222 lines
37 KiB
XML
222 lines
37 KiB
XML
<document id="1A3F6DB60A518240831CD22D65562AE7" ID-DOI="10.11646/zootaxa.4040.1.7" ID-GBIF-Dataset="a33544e3-4041-419b-902c-b0a26d3e01fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="289991" ID-ZooBank="42E61C33-C8C5-45E9-8CED-6B4D68A38538" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333966621" checkinUser="plazi" docAuthor="Houben, Albrecht M., Schwank, Peter, Proesmans, Willem, Bert, Wim & Artois, Tom J." docDate="2015" docId="039687BDFFEA043C7AFA46E1FB34C55E" docLanguage="en" docName="zt04040p092.pdf" docOrigin="Zootaxa 4040 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carcharodopharynx arcanus (Reisinger, 1924) Poche 1926" docType="treatment" docVersion="5" lastPageNumber="88" masterDocId="FFAFFFC5FFEF04397A6D4676FF88C333" masterDocTitle="Notes on some enigmatic taxa of limnoterrestrial rhabdocoels, with the description of two new species" masterLastPageNumber="92" masterPageNumber="83" pageNumber="88" updateTime="1698618935067" updateUser="plazi">
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<mods:title id="1C329CE28A2A1A5E7C4C6E196B9A316C">Notes on some enigmatic taxa of limnoterrestrial rhabdocoels, with the description of two new species</mods:title>
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<mods:namePart id="E0342E2033C91C915A891CF92DA0C526">Houben, Albrecht M.</mods:namePart>
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<mods:namePart id="7ECABCA8977082328EAE6D80153CB94D">Schwank, Peter</mods:namePart>
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<mods:namePart id="EBE80FB9E8E94D66E001CDEB30629E2F">Proesmans, Willem</mods:namePart>
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<mods:namePart id="F28C58905B2E7694291BC4480DF3A5BE">Bert, Wim</mods:namePart>
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<mods:namePart id="F6A4FF0BCD652ED80E6BA34000525B1F">Artois, Tom J.</mods:namePart>
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<mods:title id="F3FB674D7CD94373C1C4C7D5B4FF8B3F">Zootaxa</mods:title>
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<mods:date id="2A219C7C656C37F3798D3ACA0D218E37">2015</mods:date>
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<mods:number id="3A20305E3BF2224052E8E025A134DE63">4040</mods:number>
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<treatment id="039687BDFFEA043C7AFA46E1FB34C55E" ID-DOI="http://doi.org/10.5281/zenodo.6112826" ID-GBIF-Taxon="127694093" ID-Zenodo-Dep="6112826" LSID="urn:lsid:plazi:treatment:039687BDFFEA043C7AFA46E1FB34C55E" httpUri="http://treatment.plazi.org/id/039687BDFFEA043C7AFA46E1FB34C55E" lastPageNumber="88" pageId="5" pageNumber="88">
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<subSubSection id="C3256520FFEA043C7AFA46E1FE48C122" pageId="5" pageNumber="88" type="nomenclature">
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<paragraph id="8B8036ABFFEA043C7AFA46E1FCE2C381" blockId="5.[151,874,151,211]" box="[151,874,151,178]" pageId="5" pageNumber="88">
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<heading id="D0C881C7FFEA043C7AFA46E1FCE2C381" bold="true" box="[151,874,151,178]" fontSize="11" level="1" pageId="5" pageNumber="88" reason="1">
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<taxonomicName id="4C3F4D28FFEA043C7AFA46E1FCE2C381" authority="(Reisinger, 1924) Poche, 1926" authorityName="(Reisinger, 1924) Poche" authorityYear="1926" box="[151,874,151,178]" class="Rhabditophora" family="Carcharodopharyngidae" genus="Carcharodopharynx" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="88" phylum="Platyhelminthes" rank="species" species="arcanus">
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<emphasis id="B94BEAB9FFEA043C7AFA46E1FCE2C381" bold="true" box="[151,874,151,178]" pageId="5" pageNumber="88">
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<emphasis id="B94BEAB9FFEA043C7AFA46E1FE7EC382" bold="true" box="[151,502,151,177]" italics="true" pageId="5" pageNumber="88">Carcharodopharynx arcanus</emphasis>
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(
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<bibRefCitation id="EFAE4B5AFFEA043C786946EEFD4DC381" author="Reisinger" box="[516,709,152,178]" pageId="5" pageNumber="88" refString="Reisinger, E. (1924) Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger, 59, 1 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
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)
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<bibRefCitation id="EFAE4B5AFFEA043C78B846EEFCE2C381" author="Poche" box="[725,874,152,178]" pageId="5" pageNumber="88" refString="Poche, F. (1926) Das System der Platodaria. Archiv fur Naturgeschichte, 91, 1 - 458." type="journal article" year="1926">Poche, 1926</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8036ABFFEA043C7AFA46CCFEA3C3E0" blockId="5.[151,874,151,211]" box="[151,299,186,211]" pageId="5" pageNumber="88">
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(
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<figureCitation id="13042A2EFFEA043C7AF246CCFF64C3E0" box="[159,236,186,211]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Figs. 2</figureCitation>
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A–C)
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</paragraph>
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<paragraph id="8B8036ABFFEA043C7AFA4774FDE7C22B" blockId="5.[151,623,258,280]" box="[151,623,258,280]" pageId="5" pageNumber="88">
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Syn.
