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<document id="F21C02380F0A4A2AF7ED5F13BC1C81DE" ID-CLB-Dataset="21713" ID-DOI="10.5852/ejt.2020.655" ID-GBIF-Dataset="fadee126-7da7-47d9-b9ac-973697ecf9f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3888797" ID-ZooBank="0042EFAE-2E4F-444B-AFB9-E321D16116E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591816327545" checkinUser="valdenar" docAuthor="Bello, Giambattista" docDate="2020" docId="03968791BD3CFFD6911995E0FE47FC44" docLanguage="en" docName="ejt-655_bello.pdf.imf" docOrigin="European Journal of Taxonomy 655" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Euprymna Steenstrup 1887" docType="treatment" docVersion="6" lastPageNumber="40" masterDocId="FFAFFFE9BD1AFFF1936A9019FFD7FFFC" masterDocTitle="Evolution of the hectocotylus in Sepiolinae (Cephalopoda: Sepiolidae) and description of four new genera" masterLastPageNumber="53" masterPageNumber="1" pageNumber="39" updateTime="1698842281433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B5C6E1DF64FC461F815DF953B688793E">Evolution of the hectocotylus in Sepiolinae (Cephalopoda: Sepiolidae) and description of four new genera</mods:title>
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<mods:title id="652C09F7B4617A37C098716A81D67B79">European Journal of Taxonomy</mods:title>
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<mods:date id="1C5C72D0ECB2185A1792464B5BC86F42">2020</mods:date>
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<mods:classification id="711183CF13987D055C0B8ABE355485C3">journal article</mods:classification>
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<treatment id="03968791BD3CFFD6911995E0FE47FC44" ID-DOI="http://doi.org/10.5281/zenodo.5920277" ID-GBIF-Taxon="164500954" ID-Zenodo-Dep="5920277" LSID="urn:lsid:plazi:treatment:03968791BD3CFFD6911995E0FE47FC44" httpUri="http://treatment.plazi.org/id/03968791BD3CFFD6911995E0FE47FC44" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="C325650CBD3CFFD7911995E0FC16F9EE" box="[627,961,1528,1555]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8B803687BD3CFFD7911995E0FC16F9EE" blockId="38.[627,961,1528,1591]" box="[627,961,1528,1555]" pageId="38" pageNumber="39">
<heading id="D0C881EBBD3CFFD7911995E0FC16F9EE" box="[627,961,1528,1555]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<taxonomicName id="4C3F4D04BD3CFFD7911995E0FC16F9EE" authority="Steenstrup, 1887" authorityName="Steenstrup" authorityYear="1887" box="[627,961,1528,1555]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD3CFFD7911995E0FD23F9EF" bold="true" box="[627,756,1529,1555]" italics="true" pageId="38" pageNumber="39">Euprymna</emphasis>
Steenstrup, 1887
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD3CFFD7919D9605FCEBF9CB" box="[759,828,1564,1591]" pageId="38" pageNumber="39" type="description">
<paragraph id="8B803687BD3CFFD7919D9605FCEBF9CB" blockId="38.[627,961,1528,1591]" box="[759,828,1564,1591]" pageId="38" pageNumber="39">
<figureCitation id="13042A02BD3CFFD7919D9605FCEBF9CB" box="[759,828,1564,1591]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[441,1148,1083,1882]" captionTargetId="figure@7.[403,1183,1083,1885]" captionTargetPageId="7" captionText="Fig. 7. Schematic drawing of the hectocotylus of Euprymna Steenstrup, 1887. Abbreviations: cop. app. = copulatory apparatus (after Norman &amp; Lu 1997, modified)." figureDoi="http://doi.org/10.5281/zenodo.3888811" httpUri="https://zenodo.org/record/3888811/files/figure.png" pageId="38" pageNumber="39">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD3CFFD793D79646FD92F984" box="[189,581,1630,1656]" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="8B803687BD3CFFD793D79646FD92F984" blockId="38.[189,581,1630,1656]" box="[189,581,1630,1656]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="AB2F11A9BD3CFFD793D79646FD92F984" box="[189,581,1630,1656]" pageId="38" pageNumber="39">
<treatmentCitation id="0A9E1096BD3CFFD793D79646FD96F984" author="Steenstrup J." box="[189,577,1630,1656]" page="43" pageId="38" pageNumber="39" year="1887">
<taxonomicName id="4C3F4D04BD3CFFD793D79646FD96F984" authority="Steenstrup, 1887 a: 43" authorityName="Steenstrup" authorityPageNumber="43" authorityYear="1887" box="[189,577,1630,1656]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD3CFFD793D79646FEE0F984" box="[189,311,1631,1656]" italics="true" pageId="38" pageNumber="39">Euprymna</emphasis>
<bibRefCitation id="EFAE4B76BD3CFFD792559647FD96F984" author="Steenstrup J." box="[319,577,1630,1656]" pageId="38" pageNumber="39" pagination="47 - 66" refId="ref28822" refString="Steenstrup J. 1887 a. Notae Teuthologicae. 6. Species generis Sepiolae Maris Mediterranei. Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger 1887: 47 - 66." type="journal article" year="1887">Steenstrup, 1887a: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD3CFFD793D796B8FCAEF916" pageId="38" pageNumber="39" type="type_taxon">
<paragraph id="8B803687BD3CFFD793D796B8FE8FF946" blockId="38.[189,344,1696,1723]" box="[189,344,1696,1723]" pageId="38" pageNumber="39">
<heading id="D0C881EBBD3CFFD793D796B8FE8FF946" bold="true" box="[189,344,1696,1723]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B94BEA95BD3CFFD793D796B8FE8FF946" bold="true" box="[189,344,1696,1723]" pageId="38" pageNumber="39">
<typeStatus id="54848825BD3CFFD793D796B8FF2DF947" box="[189,250,1697,1723]" pageId="38" pageNumber="39">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD3CFFD793D796C9FCAEF916" blockId="38.[189,889,1744,1770]" box="[189,889,1744,1770]" pageId="38" pageNumber="39">
<taxonomicName id="4C3F4D04BD3CFFD793D796C9FDF8F916" authority="Verrill, 1881" authorityName="Verrill" authorityYear="1881" box="[189,559,1744,1770]" class="Cephalopoda" family="Sepiolidae" genus="Inioteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Sepiida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="morsei">
<emphasis id="B94BEA95BD3CFFD793D796C9FE45F915" box="[189,402,1744,1770]" italics="true" pageId="38" pageNumber="39">Inioteuthis morsei</emphasis>
<bibRefCitation id="EFAE4B76BD3CFFD792F296C9FDF8F916" author="Verrill A. E." box="[408,559,1744,1770]" pageId="38" pageNumber="39" pagination="259 - 446" refId="ref29377" refString="Verrill A. E. 1881. The Cephalopods of the Northeastern Coast of America. Part II. The smaller cephalopods, including the &quot; squids &quot; and the octopi, with other allied forms. Transactions of the Connecticut Academy of Sciences 5 (6): 259 - 446." type="journal article" year="1881">Verrill, 1881</bibRefCitation>
</taxonomicName>
, by subsequent designation.
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD3CFFD693D7970BFE5EFDE0" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8B803687BD3CFFD793D7970BFEE1F8D0" blockId="38.[189,310,1810,1836]" box="[189,310,1810,1836]" pageId="38" pageNumber="39">
<heading id="D0C881EBBD3CFFD793D7970BFEE1F8D0" bold="true" box="[189,310,1810,1836]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B94BEA95BD3CFFD793D7970BFEE1F8D0" bold="true" box="[189,310,1810,1836]" pageId="38" pageNumber="39">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD3CFFD693D79758FE16FE26" blockId="38.[189,1399,1857,2027]" lastBlockId="39.[189,1399,266,474]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<taxonomicName id="4C3F4D04BD3CFFD793D79758FEEEF8A7" authorityName="Leach" authorityYear="1817" box="[189,313,1857,1883]" class="Cephalopoda" family="Sepiolidae" kingdom="Animalia" order="Sepiolida" pageId="38" pageNumber="39" phylum="Mollusca" rank="subFamily" subFamily="Sepiolinae">Sepiolinae</taxonomicName>
with fins rounded; their length about half mantle length. Arm suckers in 4, exceptionally 8 longitudinal rows. Tentacle club with numerous minute suckers in a few tens of longitudinal rows. Mantle-head occipital band broad, extending over the ocular globes.A pair of kidney-shaped photophores on ventral surface of ink-sac. Gladius absent. Ventral mantle margin slightly sinuate, without any deep funnel indentation. Hectocotylus (male left arm I) thicker than right arm I, bipartite: proximal part, occupying 1⁄5 to ½ of arm, with regular suckers, except for 1–3 modified pedicels (may bear vestigial suckers), lengthened papilla-like, in 3
<superScript id="7C4A9BCFBD3DFFD6911D9137FD52FEC0" attach="left" box="[631,645,302,316]" fontSize="6" pageId="39" pageNumber="40">rd</superScript>
to 5
<superScript id="7C4A9BCFBD3DFFD691D09137FD10FEC0" attach="left" box="[698,711,302,316]" fontSize="6" pageId="39" pageNumber="40">th</superScript>
position of ventral sucker row; distalmost part, occupying ½ to *⁄5 of arm distally, with lengthened columnar sucker pedicels, closely packed to form longitudinal palisades, bearing at tip embedded suckers that are partially covered by fleshy caps, number of palisades proximally equal to that of regular sucker rows but reduced toward arm extremity. Female bursa copulatrix pouch-like.
</paragraph>
<paragraph id="8B803687BD3DFFD693D7921BFE5EFDE0" blockId="39.[189,393,514,540]" box="[189,393,514,540]" pageId="39" pageNumber="40">
<heading id="D0C881EBBD3DFFD693D7921BFE5EFDE0" bold="true" box="[189,393,514,540]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B94BEA95BD3DFFD693D7921BFE5EFDE0" bold="true" box="[189,393,514,540]" pageId="39" pageNumber="40">Included species</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD3DFFD693D7922BFE47FC44" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8B803687BD3DFFD693D7922BFDC5FD4B" blockId="39.[189,1399,561,695]" pageId="39" pageNumber="40">
<taxonomicName id="4C3F4D04BD3DFFD693D7922BFDEDFDB7" authority="(Verrill, 1881)" baseAuthorityName="Verrill" baseAuthorityYear="1881" box="[189,570,561,587]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="morsei">
<emphasis id="B94BEA95BD3DFFD693D7922BFE5BFDB7" box="[189,396,562,587]" italics="true" pageId="39" pageNumber="40">Euprymna morsei</emphasis>
(
<bibRefCitation id="EFAE4B76BD3DFFD692F39228FDE6FDB7" author="Verrill A. E." box="[409,561,561,587]" pageId="39" pageNumber="40" pagination="259 - 446" refId="ref29377" refString="Verrill A. E. 1881. The Cephalopods of the Northeastern Coast of America. Part II. The smaller cephalopods, including the &quot; squids &quot; and the octopi, with other allied forms. Transactions of the Connecticut Academy of Sciences 5 (6): 259 - 446." type="journal article" year="1881">Verrill, 1881</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD6912E922BFCA0FDB7" authority=", Voss, 1962" authorityName=", Voss" authorityYear="1962" box="[580,887,561,588]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="albatrossae">
<emphasis id="B94BEA95BD3DFFD6912E922BFD3DFDB7" box="[580,746,561,587]" italics="true" pageId="39" pageNumber="40">E. albatrossae</emphasis>
, Voss, 1962
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD690EB922BFB54FDB7" authority="Sasaki, 1929" authorityName="Sasaki" authorityYear="1929" box="[897,1155,561,587]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="berryi">
<emphasis id="B94BEA95BD3DFFD690EB922BFC3FFDB7" box="[897,1000,561,587]" italics="true" pageId="39" pageNumber="40">E. berryi</emphasis>
<bibRefCitation id="EFAE4B76BD3DFFD690879228FB54FDB7" author="Sasaki M." box="[1005,1155,561,587]" pageId="39" pageNumber="40" pagination="1 - 357" refId="ref28713" refString="Sasaki M. 1929. A monograph of the dibranchiate cephalopods of the Japanese and adjacent waters. Journal of the College of Agriculture, Hokkaido Imperial University 20 suppl: 1 - 357." type="journal article" year="1929">Sasaki, 1929</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD697E7922BFE90FD93" authority="Sanchez et al., 2019" authorityName="Sanchez" authorityYear="2019" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="brenneri">
<emphasis id="B94BEA95BD3DFFD697E7922BFAC7FDB7" box="[1165,1296,561,587]" italics="true" pageId="39" pageNumber="40">E. brenneri</emphasis>
<bibRefCitation id="EFAE4B76BD3DFFD6967F9228FE90FD93" author="Sanchez G. &amp; Jolly J. &amp; Reid A. &amp; Sugimoto C. &amp; Azama C. &amp; Marletaz F. &amp; Simakov O. &amp; Rokhsar D. S." pageId="39" pageNumber="40" pagination="465" refId="ref28636" refString="Sanchez G., Jolly J., Reid A., Sugimoto C., Azama C., Marletaz F., Simakov O. &amp; Rokhsar D. S. 2019. New bobtail squid (Sepiolidae: Sepiolinae) from the Ryukyu islands revealed by molecular and morphological analysis. Communications Biology 2: 465. https: // doi. org / 10.1038 / s 42003 - 019 - 0661 - 6" type="journal article" year="2019">
Sanchez
<emphasis id="B94BEA95BD3DFFD693D7924FFF2EFD93" box="[189,249,597,623]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
, 2019
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD6923E924FFD8DFD93" authority="Adam, 1986" authorityName="Adam" authorityYear="1986" box="[340,602,597,623]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="hoylei">
<emphasis id="B94BEA95BD3DFFD6923E924FFE69FD93" box="[340,446,597,623]" italics="true" pageId="39" pageNumber="40">E. hoylei</emphasis>
Adam, 1986
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD6910D924FFBD4FD93" authority=", Nateewathana, 1997" authorityName=", Nateewathana" authorityYear="1997" box="[615,1027,597,624]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="hyllebergi">
<emphasis id="B94BEA95BD3DFFD6910D924FFCD7FD93" box="[615,768,597,623]" italics="true" pageId="39" pageNumber="40">E. hyllebergi</emphasis>
,
<bibRefCitation id="EFAE4B76BD3DFFD69065924CFBD4FD93" author="Nateewathana A." box="[783,1027,597,623]" pageId="39" pageNumber="40" pagination="465 - 481" refId="ref28215" refString="Nateewathana A. 1997. The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description of Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication 17: 465 - 481." type="journal article" year="1997">Nateewathana, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD6977B924FFEBCFD6F" authority="Kubodera &amp; Okutani, 2002" authorityName="Kubodera &amp; Okutani" authorityYear="2002" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="megaspadicea">
<emphasis id="B94BEA95BD3DFFD6977B924FFB0CFD93" box="[1041,1243,597,623]" italics="true" pageId="39" pageNumber="40">E. megaspadicea</emphasis>
Kubodera &amp; Okutani, 2002
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD692139263FD49FD6F" authority="(Gray, 1849)" baseAuthorityName="Gray" baseAuthorityYear="1849" box="[377,670,633,659]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="penares">
<emphasis id="B94BEA95BD3DFFD692139263FE2EFD6F" box="[377,505,634,659]" italics="true" pageId="39" pageNumber="40">E. penares</emphasis>
(Gray, 1849)
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD691C69263FC19FD6F" authority="Berry, 1913" authorityName="Berry" authorityYear="1913" box="[684,974,633,659]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="scolopes">
<emphasis id="B94BEA95BD3DFFD691C69263FCE2FD6F" box="[684,821,633,659]" italics="true" pageId="39" pageNumber="40">E. scolopes</emphasis>
Berry, 1913
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD690B49263FA96FD6F" authority="(Grant, 1833)" baseAuthorityName="Grant" baseAuthorityYear="1833" box="[990,1345,633,659]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="stenodactyla">
<emphasis id="B94BEA95BD3DFFD690B49263FB42FD6F" box="[990,1173,633,659]" italics="true" pageId="39" pageNumber="40">E. stenodactyla</emphasis>
(Grant, 1833)
</taxonomicName>
and
<taxonomicName id="4C3F4D04BD3DFFD693D79287FDD9FD4B" authority="(Pfeffer, 1884)" baseAuthorityName="Pfeffer" baseAuthorityYear="1884" box="[189,526,669,695]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="tasmanica">
<emphasis id="B94BEA95BD3DFFD693D79287FE80FD4B" box="[189,343,670,695]" italics="true" pageId="39" pageNumber="40">E. tasmanica</emphasis>
(Pfeffer, 1884)
</taxonomicName>
.
</paragraph>
<paragraph id="8B803687BD3DFFD693D792C6FEF8FD05" blockId="39.[189,303,735,761]" box="[189,303,735,761]" pageId="39" pageNumber="40">
<heading id="D0C881EBBD3DFFD693D792C6FEF8FD05" bold="true" box="[189,303,735,761]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B94BEA95BD3DFFD693D792C6FEF8FD05" bold="true" box="[189,303,735,761]" pageId="39" pageNumber="40">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD3DFFD693D79317FE47FC44" blockId="39.[189,1399,782,952]" pageId="39" pageNumber="40">
Some nominal tetraseriate sucker species are deemed unresolved (cf.
<bibRefCitation id="EFAE4B76BD3DFFD690899316FB1FFCD4" author="Norman M. D. &amp; Lu C. C." box="[995,1224,782,809]" pageId="39" pageNumber="40" pagination="1109 - 1137" refId="ref28320" refString="Norman M. D. &amp; Lu C. C. 1997. Redescription of the southern dumpling squid Euprymna tasmanica and a revision of the genus Euprymna (Cephalopoda: Sepiolidae). Journal of the Marine Biological Association of the United Kingdom 77: 1109 - 1137. https: // doi. org / 10.1017 / S 0025315400038662" type="journal article" year="1997">Norman &amp; Lu 1997</bibRefCitation>
). The biseriate sucker arm
<taxonomicName id="4C3F4D04BD3DFFD69227932AFE62FCB0" authorityName="Bello" authorityYear="2020" baseAuthorityName="Sasaki" baseAuthorityYear="1914" box="[333,437,819,844]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="parva">
<emphasis id="B94BEA95BD3DFFD69227932AFE62FCB0" box="[333,437,819,844]" italics="true" pageId="39" pageNumber="40">E. parva</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD3DFFD692A2932AFD88FCB0" authorityName="Bello" authorityYear="2020" baseAuthorityName="Bello" baseAuthorityYear="2011" box="[456,607,818,844]" class="Cephalopoda" family="Sepiolidae" genus="Eumandya" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pardalota">
<emphasis id="B94BEA95BD3DFFD692A2932AFD88FCB0" box="[456,607,818,844]" italics="true" pageId="39" pageNumber="40">E. pardalota</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D04BD3DFFD691F5932AFCCFFCB0" authorityName="Bello" authorityYear="2020" baseAuthorityName="Voss" baseAuthorityYear="1962" box="[671,792,818,844]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="phenax">
<emphasis id="B94BEA95BD3DFFD691F5932AFCCFFCB0" box="[671,792,818,844]" italics="true" pageId="39" pageNumber="40">E. phenax</emphasis>
</taxonomicName>
, were removed from this genus to be placed into
<taxonomicName id="4C3F4D04BD3DFFD693D7934FFEEDFC8C" authority="Bello, 2020" authorityName="Bello" authorityYear="2020" box="[189,314,854,880]" class="Cephalopoda" family="Sepiolidae" genus="Eumandya" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B94BEA95BD3DFFD693D7934FFEEDFC8C" box="[189,314,854,880]" italics="true" pageId="39" pageNumber="40">Eumandya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857EEBD3DFFD6922F934FFE64FC8C" box="[325,435,854,880]" pageId="39" pageNumber="40" rank="genus">gen. nov.</taxonomicNameLabel>
(see above). The species in
<taxonomicName id="4C3F4D04BD3DFFD69065934EFC5EFC8C" authorityName="Steenstrup" authorityYear="1887" box="[783,905,855,880]" class="Cephalopoda" family="Sepiolidae" genus="Euprymna" kingdom="Animalia" order="Sepiolida" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD3DFFD69065934EFC5EFC8C" box="[783,905,855,880]" italics="true" pageId="39" pageNumber="40">Euprymna</emphasis>
</taxonomicName>
are distributed in tropical and temperate waters of the Indo-West Pacific region (
<bibRefCitation id="EFAE4B76BD3DFFD691FA9363FCAFFC68" author="Norman M. D. &amp; Lu C. C." box="[656,888,890,916]" pageId="39" pageNumber="40" pagination="1109 - 1137" refId="ref28320" refString="Norman M. D. &amp; Lu C. C. 1997. Redescription of the southern dumpling squid Euprymna tasmanica and a revision of the genus Euprymna (Cephalopoda: Sepiolidae). Journal of the Marine Biological Association of the United Kingdom 77: 1109 - 1137. https: // doi. org / 10.1017 / S 0025315400038662" type="journal article" year="1997">Norman &amp; Lu 1997</bibRefCitation>
;
<bibRefCitation id="EFAE4B76BD3DFFD690ED9363FBA7FC68" author="Nateewathana A." box="[903,1136,890,916]" pageId="39" pageNumber="40" pagination="465 - 481" refId="ref28215" refString="Nateewathana A. 1997. The Sepiolidae (Cephalopoda) of the Andaman Sea, Thailand, with description of Euprymna hyllebergi sp. nov. Phuket Marine Biological Center Special Publication 17: 465 - 481." type="journal article" year="1997">Nateewathana 1997</bibRefCitation>
; Kubodera &amp; Okutani 2002;
<bibRefCitation id="EFAE4B76BD3DFFD6926D9387FE57FC44" author="Reid A." box="[263,384,926,952]" pageId="39" pageNumber="40" pagination="135 - 142" refId="ref28431" refString="Reid A. 2011. Euprymna pardalota sp. nov. (Cephalopoda: Sepiolidae), a new dumpling squid from northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory 27: 135 - 142." type="journal article" year="2011">Reid 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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