<emphasis id="B94BEAD9FFCC8D5AFBD1B126FBAC78A4" box="[1072,1118,1389,1414]" italics="true" pageId="27" pageNumber="28">
JV
<quantity id="4CC79B2EFFCC8D5AFBB2B126FBAC78A4" box="[1107,1118,1389,1414]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="28" unit="in" value="1.0">1</quantity>
</emphasis>
in front of anterior margin of shield. Exopodal plate strips reduced to triangular fragments between coxae II and III, and between coxae III and IV, not discernible in some specimens.
</paragraph>
<paragraph id="8B8036CBFFCC8D5AFF24B1AAFC7C7B4D" blockId="27.[151,1437,152,2034]" pageId="27" pageNumber="28">
<emphasis id="B94BEAD9FFCC8D5AFF24B1AAFD9278D8" box="[197,608,1505,1530]" italics="true" pageId="27" pageNumber="28">Peritrematal shield and peritreme</emphasis>
(
<figureCitation id="13042A4EFFCC8D5AFD90B1AAFD0878D9" box="[625,762,1505,1531]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="27" pageNumber="28">Fig. 6061</figureCitation>
): Peritrematal shield interrupted between levels of setal alveoli
<emphasis id="B94BEAD9FFCC8D5AFF13B243FEFF7B03" box="[242,269,1544,1569]" italics="true" pageId="27" pageNumber="28">r3</emphasis>
and
<emphasis id="B94BEAD9FFCC8D5AFEA8B243FE967B03" box="[329,356,1544,1569]" italics="true" pageId="27" pageNumber="28">s2</emphasis>
, with separate triangular anterior piece abutting but not fused to dorsal shield at level of paravertical poroids and setae
<emphasis id="B94BEAD9FFCC8D5AFDEAB264FDD77B6A" box="[523,549,1583,1608]" italics="true" pageId="27" pageNumber="28">z1</emphasis>
; peritrematal shield narrow posteriorly, not fused to exopodal fragments alongside coxae IIIV; peritremes extending to bases of setae
<emphasis id="B94BEAD9FFCC8D5AFC8CB21EFC747B4C" box="[877,902,1621,1646]" italics="true" pageId="27" pageNumber="28">r3</emphasis>
.
</paragraph>
<paragraph id="8B8036CBFFCC8D5AFF24B237FBA47BB4" blockId="27.[151,1437,152,2034]" box="[197,1110,1660,1686]" pageId="27" pageNumber="28">
<emphasis id="B94BEAD9FFCC8D5AFF24B237FE167BB7" box="[197,484,1660,1685]" italics="true" pageId="27" pageNumber="28">Spermathecal apparatus</emphasis>
: Unsclerotized, not discernible in specimens at hand.
</paragraph>
<paragraph id="8B8036CBFFCC8D5AFF24B2E8FCA67AE9" blockId="27.[151,1437,152,2034]" pageId="27" pageNumber="28">
<taxonomicName id="4C3F4D48FFCC8D5AFF24B2E8FEAC7B9E" box="[197,350,1699,1724]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="27" pageNumber="28" phylum="Nematoda" rank="genus">
<emphasis id="B94BEAD9FFCC8D5AFF24B2E8FEAC7B9E" box="[197,350,1699,1724]" italics="true" pageId="27" pageNumber="28">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13042A4EFFCC8D5AFE8FB2E8FE0F7B9F" box="[366,509,1699,1725]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="27" pageNumber="28">Figs 6264</figureCitation>
): Anterior margin of tectum denticulate, irregularly triangular. Fixed digit of chelicera with
<emphasis id="B94BEAD9FFCC8D5AFEA2B282FE2B7BC0" box="[323,473,1737,1762]" italics="true" pageId="27" pageNumber="28">pilus dentilis</emphasis>
, with large offset subapical tooth and row of 1113 evenly sized and spaced teeth; movable digit 35 µm long, with 2 well separated, slightly retrorse teeth, the proximal larger than subapical one. Deutosternum with 7 narrow, connected rows of denticles, anteriormost 2 or 3 rows with single median denticle, other rows with 24 denticles; corniculus stout, reaching to mid-level of palpfemur; internal mala slender, with lateral margin slightly fimbriate, hardly widened basally, extending to tips of corniculus. Hypostomatic setae
<emphasis id="B94BEAD9FFCC8D5AFE69B3C0FE547A86" box="[392,422,1931,1956]" italics="true" pageId="27" pageNumber="28">h1</emphasis>
(20 µm) similar in slender form to, but longer than,
<emphasis id="B94BEAD9FFCC8D5AFBFDB3C0FBCB7A86" box="[1052,1081,1931,1956]" italics="true" pageId="27" pageNumber="28">h2</emphasis>
(15 µm). Palptrochanter with apical seta slightly stout, rigid, inserted on low prominence.
</paragraph>
<paragraph id="8B8036CBFFCC8D5DFF24B393FAF37F16" blockId="27.[151,1437,152,2034]" lastBlockId="28.[151,1437,152,1028]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B94BEAD9FFCC8D5AFF24B393FF0E7AD3" box="[197,252,2008,2033]" italics="true" pageId="27" pageNumber="28">Legs</emphasis>
(
<figureCitation id="13042A4EFFCC8D5AFEEAB393FE337AD0" box="[267,449,2008,2034]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="27" pageNumber="28">Figs 61, 6567</figureCitation>
): Leg I (268275 µm) nearly as long as combined length of the 2 dorsal shields; legs II (163168 µm), III (132140 µm) and IV (200212 µm) considerably shorter, with III notably smaller than II. Coxa I with cluster of 24 small spine-like processes on dorsal distal margin, and sparsely lineate ventrally; coxa II with 2 lines posteroventrally; coxae IIIIV unornamented (processes and lines of coxae I and II not shown in
<figureCitation id="13042A4EFFCB8D5DFEE8B547FE927C04" box="[265,352,268,294]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="28" pageNumber="29">Fig. 61</figureCitation>
). Legs IIV with normally developed pretarsi (length
<emphasis id="B94BEAD9FFCB8D5DFC3DB547FC057C07" box="[988,1015,268,293]" italics="true" pageId="28" pageNumber="29">ca</emphasis>
. 5 µm on I, 811 µm on IIIV) and claws. Tarsi IIIV with apical setal processes
<emphasis id="B94BEAD9FFCB8D5DFD52B579FD1B7C69" box="[691,745,306,332]" italics="true" pageId="28" pageNumber="29">ad -1</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFD19B579FCDC7C69" box="[760,814,306,332]" italics="true" pageId="28" pageNumber="29">pd -1</emphasis>
inconspicuous, thin (length ca. 10 µm). Leg IV with tarsus (4750 µm)
<emphasis id="B94BEAD9FFCB8D5DFE97B512FE607C50" box="[374,402,345,370]" italics="true" pageId="28" pageNumber="29">ca</emphasis>
. 2.3 as long as tibia (3538 µm); tarsus IV with seta
<emphasis id="B94BEAD9FFCB8D5DFBE5B512FBC87C50" box="[1028,1082,345,371]" italics="true" pageId="28" pageNumber="29">ad -2</emphasis>
(also denoted as
<emphasis id="B94BEAD9FFCB8D5DFAE7B512FAD87C50" box="[1286,1322,345,370]" italics="true" pageId="28" pageNumber="29">md</emphasis>
) (22 µm) and
<emphasis id="B94BEAD9FFCB8D5DFF28B5CBFF0D7CBB" box="[201,255,384,410]" italics="true" pageId="28" pageNumber="29">ad -3</emphasis>
(2730 µm) erect and longer than other setae on same segment (
<figureCitation id="13042A4EFFCB8D5DFC1DB5CBFBA77CB8" box="[1020,1109,384,410]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="28" pageNumber="29">Fig. 67</figureCitation>
). Setation of femora of legs I-II-III-IV, respectively,
<date id="FF81100BFFCB8D5DFE53B5EDFDDB7CE2" box="[434,553,422,448]" pageId="28" pageNumber="29" value="1912-11-06" valueMax="1912-11-06" valueMin="1912-11-06">12-11-6-6</date>
; that of genua,
<date id="FF81100BFFCB8D5DFD3CB5EDFCA27CE2" box="[733,848,422,448]" pageId="28" pageNumber="29" value="1913-11-08" valueMax="1913-11-08" valueMin="1913-11-08">13-11-8-8</date>
(
<emphasis id="B94BEAD9FFCB8D5DFC80B5ECFC647CE2" box="[865,918,422,448]" italics="true" pageId="28" pageNumber="29">pv -1</emphasis>
absent on genu III,
<emphasis id="B94BEAD9FFCB8D5DFB9FB5ECFB5C7CE2" box="[1150,1198,422,448]" italics="true" pageId="28" pageNumber="29">pl -1</emphasis>
absent on genu IV); that of tibiae,
<date id="FF81100BFFCB8D5DFEDFB586FE477CC5" box="[318,437,461,487]" pageId="28" pageNumber="29" value="1913-08-10" valueMax="1913-09-10" valueMin="1913-08-10">13-10-8-9</date>
(
<emphasis id="B94BEAD9FFCB8D5DFE27B586FE047CC4" box="[454,502,461,487]" italics="true" pageId="28" pageNumber="29">pl -2</emphasis>
absent on tibia IV); leg setae smooth, mostly slender, except
<emphasis id="B94BEAD9FFCB8D5DFB39B586FAFC7CC4" box="[1240,1294,461,487]" italics="true" pageId="28" pageNumber="29">ad -3</emphasis>
on femur I,
<emphasis id="B94BEAD9FFCB8D5DFF76B5BFFF3C7F2F" box="[151,206,500,526]" italics="true" pageId="28" pageNumber="29">ad -2</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFF3FB5BFFEE77F2F" box="[222,277,500,526]" italics="true" pageId="28" pageNumber="29">ad -3</emphasis>
on femur II, and
<emphasis id="B94BEAD9FFCB8D5DFE0CB5BFFDD67F2F" box="[493,548,500,526]" italics="true" pageId="28" pageNumber="29">ad -2</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFDD4B5BFFD9E7F2F" box="[565,620,500,526]" italics="true" pageId="28" pageNumber="29">pd -1</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFD9DB5BFFD5F7F2F" box="[636,685,500,526]" italics="true" pageId="28" pageNumber="29">pl -1</emphasis>