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<taxonomicName id="4C3F4D28FFEA043C7AA64774FDE7C22B" authority="Reisinger, 1924" authorityName="Reisinger" authorityYear="1924" box="[203,623,258,280]" class="Adenophorea" family="Desmodoridae" genus="Acanthopharynx" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="88" phylum="Nematoda" rank="species" species="arcanus">
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<emphasis id="B94BEAB9FFEA043C7AA64774FE41C22B" box="[203,457,258,280]" italics="true" pageId="5" pageNumber="88">Acanthopharynx arcanus</emphasis>
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<bibRefCitation id="EFAE4B5AFFEA043C7BA24774FDE7C22B" author="Reisinger" box="[463,623,258,280]" pageId="5" pageNumber="88" refString="Reisinger, E. (1924) Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger, 59, 1 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8B8036ABFFEA043C7AFA4733FB45C2B2" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
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<emphasis id="B94BEAB9FFEA043C7AFA4733FECCC26D" bold="true" box="[151,324,325,350]" pageId="5" pageNumber="88">New localities.</emphasis>
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Oberau, Bavaria,
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<collectingCountry id="F328763BFFEA043C787A4730FDF7C26D" box="[535,639,326,350]" name="Germany" pageId="5" pageNumber="88">Germany</collectingCountry>
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(
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<geoCoordinate id="EE0B506CFFEA043C78FC4730FC95C26D" box="[657,797,326,350]" direction="north" orientation="latitude" pageId="5" pageNumber="88" precision="15" value="47.559166">47°33’33”N</geoCoordinate>
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;
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<geoCoordinate id="EE0B506CFFEA043C79414733FC3AC26D" box="[812,946,325,350]" direction="east" orientation="longitude" pageId="5" pageNumber="88" precision="15" value="11.115833">11°06’57”E</geoCoordinate>
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) in forest litter (
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<date id="FF81106BFFEA043C7E1F4730FA83C26D" box="[1138,1291,325,350]" pageId="5" pageNumber="88" value="2011-07-13">13 July 2011</date>
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). Kreuzberg, Weyer,
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<collectingCountry id="F328763BFFEA043C7A81471FFEB6C2B1" box="[236,318,361,386]" name="Austria" pageId="5" pageNumber="88">Austria</collectingCountry>
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(
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<geoCoordinate id="EE0B506CFFEA043C7B23471FFE53C2B1" box="[334,475,361,386]" direction="north" orientation="latitude" pageId="5" pageNumber="88" precision="15" value="47.859997">47°51’36”N</geoCoordinate>
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;
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<geoCoordinate id="EE0B506CFFEA043C7B8A471FFDE5C2B2" box="[487,621,361,385]" direction="east" orientation="longitude" pageId="5" pageNumber="88" precision="15" value="14.652499">14°39’09”E</geoCoordinate>
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) in humid mosses and leafy humus (
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<date id="FF81106BFFEA043C7E64471FFB49C2B2" box="[1033,1217,361,385]" pageId="5" pageNumber="88" value="2011-08-29">29 August 2011</date>
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).
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</paragraph>
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<paragraph id="8B8036ABFFEA043C7AAA47FBFE48C122" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
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<emphasis id="B94BEAB9FFEA043C7AAA47FBFE32C295" bold="true" box="[199,442,397,422]" pageId="5" pageNumber="88">Known distribution.</emphasis>
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Graz,
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<collectingCountry id="F328763BFFEA043C786847FBFDD2C295" box="[517,602,397,422]" name="Austria" pageId="5" pageNumber="88">Austria</collectingCountry>
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, widespread in forest soils (
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<bibRefCitation id="EFAE4B5AFFEA043C79FE47FBFBC1C295" author="Reisinger" box="[915,1097,397,422]" pageId="5" pageNumber="88" refString="Reisinger, E. (1924) Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger, 59, 1 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
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); Pallanza,
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<collectingCountry id="F328763BFFEA043C7EA747FBFB75C295" box="[1226,1277,397,422]" name="Italy" pageId="5" pageNumber="88">Italy</collectingCountry>
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in forest soils (
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<bibRefCitation id="EFAE4B5AFFEA043C7AF347C7FEDCC2F9" author="Steinbock" box="[158,340,433,458]" pageId="5" pageNumber="88" refString="Steinbock, O. (1951) Turbellarienstudien am Lago Maggiore I. Memorie dell'istituto Italiano di idrobiologia dott. Marco de Marchi, 6, 137 - 164." type="journal article" year="1951">Steinböck 1951</bibRefCitation>
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); Tvärminne,
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<collectingCountry id="F328763BFFEA043C7B9847C7FDC3C2F9" box="[501,587,433,458]" name="Finland" pageId="5" pageNumber="88">Finland</collectingCountry>
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in mosses (
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<bibRefCitation id="EFAE4B5AFFEA043C78BF47C7FCEBC2F9" author="Luther" box="[722,867,433,458]" pageId="5" pageNumber="88" refString="Luther, A. (1963) Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica, 16, 1 - 163, Taf. I - II." type="journal article" year="1963">Luther 1963</bibRefCitation>
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); Sierra de Carzola and Finca Caleron,
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<collectingCountry id="F328763BFFEA043C7F4547C7FAE0C2F9" box="[1320,1384,433,458]" name="Spain" pageId="5" pageNumber="88">Spain</collectingCountry>
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(An der Lan 1963); Schlitz,
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<collectingCountry id="F328763BFFEA043C7BDC47A0FD92C2DD" box="[433,538,470,494]" name="Germany" pageId="5" pageNumber="88">Germany</collectingCountry>
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, in beech litter near helocrene springs (
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<bibRefCitation id="EFAE4B5AFFEA043C798347A3FB16C2DD" author="Schwank" box="[1006,1182,469,494]" pageId="5" pageNumber="88" refString="Schwank, P. (1981) Seltene und wenig bekannte Strudelwurmer (Turbellaria) aus Quellen und Bergbachen Mitteleuropas. Beitrage zur Naturkunde in Osthessen, 17, 101 - 131." type="journal article" year="1981">Schwank 1981</bibRefCitation>
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); Carpathians,
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<collectingCountry id="F328763BFFEA043C7F2347A3FA14C2DD" box="[1358,1436,469,494]" name="Poland" pageId="5" pageNumber="88">Poland</collectingCountry>
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(Kolasa in
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<bibRefCitation id="EFAE4B5AFFEA043C7B7F478FFE30C122" author="Schwank" box="[274,440,505,530]" pageId="5" pageNumber="88" refString="Schwank, P. (1981) Seltene und wenig bekannte Strudelwurmer (Turbellaria) aus Quellen und Bergbachen Mitteleuropas. Beitrage zur Naturkunde in Osthessen, 17, 101 - 131." type="journal article" year="1981">Schwank 1981</bibRefCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3256520FFEA043C7AAA446BFBC9C579" pageId="5" pageNumber="88" type="materials_examined">
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<paragraph id="8B8036ABFFEA043C7AAA446BFD37C169" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
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<emphasis id="B94BEAB9FFEA043C7AAA446BFEBEC105" bold="true" box="[199,310,541,566]" pageId="5" pageNumber="88">Material.</emphasis>
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Three adult specimens studied alive. One sagittally-sectioned specimen from Oberau, Bavaria,
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<collectingCountry id="F328763BFFEA043C7AFA4437FF77C16A" box="[151,255,577,601]" name="Germany" pageId="5" pageNumber="88">Germany</collectingCountry>
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designated
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<typeStatus id="54848809FFEA043C7BE84434FE56C169" box="[389,478,578,602]" pageId="5" pageNumber="88" type="neotype">neotype</typeStatus>
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(
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<collectionCode id="ED2EAE6EFFEA043C7B814437FDC9C16A" box="[492,577,577,601]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="5" pageNumber="88">SMNH</collectionCode>
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, no. 8760).
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</paragraph>
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<paragraph id="8B8036ABFFEA043C7AAA4413FBA3C761" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
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<emphasis id="B94BEAB9FFEA043C7AAA4413FE14C14D" bold="true" box="[199,412,613,638]" pageId="5" pageNumber="88">Descriptive notes.</emphasis>
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Animals about
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<quantity id="4CC79B4EFFEA043C783C4413FD51C14E" box="[593,729,613,638]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="5" pageNumber="88" unit="mm" value="1.15" valueMax="1.5" valueMin="0.8">0.8–1.5 mm</quantity>
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long and usually very dark because of gut contents. The body shape is rounded anteriorly and ends in a small tail with small eosinophilic tail glands. Small, closely-packedtogether dermal rhabdites are present all over the epidermis, adenal rhabdites are lacking. Two separate protonephridiopores (
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<figureCitation id="13042A2EFFEA043C7BE244A7FE50C1D9" box="[399,472,721,746]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
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A: pp) open ventrally at ±35 % of the body. The mouth (
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<figureCitation id="13042A2EFFEA043C7EE044A7FB5DC1D9" box="[1165,1237,721,746]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
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A, 2C: m) has a subterminal position and opens into the prepharyngeal cavity, which is surrounded with longitudinal muscles and is lined with a very thin epithelium. The 75-µm-long pharynx (
|
||
<figureCitation id="13042A2EFFEA043C7953456FFC08C001" box="[830,896,793,818]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C) is situated at ±25 % and is directed forwards. It is divided into an anterior and a posterior part. The anterior part protrudes into the prepharyngeal cavity and contains many longitudinal muscles (
|
||
<figureCitation id="13042A2EFFEA043C78554517FDF5C049" box="[568,637,865,890]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: lm). The distal wall of this anterior part is provided with many small spines (
|
||
<figureCitation id="13042A2EFFEA043C7A8345F3FEBBC0AD" box="[238,307,901,926]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: ss), while its proximal wall is lined by a thin epithelium. The posterior part is a typical pharynx rosulatus, with the musculature consisting of outer circular muscles (
|
||
<figureCitation id="13042A2EFFEA043C79C345DFFC7CC0F1" box="[942,1012,937,962]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: ocm), which also are found in the anterior part, outer longitudinal muscles (
|
||
<figureCitation id="13042A2EFFEA043C780145BBFD38C0D5" box="[620,688,973,998]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: olm), radial muscles (
|
||
<figureCitation id="13042A2EFFEA043C79A545BBFB84C0D5" box="[968,1036,973,998]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: rm), inner circular muscles (
|
||
<figureCitation id="13042A2EFFEA043C7F0245BBFF2FC739" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: icm) and inner longitudinal muscles (
|
||
<figureCitation id="13042A2EFFEA043C78064587FD38C739" box="[619,688,1009,1034]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: ilm). The pharyngeal glands (
|
||
<figureCitation id="13042A2EFFEA043C7E704587FBE8C739" box="[1053,1120,1009,1034]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
C: pg) have intra- as well as extracapsular cell bodies. Live animals often retract and protrude the spiny anterior part, so that the shape of this part can vary from a short and broad trunk to a very elongate and narrow channel.
|
||
</paragraph>
|
||
<paragraph id="8B8036ABFFEA043C7AAA422BFB67C78D" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
|
||
The gonopore (
|
||
<figureCitation id="13042A2EFFEA043C7B1B422BFE33C745" box="[374,443,1117,1142]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
A, 2B: gp) is situated at ±50 % of the body and is connected with the common genital atrium by means of a long gonoduct (
|
||
<figureCitation id="13042A2EFFEA043C782342F7FD1EC7A9" box="[590,662,1153,1178]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: gd). Both are surrounded by circular muscles, which increase in strength towards the genital atrium. The genital atrium is lined with a high eosinophilic epithelium.
|
||
</paragraph>
|
||
<paragraph id="8B8036ABFFEA043C7AAA42BFFED3C689" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
|
||
The small, round testes (
|
||
<figureCitation id="13042A2EFFEA043C7BB642BFFD97C7D1" box="[475,543,1225,1250]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
A: t) lie at ±60 % of the body and are situated ventrally from the vitellaria (
|
||
<figureCitation id="13042A2EFFEA043C7F0342BFFF2EC635" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
A: vi). The paired vasa deferentia (
|
||
<figureCitation id="13042A2EFFEA043C785B429BFDF4C635" box="[566,636,1261,1286]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: vde) enter the copulatory organ separately at its proximal end. Two inner spiral muscle layers and an outer longitudinal layer surround this 57-µm-long, oval copulatory bulb, which is filled with prostate secretion produced by large, eosinophilic, extracapsular prostate glands (
|
||
<figureCitation id="13042A2EFFEA043C7E8F4343FAA4C67D" box="[1250,1324,1333,1358]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: gg). It contains a large seminal vesicle (
|
||
<figureCitation id="13042A2EFFEA043C7867432FFDC6C641" box="[522,590,1369,1394]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: vs) in its proximal part. The unarmed cirrus (
|
||
<figureCitation id="13042A2EFFEA043C7E05432FFB24C641" box="[1128,1196,1369,1394]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: cir) is found in the distal third of the copulatory organ. The male duct (
|
||
<figureCitation id="13042A2EFFEA043C78B0430BFCA9C6A5" box="[733,801,1405,1430]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: md) is surrounded by circular muscles and is lined by a low epithelium.
|
||
</paragraph>
|
||
<paragraph id="8B8036ABFFEA043C7AAA43B3FBC9C579" blockId="5.[151,1437,325,1646]" pageId="5" pageNumber="88">
|
||
Paired vitellaria (
|
||
<figureCitation id="13042A2EFFEA043C7BE643B3FE58C6ED" box="[395,464,1477,1502]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
A: vi) extend from the posterior end to ±40 % of the body. They are located dorsally, but extend to the ventral side at their anterior end. Paired vitelloducts unite to form a single vitelloduct (
|
||
<figureCitation id="13042A2EFFEA043C7F53439FFA0CC531" box="[1342,1412,1513,1538]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: vd), which fuses with a short oviduct (
|
||
<figureCitation id="13042A2EFFEA043C782F407BFD0EC515" box="[578,646,1549,1574]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: od) to form the female duct (
|
||
<figureCitation id="13042A2EFFEA043C7988407BFBA1C515" box="[997,1065,1549,1574]" captionStart="FIGURE 2" captionStartId="6.[151,250,1896,1918]" captionTargetBox="[603,1393,1165,1871]" captionTargetId="figure@6.[976,1144,1420,1597]" captionTargetPageId="6" captionText="FIGURE 2. Carcharodopharynx arcanus: (A) Habitus of live animal from Oberau. (B) Reconstruction from sagittal sections of the reproductive organs (SMNH, no. 8760). (C) Reconstruction from sagittal sections of the pharynx (SMNH, no. 8760); C. schlitzensis n. sp.: (D) Habitus of live animal. (E) Free swimming animal. (F) Live animal. (G) Male copulatory organ. D – E, after Schwank (1981)." httpUri="https://zenodo.org/record/289992/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
|
||
B: fd). The latter narrows distally and its surrounding circular muscles increase in strength towards the genital atrium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3256520FFEA043C7AAA4023FB34C55E" box="[199,1212,1621,1646]" pageId="5" pageNumber="88" type="discussion">
|
||
<paragraph id="8B8036ABFFEA043C7AAA4023FB34C55E" blockId="5.[151,1437,325,1646]" box="[199,1212,1621,1646]" pageId="5" pageNumber="88">
|
||
<emphasis id="B94BEAB9FFEA043C7AAA4023FEC4C55D" bold="true" box="[199,332,1621,1646]" pageId="5" pageNumber="88">Discussion.</emphasis>
|
||
See the discussion following the descriptive notes and remarks on
|
||
<taxonomicName id="4C3F4D28FFEA043C7E514020FB3DC55E" box="[1084,1205,1622,1645]" class="Adenophorea" family="Desmodoridae" genus="Acanthopharynx" kingdom="Animalia" order="Desmodorida" pageId="5" pageNumber="88" phylum="Nematoda" rank="species" species="arcanus">
|
||
<emphasis id="B94BEAB9FFEA043C7E514020FB3DC55E" box="[1084,1205,1622,1645]" italics="true" pageId="5" pageNumber="88">C. arcanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |