397 lines
45 KiB
XML
397 lines
45 KiB
XML
<document id="10B16B1259DC94160BCADF8838701158" ID-DOI="10.5281/zenodo.293364" ID-GBIF-Dataset="45dbc0b6-9ae8-4970-984e-cfe34dcae5f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="293364" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460246409096" checkinUser="plazi" docAuthor="Frolov, Andrey" docDate="2010" docId="039687DBFFE3FFF0FE906FEBFAF5FCBE" docLanguage="en" docName="zt02531p068.pdf" docOrigin="Zootaxa 2531" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Madecorphnus cuccodoroi Frolov, sp. n." docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFAFFFA3FFE3FFF4FE07696CFF8DFFB2" masterDocTitle="A new species of scarab beetle, Madecorphnus cuccodoroi (Coleoptera: Scarabaeidae: Orphninae), from northern Madagascar" masterLastPageNumber="68" masterPageNumber="64" pageNumber="64" updateTime="1698241956307" updateUser="plazi">
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<mods:titleInfo id="774A6DB04DE965115C856EE75221D60E">
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<mods:title id="E18FA778150E09ED62A14704DD7255F0">A new species of scarab beetle, Madecorphnus cuccodoroi (Coleoptera: Scarabaeidae: Orphninae), from northern Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="E268FF7483484276F22D8CBD36F0F4D7" type="personal">
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<mods:roleTerm id="7C5FC7AC36050CFA23D590BDBCF850C3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6538059B82E3B68E2419D4BD41F41FB0">Frolov, Andrey</mods:namePart>
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<mods:typeOfResource id="F371B6EB85C47965CF9038942221E1EA">text</mods:typeOfResource>
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<mods:title id="DAB37FFAAE837FF63E771BC67B46A769">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CF46C4B2CF1E34E178763F6837AFA192">
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<mods:date id="2221ADA01A97B4429731979CCF52CA22">2010</mods:date>
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<mods:detail id="F1A36CFD15C2AABC87E65717A961740D" type="volume">
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<mods:number id="15DF7AB543E97869740E9EABF6AD7756">2531</mods:number>
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<mods:start id="CC4996D862F18457EAABA94D66A6F1D3">64</mods:start>
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<mods:end id="23397AF756FA38F4EAB677DFCB627A9A">68</mods:end>
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<mods:classification id="D92568ACAA78D91F0BC7B9E1ECC6FCFD">journal article</mods:classification>
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<mods:identifier id="B8EC4C208EDFC57BF05DC014C84C31A0" type="DOI">10.5281/zenodo.293364</mods:identifier>
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<mods:identifier id="E8030D7026D6567DA3C06EA7314C5A6C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="681093162A2B9120B35B12746F52E438" type="Zenodo-Dep">293364</mods:identifier>
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<treatment id="039687DBFFE3FFF0FE906FEBFAF5FCBE" ID-DOI="http://doi.org/10.5281/zenodo.3505135" ID-GBIF-Taxon="119390558" ID-Zenodo-Dep="3505135" LSID="urn:lsid:zoobank.org:act:CF051797-A4EE-4172-A537-8731D86D92D1" httpUri="http://treatment.plazi.org/id/039687DBFFE3FFF0FE906FEBFAF5FCBE" lastPageId="4" lastPageNumber="68" pageId="0" pageNumber="64">
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<subSubSection id="C3256546FFE3FFF4FE906FEBFEB2F976" pageId="0" pageNumber="64" type="nomenclature">
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<paragraph id="8B8036CDFFE3FFF4FE906FEBFD0EF913" blockId="0.[151,643,1671,1732]" box="[151,643,1671,1697]" pageId="0" pageNumber="64">
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<heading id="D0C881A1FFE3FFF4FE906FEBFD0EF913" bold="true" box="[151,643,1671,1697]" fontSize="11" level="1" pageId="0" pageNumber="64" reason="1">
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<emphasis id="B94BEADFFFE3FFF4FE906FEBFD0EF913" bold="true" box="[151,643,1671,1697]" pageId="0" pageNumber="64">
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<taxonomicName id="4C3F4D4EFFE3FFF4FE906FEBFDBEF913" authority="Frolov" authorityName="Frolov" box="[151,563,1671,1697]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="64" phylum="Arthropoda" rank="species" species="cuccodoroi" status="sp. nov.">
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<emphasis id="B94BEADFFFE3FFF4FE906FEBFE57F913" bold="true" box="[151,474,1671,1697]" italics="true" pageId="0" pageNumber="64">Madecorphnus cuccodoroi</emphasis>
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Frolov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27857A4FFE3FFF4FC466FEBFD0EF913" box="[577,643,1671,1697]" pageId="0" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8036CDFFE3FFF4FE906FC6FEB2F976" blockId="0.[151,643,1671,1732]" box="[151,319,1706,1732]" pageId="0" pageNumber="64">
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<figureCitation id="13042A48FFE3FFF4FE906FC6FEA9F976" box="[151,292,1706,1732]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 8" captionStartId-0="1.[153,271,1985,2009]" captionStartId-1="2.[153,271,1626,1650]" captionTargetBox-0="[192,1390,1192,1949]" captionTargetBox-1="[185,1397,201,1594]" captionTargetId-0="figure@1.[189,1397,1192,1960]" captionTargetId-1="figure@2.[184,1404,194,1605]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 2. Madecorphnus cuccodoroi Frolov dorsal habitus: (1) holotype, male, (2) paratype, female." captionText-1="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri-0="https://zenodo.org/record/294098/files/figure.png" httpUri-1="https://zenodo.org/record/294099/files/figure.png" pageId="0" pageNumber="64">Figs. 1–4, 6</figureCitation>