of femur IV short, spine-like, and
<emphasis id="B94BEAD9FFCB8D5DFBB9B5BFFB7C7F2F" box="[1112,1166,500,526]" italics="true" pageId="28" pageNumber="29">av -2</emphasis>
, to lesser extent other ventral setae on tarsus II (
<figureCitation id="13042A4EFFCB8D5DFE24B651FDE97F16" box="[453,539,538,564]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="28" pageNumber="29">Fig. 65</figureCitation>
) and
<emphasis id="B94BEAD9FFCB8D5DFDBDB651FD627F11" box="[604,656,538,564]" italics="true" pageId="28" pageNumber="29">av -2</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFD7EB651FD207F11" box="[671,722,538,564]" italics="true" pageId="28" pageNumber="29">pv -2</emphasis>
,
<emphasis id="B94BEAD9FFCB8D5DFD00B651FCF67F11" box="[737,772,538,563]" italics="true" pageId="28" pageNumber="29">mv</emphasis>
on tarsus IV thickened, spinelike (
<figureCitation id="13042A4EFFCB8D5DFB7DB651FB067F16" box="[1180,1268,538,564]" captionStart="FIGURES 60 67" captionStartId="26.[151,273,1938,1962]" captionTargetBox="[189,1410,186,1907]" captionTargetId="figure@26.[171,1426,172,1936]" captionTargetPageId="26" captionText="FIGURES 60 67. Protogamasellus sigillophorus n. sp. Adult female, dorsal idiosoma (60), ventral idiosoma (61), tectum (62), chelicera (63), ventral gnathosoma (64), ventral tarsus II (65), dorsal tarsus II (66) and tibia and tarsus IV (67)." httpUri="https://zenodo.org/record/186147/files/figure.png" pageId="28" pageNumber="29">Fig. 67</figureCitation>
).
</paragraph>
<paragraph id="8B8036CBFFCB8D5DFF24B60AFE227F79" blockId="28.[151,1437,152,1028]" box="[197,464,577,603]" pageId="28" pageNumber="29">
<emphasis id="B94BEAD9FFCB8D5DFF24B60AFEB77F78" box="[197,325,577,602]" italics="true" pageId="28" pageNumber="29">Adult male</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3256540FFCB8D5DFF24B62CFCE67FD4" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B8036CBFFCB8D5DFF24B62CFCE67FD4" blockId="28.[151,1437,152,1028]" pageId="28" pageNumber="29">
<emphasis id="B94BEAD9FFCB8D5DFF24B62CFE397FA3" bold="true" box="[197,459,615,641]" pageId="28" pageNumber="29">Material examined:</emphasis>
<typeStatus id="54848869FFCB8D5DFE35B623FDBA7FA0" box="[468,584,616,642]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
female collected,
<date id="FF81100BFFCB8D5DFCD3B623FC517FA0" box="[818,931,616,642]" pageId="28" pageNumber="29" value="1998-10-02">2.x.1998</date>
, from soil of a corn field, ESALQ-USP, Piracicaba, State of São Paulo, col. J.L. de C. Mineiro, deposited at ESALQ-USP. Three
<typeStatus id="54848869FFCB8D5DFB2EB6C5FAC57F8A" box="[1231,1335,654,680]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
females including two with same data as
<typeStatus id="54848869FFCB8D5DFDC0B6FEFD7B7FED" box="[545,649,693,719]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
and one with same data except that collected on
<date id="FF81100BFFCB8D5DFB39B6FEFA917FED" box="[1240,1379,693,719]" pageId="28" pageNumber="29" value="1998-12-21">21.xii.1998</date>
, two deposited at
<collectionCode id="ED2EAE0EFFCB8D5DFECAB697FE7B7FD4" box="[299,393,732,758]" pageId="28" pageNumber="29">ESALQ</collectionCode>
/
<collectionCode id="ED2EAE0EFFCB8D5DFE73B697FE357FD4" box="[402,455,732,758]" httpUri="http://grbio.org/cool/hb24-jpsb" name="University of the South Pacific" pageId="28" pageNumber="29">USP</collectionCode>
and one deposited at
<collectionCode id="ED2EAE0EFFCB8D5DFD27B697FCE27FD4" box="[710,784,732,758]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="28" pageNumber="29">CNCI</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256540FFCB8D5DFF27B749FC627E95" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B8036CBFFCB8D5DFF27B749FC627E95" blockId="28.[151,1437,152,1028]" pageId="28" pageNumber="29">
<emphasis id="B94BEAD9FFCB8D5DFF27B749FEB07E3E" bold="true" box="[198,322,770,796]" pageId="28" pageNumber="29">Remarks:</emphasis>
Our description of this species as distinct from
<taxonomicName id="4C3F4D48FFCB8D5DFC63B749FBF37E39" box="[898,1025,770,796]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="massula">
<emphasis id="B94BEAD9FFCB8D5DFC63B749FC637E39" box="[898,913,770,795]" italics="true" pageId="28" pageNumber="29">P</emphasis>
.
<emphasis id="B94BEAD9FFCB8D5DFC40B749FBF37E39" box="[929,1025,770,795]" italics="true" pageId="28" pageNumber="29">massula</emphasis>
</taxonomicName>
is based in part on notes taken by one of us (EEL, unpublished, 1972) during a study of Athias-Henriots
<typeStatus id="54848869FFCB8D5DFBFDB762FBA07E61" box="[1052,1106,809,835]" pageId="28" pageNumber="29">type</typeStatus>
material as well as on the subsequent redescription of that species by
<bibRefCitation id="EFAE4B3AFFCB8D5DFD7EB71BFCB17E48" author="Evans" box="[671,835,848,874]" pageId="28" pageNumber="30" refString="Evans, G. O. (1982) Observations of the genus Protogamasellus Karg (Acari: Mesostigmata) with a description of a new species. Acarologia, 23, 303 - 313." type="journal article" year="1982">Evans (1982)</bibRefCitation>
. The setal deficiencies noted for genua IIIIV and tibia IV in the above description are typical for the leg chaetotaxy of the genus
<taxonomicName id="4C3F4D48FFCB8D5DFBA3B73CFAFA7EB2" box="[1090,1288,887,912]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAD9FFCB8D5DFBA3B73CFAFA7EB2" box="[1090,1288,887,912]" italics="true" pageId="28" pageNumber="29">Protogamasellus</emphasis>
</taxonomicName>
as indicated by
<bibRefCitation id="EFAE4B3AFFCB8D5DFF5DB7D6FE1C7E95" author="Lindquist" box="[188,494,925,951]" pageId="28" pageNumber="32" refString="Lindquist, E. E. &amp; Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist &amp; Evans (1965)</bibRefCitation>
and further noted by
<bibRefCitation id="EFAE4B3AFFCB8D5DFD0AB7D6FC7E7E95" author="Evans" box="[747,908,925,951]" pageId="28" pageNumber="30" refString="Evans, G. O. (1982) Observations of the genus Protogamasellus Karg (Acari: Mesostigmata) with a description of a new species. Acarologia, 23, 303 - 313." type="journal article" year="1982">Evans (1982)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256540FFCB8D5DFF27B788FEA27926" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B8036CBFFCB8D5DFF27B788FEA27926" blockId="28.[151,1437,152,1028]" pageId="28" pageNumber="29">
<emphasis id="B94BEAD9FFCB8D5DFF27B788FEA57EFF" bold="true" box="[198,343,963,989]" pageId="28" pageNumber="29">Etymology:</emphasis>
The name
<taxonomicName id="4C3F4D48FFCB8D5DFE01B78FFD897EFF" box="[480,635,964,989]" class="Arachnida" family="Ascidae" genus="Protogamasellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sigillophorus">
<emphasis id="B94BEAD9FFCB8D5DFE01B78FFD897EFF" box="[480,635,964,989]" italics="true" pageId="28" pageNumber="29">sigillophorus</emphasis>
</taxonomicName>
refers to the presence of median sigillar scar between the bases of setae
<emphasis id="B94BEAD9FFCB8D5DFF38B7A1FF077921" box="[217,245,1002,1027]" italics="true" pageId="28" pageNumber="29">J1</emphasis>
and
<emphasis id="B94BEAD9FFCB8D5DFECCB7A1FEBB7921" box="[301,329,1002,1027]" italics="true" pageId="28" pageNumber="29">J2</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>