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,
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<figureCitation id="13042A48FFE3FFF4FF376FC6FEB2F976" box="[304,319,1706,1732]" captionStart="FIGURE 9. D" captionStartId="3.[151,256,1909,1933]" captionTargetBox="[177,1413,351,1886]" captionTargetId="figure@3.[169,1417,343,1888]" captionTargetPageId="3" captionText="FIGURE 9. D istribution map of Madecorphnus cuccodoroi Frolov, M. perinetensis Frolov, M. montreuili Frolov, M. falcatus Paulian, and M. peyrierasi Frolov." httpUri="https://zenodo.org/record/293367/files/figure.png" pageId="0" pageNumber="64">9</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3256546FFE3FFF4FE906F86FB1DF856" pageId="0" pageNumber="64" type="materials_examined">
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<paragraph id="8B8036CDFFE3FFF4FE906F86FB1DF856" blockId="0.[151,1436,1770,2020]" pageId="0" pageNumber="64">
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<emphasis id="B94BEADFFFE3FFF4FE906F86FEDEF8B6" bold="true" box="[151,339,1770,1796]" pageId="0" pageNumber="64">
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<typeStatus id="5484886FFFE3FFF4FE906F86FF5BF8B6" box="[151,214,1770,1796]" pageId="0" pageNumber="64">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="5484886FFFE3FFF4FF5B6F86FE5DF8B6" box="[348,464,1770,1796]" pageId="0" pageNumber="64" type="holotype">Holotype</typeStatus>
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||
, male: North
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<collectingCountry id="F328765DFFE3FFF4FC706F86FC86F8B6" box="[631,779,1770,1796]" name="Madagascar" pageId="0" pageNumber="64">Madagascar</collectingCountry>
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||
, Amber Mountain National Park,
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||
NE camp site,
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<geoCoordinate id="EE0B500AFFE3FFF4FE906E7CFE92F898" box="[151,287,1808,1834]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.5525">12°33'09''S</geoCoordinate>
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,
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<geoCoordinate id="EE0B500AFFE3FFF4FF2A6E7CFE37F898" box="[301,442,1808,1834]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.161114">49°09'40''N</geoCoordinate>
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,
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<quantity id="4CC79B28FFE3FFF4FFC06E7CFDABF898" box="[455,550,1808,1834]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="0" pageNumber="64" unit="m" value="1300.0">1300 m</quantity>
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.
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<date id="FF81100DFFE3FFF4FC356E7CFD3CF898" box="[562,689,1808,1834]" pageId="0" pageNumber="64" value="2003-02-28">28.II.2003</date>
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, G. Cuccodoro leg. (
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<collectionCode id="ED2EAE08FFE3FFF4FDA16E7CFB85F898" box="[934,1032,1808,1834]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
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).
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<typeStatus id="5484886FFFE3FFF4FA1B6E7CFB1EF898" box="[1052,1171,1808,1834]" pageId="0" pageNumber="64" type="paratype">Paratypes</typeStatus>
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,
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<specimenCount id="9D39FD44FFE3FFF4FA986E7CFABFF898" box="[1183,1330,1808,1834]" pageId="0" pageNumber="64" type="generic">7 specimens</specimenCount>
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with the same data as the
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||
<typeStatus id="5484886FFFE3FFF4FF5C6E59FE4CF8FD" box="[347,449,1845,1871]" pageId="0" pageNumber="64" type="holotype">holotype</typeStatus>
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||
except with the exact locality and date as follows:
|
||
<quantity id="4CC79B28FFE3FFF4FA146E59FBEEF8FD" box="[1043,1123,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="64" unit="km" value="2.5">2,5 km</quantity>
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SE camp site,
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<geoCoordinate id="EE0B500AFFE3FFF4FB166E59FA1BF8FD" box="[1297,1430,1845,1871]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.5199995">12°31'12''S</geoCoordinate>
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,
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<geoCoordinate id="EE0B500AFFE3FFF4FE906E36FEAEF8C6" box="[151,291,1882,1908]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.195">49°11'42''N</geoCoordinate>
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,
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<quantity id="4CC79B28FFE3FFF4FF366E36FE02F8C6" box="[305,399,1882,1908]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="0" pageNumber="64" unit="m" value="1050.0">1050 m</quantity>
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.
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<date id="FF81100DFFE3FFF4FF9C6E36FDAEF8C6" box="[411,547,1882,1908]" pageId="0" pageNumber="64" value="2003-03-11">11.III.2003</date>
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,
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<specimenCount id="9D39FD44FFE3FFF4FC376E36FD0CF8C6" box="[560,641,1882,1908]" pageId="0" pageNumber="64" type="male">1 male</specimenCount>
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(
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<collectionCode id="ED2EAE08FFE3FFF4FC956E36FD79F8C6" box="[658,756,1882,1908]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
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);
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<quantity id="4CC79B28FFE3FFF4FD0F6E36FCC8F8C6" box="[776,837,1882,1908]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="64" unit="km" value="1.0">1 km</quantity>
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||
NW camp site,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FA0E6E36FB1CF8C6" box="[1033,1169,1882,1908]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.531944">12°31'55''S</geoCoordinate>
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,
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<geoCoordinate id="EE0B500AFFE3FFF4FA986E36FAA0F8C6" box="[1183,1325,1882,1908]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.175556">49°10'32''N</geoCoordinate>
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,
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||
<quantity id="4CC79B28FFE3FFF4FB3D6E36FA14F8C6" box="[1338,1433,1882,1908]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="64" unit="m" value="1000.0">1000 m</quantity>
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||
.
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||
<date id="FF81100DFFE3FFF4FE906EECFE99F828" box="[151,276,1920,1946]" pageId="0" pageNumber="64" value="2003-03-02">2.III.2003</date>
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||
,
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||
<specimenCount id="9D39FD44FFE3FFF4FF256EECFEF8F828" box="[290,373,1920,1946]" pageId="0" pageNumber="64" type="male">1 male</specimenCount>
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||
(
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||
<collectionCode id="ED2EAE08FFE3FFF4FF816EECFE64F828" box="[390,489,1920,1946]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
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);
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||
<quantity id="4CC79B28FFE3FFF4FFF86EECFDB0F828" box="[511,573,1920,1946]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="64" unit="km" value="3.0">3 km</quantity>
|
||
NE camp site,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FCFB6EECFC05F828" box="[764,904,1920,1946]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.541667">12°32'30''S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FD906EECFBA4F828" box="[919,1065,1920,1946]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.167225">49°10'02''N</geoCoordinate>
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,
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<quantity id="4CC79B28FFE3FFF4FA316EECFB1BF828" box="[1078,1174,1920,1946]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="0" pageNumber="64" unit="m" value="1250.0">1250 m</quantity>
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||
.
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||
<date id="FF81100DFFE3FFF4FAA46EECFABDF828" box="[1187,1328,1920,1946]" pageId="0" pageNumber="64" value="2003-03-15">15.III.2003</date>
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||
,
|
||
<specimenCount id="9D39FD44FFE3FFF4FB3A6EECFA11F828" box="[1341,1436,1920,1946]" pageId="0" pageNumber="64" type="male">2 males</specimenCount>
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||
(
|
||
<collectionCode id="ED2EAE08FFE3FFF4FE996EC9FF74F80D" box="[158,249,1957,1983]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
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||
,
|
||
<collectionCode id="ED2EAE08FFE3FFF4FF026EC9FEB6F80D" box="[261,315,1957,1983]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="0" pageNumber="64">ZIN</collectionCode>
|
||
);
|
||
<quantity id="4CC79B28FFE3FFF4FF486EC9FE2DF80D" box="[335,416,1957,1983]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="64" unit="km" value="2.5">2,5 km</quantity>
|
||
SE camp site,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FC486EC9FD59F80D" box="[591,724,1957,1983]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.520833">12°31'15''S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FCE66EC9FCE6F80D" box="[737,875,1957,1983]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.194447">49°11'40''N</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D39FD44FFE3FFF4FD706EC9FC45F80D" box="[887,968,1957,1983]" pageId="0" pageNumber="64" type="generic">
|
||
<quantity id="4CC79B28FFE3FFF4FD706EC9FC49F80D" box="[887,964,1957,1983]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="0" pageNumber="64" unit="m" value="950.0">950 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
<date id="FF81100DFFE3FFF4FDC86EC9FBDBF80D" box="[975,1110,1957,1983]" pageId="0" pageNumber="64" value="2003-03-03">03.III.2003</date>
|
||
,
|
||
<specimenCount id="9D39FD44FFE3FFF4FA656EC9FB4AF80D" box="[1122,1223,1957,1983]" pageId="0" pageNumber="64" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED2EAE08FFE3FFF4FAD16EC9FABBF80D" box="[1238,1334,1957,1983]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
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);
|
||
<quantity id="4CC79B28FFE3FFF4FB4C6EC9FA11F80D" box="[1355,1436,1957,1983]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="64" unit="km" value="0.5">0,5 km</quantity>
|
||
NE camp site,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FF446EA6FE44F856" box="[323,457,1994,2020]" direction="south" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="-12.530556">12°31'50''S</geoCoordinate>
|
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,
|
||
<geoCoordinate id="EE0B500AFFE3FFF4FFD16EA6FDECF856" box="[470,609,1994,2020]" direction="north" orientation="latitude" pageId="0" pageNumber="64" precision="15" value="49.17361">49°10'25''N</geoCoordinate>
|
||
,
|
||
<quantity id="4CC79B28FFE3FFF4FC6A6EA6FD45F856" box="[621,712,1994,2020]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="64" unit="m" value="1000.0">1000 m</quantity>
|
||
.
|
||
<date id="FF81100DFFE3FFF4FCD36EA6FCD7F856" box="[724,858,1994,2020]" pageId="0" pageNumber="64" value="2003-03-01">01.III.2003</date>
|
||
,
|
||
<specimenCount id="9D39FD44FFE3FFF4FD616EA6FC55F856" box="[870,984,1994,2020]" pageId="0" pageNumber="64" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED2EAE08FFE3FFF4FDE06EA6FBCFF856" box="[999,1090,1994,2020]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="64">MHNG</collectionCode>
|
||
,
|
||
<collectionCode id="ED2EAE08FFE3FFF4FA4A6EA6FB0EF856" box="[1101,1155,1994,2020]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="0" pageNumber="64">ZIN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256546FFE2FFF5FEC269FBFE88FBCE" pageId="1" pageNumber="65" type="description">
|
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<paragraph id="8B8036CDFFE2FFF5FEC269FBFE01FF65" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">
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<emphasis id="B94BEADFFFE2FFF5FEC269FBFED2FF03" bold="true" box="[197,351,151,177]" pageId="1" pageNumber="65">Description.</emphasis>
|
||
<typeStatus id="5484886FFFE2FFF5FF6F69F4FE57FF00" box="[360,474,152,178]" pageId="1" pageNumber="65" type="holotype">Holotype</typeStatus>
|
||
, male (
|
||
<figureCitation id="13042A48FFE2FFF5FC3669F4FDF5FF00" box="[561,632,152,178]" captionStart="FIGURES 1 – 2" captionStartId="1.[153,271,1985,2009]" captionTargetBox="[192,1390,1192,1949]" captionTargetId="figure@1.[189,1397,1192,1960]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Madecorphnus cuccodoroi Frolov dorsal habitus: (1) holotype, male, (2) paratype, female." httpUri="https://zenodo.org/record/294098/files/figure.png" pageId="1" pageNumber="65">Fig. 1</figureCitation>
|
||
). Body length 5.5 mm. Color uniformly dark brown, almost black, legs somewhat paler.
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC2698EFB69FE20" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">Right mandible slightly longer than left, without tooth behind apex. Labrum trapezoidal, with slightly rounded sides, length about 1/6 width (in dorsal view). Clypeus very slightly asymmetrical, apically obtuse, with 2 long and 2 shorter setae on the apical margin. Genae very small, not protruding past eyes. Canthus and frontal suture indistinct. Clypeus slightly depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than 4 times their diameter.</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC268F1FCD4FD95" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">Pronotum approximately 1.5 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum finer than on head, almost indistinct. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle.</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26B5EFC58FDFE" blockId="1.[151,1437,151,1148]" box="[197,981,562,588]" pageId="1" pageNumber="65">Scutellum triangular, angulate apically, about 1/12 length of elytra.</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26B34FE24FD50" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">Elytra convex, with distinct humeral and apical, widest at basal third. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra sparsely punctate with relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border from scutellum to humeral callus. Wings fully developed.</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26B81FB70FCE0" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Middle and posterior legs similar in shape to each other. Longer tibial spur shorter than 2 basal tarsomeres in middle legs and as long as 2 tarsomeres in posterior legs.</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26A31FAAAFC70" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">
|
||
Parameres with small teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (
|
||
<figureCitation id="13042A48FFE2FFF5FF326AEEFEF2FC2E" box="[309,383,898,924]" captionStart="FIGURES 3 – 8" captionStartId="2.[153,271,1626,1650]" captionTargetBox="[185,1397,201,1594]" captionTargetId="figure@2.[184,1404,194,1605]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="1" pageNumber="65">Fig. 3</figureCitation>
|
||
). Internal sac with 2 long sclerites and a large number of small spinules; some spinules more robust and form two indistinctly separated clusters in apical part (
|
||
<figureCitation id="13042A48FFE2FFF5FDDD6AC4FBACFC70" box="[986,1057,936,962]" captionStart="FIGURES 3 – 8" captionStartId="2.[153,271,1626,1650]" captionTargetBox="[185,1397,201,1594]" captionTargetId="figure@2.[184,1404,194,1605]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="1" pageNumber="65">Fig. 6</figureCitation>
|
||
, indicated by arrows).
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26AA1FE3FFBBE" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">
|
||
<emphasis id="B94BEADFFFE2FFF5FEC26AA1FEADFC55" bold="true" box="[197,288,973,999]" pageId="1" pageNumber="65">Female</emphasis>
|
||
(
|
||
<figureCitation id="13042A48FFE2FFF5FF286AA1FEFBFC55" box="[303,374,973,999]" captionStart="FIGURES 1 – 2" captionStartId="1.[153,271,1985,2009]" captionTargetBox="[192,1390,1192,1949]" captionTargetId="figure@1.[189,1397,1192,1960]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Madecorphnus cuccodoroi Frolov dorsal habitus: (1) holotype, male, (2) paratype, female." httpUri="https://zenodo.org/record/294098/files/figure.png" pageId="1" pageNumber="65">Fig. 2</figureCitation>
|
||
) differs from male in having apical spur of protibiae that is slightly longer and more robust than apical seta in male.
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE2FFF5FEC26D7BFE88FBCE" blockId="1.[151,1437,151,1148]" pageId="1" pageNumber="65">
|
||
<emphasis id="B94BEADFFFE2FFF5FEC26D7BFEC1FB83" bold="true" box="[197,332,1047,1073]" pageId="1" pageNumber="65">Variability</emphasis>
|
||
. Body length of the
|
||
<typeStatus id="5484886FFFE2FFF5FC406D74FD37FB80" box="[583,698,1048,1074]" pageId="1" pageNumber="65" type="paratype">paratypes</typeStatus>
|
||
varies from 5.0 to 5.5 mm. Left and right mandibles of male
|
||
<typeStatus id="5484886FFFE2FFF5FE906D51FE8AFBE5" box="[151,263,1085,1111]" pageId="1" pageNumber="65" type="paratype">paratypes</typeStatus>
|
||
are almost equal in length and similar to those in females. Otherwise the
|
||
<typeStatus id="5484886FFFE2FFF5FA676D51FB5DFBE5" box="[1120,1232,1085,1111]" pageId="1" pageNumber="65" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="5484886FFFE2FFF5FE906D0EFE8FFBCE" box="[151,258,1122,1148]" pageId="1" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF406645FFE2FFF5FE9E6EADFA9CF86B" box="[153,1297,1985,2009]" httpUri="https://zenodo.org/record/294098/files/figure.png" pageId="1" pageNumber="65" targetBox="[192,1390,1192,1949]" targetPageId="1">
|
||
<paragraph id="8B8036CDFFE2FFF5FE9E6EADFA9CF86B" blockId="1.[153,1297,1985,2009]" box="[153,1297,1985,2009]" pageId="1" pageNumber="65">
|
||
<emphasis id="B94BEADFFFE2FFF5FE9E6EADFEC9F86B" bold="true" box="[153,324,1985,2009]" pageId="1" pageNumber="65">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C3F4D4EFFE2FFF5FF4C6EAEFD3BF86B" authority="Frolov" authorityName="Frolov" box="[331,694,1986,2009]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="cuccodoroi">
|
||
<emphasis id="B94BEADFFFE2FFF5FF4C6EAEFDE4F86B" box="[331,617,1986,2009]" italics="true" pageId="1" pageNumber="65">Madecorphnus cuccodoroi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
dorsal habitus: (1) holotype, male, (2) paratype, female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF406645FFE1FFF6FE9E6F36FEE0F903" httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="2" pageNumber="66" targetBox="[185,1397,201,1594]" targetPageId="2">
|
||
<paragraph id="8B8036CDFFE1FFF6FE9E6F36FEE0F903" blockId="2.[151,1436,1626,1713]" pageId="2" pageNumber="66">
|
||
<emphasis id="B94BEADFFFE1FFF6FE9E6F36FEC8F9C0" bold="true" box="[153,325,1626,1650]" pageId="2" pageNumber="66">FIGURES 3–8.</emphasis>
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FF4B6F36FE61F9C3" box="[332,492,1626,1649]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEADFFFE1FFF6FF4B6F36FE61F9C3" box="[332,492,1626,1649]" italics="true" pageId="2" pageNumber="66">Madecorphnus</emphasis>
|
||
</taxonomicName>
|
||
spp., male genitalia details: (3, 6)
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FD5A6F36FBC8F9C3" authority="Frolov" authorityName="Frolov" box="[861,1093,1626,1649]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="cuccodoroi">
|
||
<emphasis id="B94BEADFFFE1FFF6FD5A6F36FC75F9C3" box="[861,1016,1626,1649]" italics="true" pageId="2" pageNumber="66">M. cuccodoroi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
holotype, (4, 7)
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FAF06F36FF52F923" authority="Frolov" authorityName="Frolov" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE1FFF6FAF06F36FA11F9C3" box="[1271,1436,1626,1649]" italics="true" pageId="2" pageNumber="66">M. perinetensis</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
, (5, 8)
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FF2E6F16FD84F923" authority="Frolov" authorityName="Frolov" box="[297,521,1658,1681]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE1FFF6FF2E6F16FE35F923" box="[297,440,1658,1681]" italics="true" pageId="2" pageNumber="66">M. peyrierasi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
; (3–5) parameres in dorsal and lateral view, (6–8) armature of the invaginated internal sac of the aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3256546FFE1FFF0FEC26FB6FA11FECB" lastPageId="4" lastPageNumber="68" pageId="2" pageNumber="66" type="diagnosis">
|
||
<paragraph id="8B8036CDFFE1FFF7FEC26FB6FB8CFF65" blockId="2.[151,1437,1754,2004]" lastBlockId="3.[151,1436,152,290]" lastPageId="3" lastPageNumber="67" pageId="2" pageNumber="66">
|
||
<emphasis id="B94BEADFFFE1FFF6FEC26FB6FECAF946" bold="true" box="[197,327,1754,1780]" pageId="2" pageNumber="66">Diagnosis.</emphasis>
|
||
This species is externally similar to
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FD046FB6FC70F946" authority="Frolov" authorityName="Frolov" box="[771,1021,1754,1780]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE1FFF6FD046FB6FC28F941" box="[771,933,1754,1779]" italics="true" pageId="2" pageNumber="66">M. peyrierasi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FA3E6FB6FAE7F946" authority="Frolov. It" authorityName="Frolov. It" box="[1081,1386,1754,1780]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE1FFF6FA3E6FB6FB7FF941" box="[1081,1266,1754,1779]" italics="true" pageId="2" pageNumber="66">M. perinetensis</emphasis>
|
||
Frolov. It
|
||
</taxonomicName>
|
||
can reliably be separated from these species only by the shape of the parameres and, especially, the internal sac armature. In
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FF316E49FE50F88C" box="[310,477,1829,1854]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE1FFF6FF316E49FE50F88C" box="[310,477,1829,1854]" italics="true" pageId="2" pageNumber="66">M. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FC1D6E49FD54F88C" box="[538,729,1829,1854]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE1FFF6FC1D6E49FD54F88C" box="[538,729,1829,1854]" italics="true" pageId="2" pageNumber="66">M. perinetensis</emphasis>
|
||
</taxonomicName>
|
||
, the internal sac armature consists of two larger sclerites basally, two smaller comma-shaped sclerites apically and an area of numerous minute spinules situated between the sclerites (
|
||
<figureCitation id="13042A48FFE1FFF6FF9C6E1CFD86F838" box="[411,523,1904,1930]" captionStart="FIGURES 3 – 8" captionStartId="2.[153,271,1626,1650]" captionTargetBox="[185,1397,201,1594]" captionTargetId="figure@2.[184,1404,194,1605]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="2" pageNumber="66">Figs. 7, 8</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C3F4D4EFFE1FFF6FC456E1CFD61F83B" box="[578,748,1904,1929]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="cuccodoroi">
|
||
<emphasis id="B94BEADFFFE1FFF6FC456E1CFD61F83B" box="[578,748,1904,1929]" italics="true" pageId="2" pageNumber="66">M. cuccodoroi</emphasis>
|
||
</taxonomicName>
|
||
, the two larger sclerites are probably homologous to those in the previous species, but they differ slightly in shape. The area composed of numerous minute spinules is larger in the new species and the spinules are somewhat coarser on average. Most prominently, a part of the apical spinules is much more robust than in the other species and they are almost tooth-shaped (
|
||
<figureCitation id="13042A48FFE0FFF7FB0F69F4FADCFF00" box="[1288,1361,152,178]" captionStart="FIGURES 3 – 8" captionStartId="2.[153,271,1626,1650]" captionTargetBox="[185,1397,201,1594]" captionTargetId="figure@2.[184,1404,194,1605]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="3" pageNumber="67">Fig. 6</figureCitation>
|
||
). This
|
||
<typeStatus id="5484886FFFE0FFF7FE9069D1FF44FF65" box="[151,201,189,215]" pageId="3" pageNumber="67">type</typeStatus>
|
||
of internal sac armature is not known in other
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FCE969D1FC13FF64" box="[750,926,189,214]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEADFFFE0FFF7FCE969D1FC13FF64" box="[750,926,189,214]" italics="true" pageId="3" pageNumber="67">Madecorphnus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE0FFF7FEC2698EFCC6FE90" blockId="3.[151,1436,152,290]" pageId="3" pageNumber="67">
|
||
The new species also differs from
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FC5E698EFD75FF49" box="[601,760,226,251]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE0FFF7FC5E698EFD75FF49" box="[601,760,226,251]" italics="true" pageId="3" pageNumber="67">M. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FD36698EFC6AFF49" box="[817,999,226,251]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE0FFF7FD36698EFC6AFF49" box="[817,999,226,251]" italics="true" pageId="3" pageNumber="67">M. perinetensis</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the parameres, which are somewhat wider and more curved at apices (
|
||
<figureCitation id="13042A48FFE0FFF7FCCB6864FCB3FE90" box="[716,830,264,290]" captionStart="FIGURES 3 – 8" captionStartId="2.[153,271,1626,1650]" captionTargetBox="[185,1397,201,1594]" captionTargetId="figure@2.[184,1404,194,1605]" captionTargetPageId="2" captionText="FIGURES 3 – 8. Madecorphnus spp., male genitalia details: (3, 6) M. cuccodoroi Frolov holotype, (4, 7) M. perinetensis Frolov, (5, 8) M. peyrierasi Frolov; (3 – 5) parameres in dorsal and lateral view, (6 – 8) armature of the invaginated internal sac of the aedeagus." httpUri="https://zenodo.org/record/294099/files/figure.png" pageId="3" pageNumber="67">Figs. 3–5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF406645FFE0FFF7FE906E19FDEDF81F" httpUri="https://zenodo.org/record/293367/files/figure.png" pageId="3" pageNumber="67" targetBox="[177,1413,351,1886]" targetPageId="3">
|
||
<paragraph id="8B8036CDFFE0FFF7FE906E19FDEDF81F" blockId="3.[151,1437,1909,1965]" pageId="3" pageNumber="67">
|
||
<emphasis id="B94BEADFFFE0FFF7FE906E19FE90F83F" bold="true" box="[151,285,1909,1933]" pageId="3" pageNumber="67">FIGURE 9.</emphasis>
|
||
<emphasis id="B94BEADFFFE0FFF7FF236E1AFEBAF83F" box="[292,311,1910,1933]" italics="true" pageId="3" pageNumber="67">D</emphasis>
|
||
istribution map of
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FC036E1AFCF7F83F" authority="Frolov" authorityName="Frolov" box="[516,890,1910,1933]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="cuccodoroi">
|
||
<emphasis id="B94BEADFFFE0FFF7FC036E1AFCA4F83F" box="[516,809,1910,1933]" italics="true" pageId="3" pageNumber="67">Madecorphnus cuccodoroi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FD826E1AFB0DF83F" authority="Frolov" authorityName="Frolov" box="[901,1152,1910,1933]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE0FFF7FD826E1AFBA3F83F" box="[901,1070,1910,1933]" italics="true" pageId="3" pageNumber="67">M. perinetensis</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FA8B6E1AFAFEF83F" authority="Frolov" authorityName="Frolov" box="[1164,1395,1910,1933]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="montreuili">
|
||
<emphasis id="B94BEADFFFE0FFF7FA8B6E1AFAAFF83F" box="[1164,1314,1910,1933]" italics="true" pageId="3" pageNumber="67">M. montreuili</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FB786E1AFEC8F81F" authority="Paulian" authorityName="Paulian" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="falcatus">
|
||
<emphasis id="B94BEADFFFE0FFF7FB786E1AFF61F81F" italics="true" pageId="3" pageNumber="67">M. falcatus</emphasis>
|
||
Paulian
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3F4D4EFFE0FFF7FF7B6EFAFDEDF81F" authority="Frolov." authorityName="Frolov." box="[380,608,1942,1965]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE0FFF7FF7B6EFAFD81F81F" box="[380,524,1942,1965]" italics="true" pageId="3" pageNumber="67">M. peyrierasi</emphasis>
|
||
Frolov.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8036CDFFE7FFF0FEC269F4FE26FF65" blockId="4.[151,1436,152,215]" pageId="4" pageNumber="68">
|
||
In the key to
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FF6169F4FD97FF03" box="[358,538,152,177]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEADFFFE7FFF0FF6169F4FD97FF03" box="[358,538,152,177]" italics="true" pageId="4" pageNumber="68">Madecorphnus</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFAE4B3CFFE7FFF0FC8E69F4FCA4FF00" author="Frolov" box="[649,809,152,178]" pageId="4" pageNumber="68" refString="Frolov, A. V. (2010) Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera, Scarabaeidae, Orphninae). Journal of Natural History, 44 (17), 1095 - 1111." type="journal article" year="2010">Frolov, 2010</bibRefCitation>
|
||
), couplet 10 should be updated to accommodate the new species as follows:
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE7FFF0FE90686EFA11FECB" blockId="4.[151,1436,258,377]" pageId="4" pageNumber="68">
|
||
10. Internal sac of aedeagus with two large separate sclerites and numerous small spinules, some of which are robust and almost tooth-shaped and may be arranged into two indistinctly separated clusters...............
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FAB8684EFAD4FE8B" box="[1215,1369,290,313]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="cuccodoroi" status="sp. nov.">
|
||
<emphasis id="B94BEADFFFE7FFF0FAB8684EFAD4FE8B" box="[1215,1369,290,313]" italics="true" pageId="4" pageNumber="68">M. cuccodoroi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94BEADFFFE7FFF0FB67684EFA16FE88" bold="true" box="[1376,1435,290,314]" pageId="4" pageNumber="68">
|
||
<taxonomicNameLabel id="A27857A4FFE7FFF0FB67684EFA16FE88" box="[1376,1435,290,314]" pageId="4" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
- Internal sac of aedeagus with two or more separate sclerites and numerous small spinules; the later are very fine and are all approximately the same size................11 (
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FCDF680EFC20FECB" authority="Frolov" authorityName="Frolov" box="[728,941,354,377]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="B94BEADFFFE7FFF0FCDF680EFCD3FECB" box="[728,862,354,377]" italics="true" pageId="4" pageNumber="68">M. brunneus</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FDBF680EFB26FECB" authority="Frolov" authorityName="Frolov" box="[952,1195,354,377]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="perinetensis">
|
||
<emphasis id="B94BEADFFFE7FFF0FDBF680EFBD0FECB" box="[952,1117,354,377]" italics="true" pageId="4" pageNumber="68">M. perinetensis</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FAB2680EFA18FECB" authority="Frolov" authorityName="Frolov" box="[1205,1429,354,377]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94BEADFFFE7FFF0FAB2680EFAC8FECB" box="[1205,1349,354,377]" italics="true" pageId="4" pageNumber="68">M. peyrierasi</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256546FFE7FFF0FEC268CEFAAAFD55" pageId="4" pageNumber="68" type="distribution">
|
||
<paragraph id="8B8036CDFFE7FFF0FEC268CEFAA4FDC5" blockId="4.[151,1436,418,780]" pageId="4" pageNumber="68">
|
||
<emphasis id="B94BEADFFFE7FFF0FEC268CEFEE8FE0E" bold="true" box="[197,357,418,444]" pageId="4" pageNumber="68">Distribution.</emphasis>
|
||
The beetles were collected near the main camp site of the Amber Mountain National Park in Antsiranana Province (northern
|
||
<collectingCountry id="F328765DFFE7FFF0FC1E68A4FD22FE50" box="[537,687,456,482]" name="Madagascar" pageId="4" pageNumber="68">Madagascar</collectingCountry>
|
||
). The park occupies the Madagascan northernmost remnant of indigenous humid forest covering about 200 square km (
|
||
<figureCitation id="13042A48FFE7FFF0FD2B6881FCE0FDB5" box="[812,877,493,519]" captionStart="FIGURE 9. D" captionStartId="3.[151,256,1909,1933]" captionTargetBox="[177,1413,351,1886]" captionTargetId="figure@3.[169,1417,343,1888]" captionTargetPageId="3" captionText="FIGURE 9. D istribution map of Madecorphnus cuccodoroi Frolov, M. perinetensis Frolov, M. montreuili Frolov, M. falcatus Paulian, and M. peyrierasi Frolov." httpUri="https://zenodo.org/record/293367/files/figure.png" pageId="4" pageNumber="68">Fig 9</figureCitation>
|
||
). It is possible that the range of the new species is limited to this forest. This locality record is now the northernmost for the genus
|
||
<emphasis id="B94BEADFFFE7FFF0FA526B7EFA81FD99" box="[1109,1292,530,555]" italics="true" pageId="4" pageNumber="68">
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FA526B7EFA85FD99" box="[1109,1288,530,555]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="genus">Madecorphnus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and is some
|
||
<quantity id="4CC79B28FFE7FFF0FE906B54FF62FDE0" box="[151,239,568,594]" metricMagnitude="5" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="68" unit="km" value="450.0">450 km</quantity>
|
||
north of the collecting localities of
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FC946B54FCFBFDE0" authority="Paulian" authorityName="Paulian" box="[659,886,568,594]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="falcatus">
|
||
<emphasis id="B94BEADFFFE7FFF0FC946B54FC9AFDE3" box="[659,791,568,593]" italics="true" pageId="4" pageNumber="68">M. falcatus</emphasis>
|
||
Paulian
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FDA86B54FB28FDE0" authority="Frolov" authorityName="Frolov" box="[943,1189,568,594]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="montreuili">
|
||
<emphasis id="B94BEADFFFE7FFF0FDA86B54FBDDFDE3" box="[943,1104,568,593]" italics="true" pageId="4" pageNumber="68">M. montreuili</emphasis>
|
||
Frolov
|
||
</taxonomicName>
|
||
in Antongil Bay. All the other species of the genus were collected south of the latitude of
|
||
<geoCoordinate id="EE0B500AFFE7FFF0FDBE6B31FB98FDC5" box="[953,1045,605,631]" direction="south" orientation="latitude" pageId="4" pageNumber="68" precision="925" value="-18.5">18°30'S</geoCoordinate>
|
||
(
|
||
<bibRefCitation id="EFAE4B3CFFE7FFF0FA236B31FA91FDC5" author="Frolov" box="[1060,1308,605,631]" pageId="4" pageNumber="68" refString="Frolov, A. V. (2010) Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera, Scarabaeidae, Orphninae). Journal of Natural History, 44 (17), 1095 - 1111." type="journal article" year="2010">Frolov 2010, Fig. 31</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8036CDFFE7FFF0FEC26BEEFAAAFD55" blockId="4.[151,1436,418,780]" pageId="4" pageNumber="68">
|
||
<emphasis id="B94BEADFFFE7FFF0FEC26BEEFEB3FD2E" bold="true" box="[197,318,642,668]" pageId="4" pageNumber="68">Bionomy.</emphasis>
|
||
The natural history of the new species was not specifically studied by the collector. All the specimens were collected by sifting forest litter. The number of collected specimens suggests that the population density of
|
||
<taxonomicName id="4C3F4D4EFFE7FFF0FF906BA1FDCCFD54" box="[407,577,717,742]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="68" phylum="Arthropoda" rank="species" species="cuccodoroi">
|
||
<emphasis id="B94BEADFFFE7FFF0FF906BA1FDCCFD54" box="[407,577,717,742]" italics="true" pageId="4" pageNumber="68">M. cuccodoroi</emphasis>
|
||
</taxonomicName>
|
||
is low at least during the season when the collecting was done.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256546FFE7FFF0FEC26B9EFAF5FCBE" box="[197,1400,754,780]" pageId="4" pageNumber="68" type="etymology">
|
||
<paragraph id="8B8036CDFFE7FFF0FEC26B9EFAF5FCBE" blockId="4.[151,1436,418,780]" box="[197,1400,754,780]" pageId="4" pageNumber="68">
|
||
<emphasis id="B94BEADFFFE7FFF0FEC26B9EFEDFFCBE" bold="true" box="[197,338,754,780]" pageId="4" pageNumber="68">Etymology.</emphasis>
|
||
The new species is named after Giulio Cuccodoro (MHNG) who collected the
|
||
<typeStatus id="5484886FFFE7FFF0FAF36B9EFAABFCBE" box="[1268,1318,754,780]" pageId="4" pageNumber="68">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |