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<document id="EEEAF92433BBC73794C9B4C3946AB45C" ID-CLB-Dataset="21713" ID-DOI="10.5852/ejt.2020.655" ID-GBIF-Dataset="fadee126-7da7-47d9-b9ac-973697ecf9f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3888797" ID-ZooBank="0042EFAE-2E4F-444B-AFB9-E321D16116E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591816327545" checkinUser="valdenar" docAuthor="Bello, Giambattista" docDate="2020" docId="03968791BD38FFD291CB96EBFF38FAFA" docLanguage="en" docName="ejt-655_bello.pdf.imf" docOrigin="European Journal of Taxonomy 655" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sepiola Leach 1817" docType="treatment" docVersion="6" lastPageNumber="36" masterDocId="FFAFFFE9BD1AFFF1936A9019FFD7FFFC" masterDocTitle="Evolution of the hectocotylus in Sepiolinae (Cephalopoda: Sepiolidae) and description of four new genera" masterLastPageNumber="53" masterPageNumber="1" pageNumber="35" updateTime="1698842281433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C6801127265A7C4B555C0130EAD67ABC">Evolution of the hectocotylus in Sepiolinae (Cephalopoda: Sepiolidae) and description of four new genera</mods:title>
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<mods:namePart id="C272CCD9ED002E0DCCC6070081E4682C">Bello, Giambattista</mods:namePart>
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<mods:title id="47BDFB9D9CD63934D483923D1905A14C">European Journal of Taxonomy</mods:title>
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<mods:date id="C55FA1ED958944F8A89656F01F712970">2020</mods:date>
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<mods:number id="36ECA367FD68698689AB80B6B2082E54">2020-05-29</mods:number>
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<treatment id="03968791BD38FFD291CB96EBFF38FAFA" ID-DOI="http://doi.org/10.5281/zenodo.5920268" ID-GBIF-Taxon="164500957" ID-Zenodo-Dep="5920268" LSID="urn:lsid:plazi:treatment:03968791BD38FFD291CB96EBFF38FAFA" httpUri="http://treatment.plazi.org/id/03968791BD38FFD291CB96EBFF38FAFA" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="C325650CBD38FFD391CB96EBFC44F8F0" box="[673,915,1778,1805]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8B803687BD38FFD391CB96EBFC44F8F0" blockId="34.[673,915,1778,1842]" box="[673,915,1778,1805]" pageId="34" pageNumber="35">
<heading id="D0C881EBBD38FFD391CB96EBFC44F8F0" box="[673,915,1778,1805]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
<taxonomicName id="4C3F4D04BD38FFD391CB96EBFC44F8F0" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[673,915,1778,1805]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="34" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD38FFD391CB96EBFD2CF8F1" bold="true" box="[673,763,1778,1805]" italics="true" pageId="34" pageNumber="35">Sepiola</emphasis>
Leach, 1817
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD38FFD391AA9701FCA3F8CE" box="[704,884,1816,1842]" pageId="34" pageNumber="35" type="description">
<paragraph id="8B803687BD38FFD391AA9701FCA3F8CE" blockId="34.[673,915,1778,1842]" box="[704,884,1816,1842]" pageId="34" pageNumber="35">
<figureCitation id="13042A02BD38FFD391AA9701FCC9F8CE" box="[704,798,1816,1842]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[189,1398,1032,1807]" captionTargetId="figure@8.[189,1399,1032,1807]" captionTargetPageId="8" captionText="Fig. 8. Basic types of bursae copulatrices in Sepiolinae Leach, 1817 (highlighted by white frames). A. Closed type, plesiomorphic condition; mated female displaying a bunch of spermatophores inserted in the bursa copulatrix and peeping out its opening (arrow) (Sepiola birostrata Sasaki, 1918). B. Open type, apomorphic condition; virgin female (Sepiola boletzkyi Bello &amp; Salman 2015). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3888813" httpUri="https://zenodo.org/record/3888813/files/figure.png" pageId="34" pageNumber="35">Figs 8B</figureCitation>
,
<figureCitation id="13042A02BD38FFD390409701FCA3F8CE" box="[810,884,1816,1842]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="25.[189,232,1860,1886]" captionStartId-1="26.[189,232,1965,1991]" captionTargetBox-0="[189,1398,1009,1803]" captionTargetBox-1="[601,979,1256,1892]" captionTargetId-0="figure@25.[189,1399,1009,1803]" captionTargetId-1="figure@26.[601,979,1256,1892]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="Fig. 19. Hectocotylus of Sepiola boletzkyi Bello &amp; Salman, 2015; a–b and c–d = modified ventral and dorsal pedicels, respectively, forming the copulatory apparatus proper; * = lobe in the dorsal sucker row; 1–11 = heteromorphous suckers in the dorsal row of distal part. A. Hectocotylus, oral-lateral view. B. Copulatory apparatus, mesial view. C. Copulatory apparatus, oblique mesial view. Scale bars: 1 mm (after Bello &amp; Salman 2015)." captionText-1="Fig. 20. Hectocotylus of Sepiola bursadhaesa Bello, 2013; mesial view. The square bracket encompasses the groove in the mesial lateral side (the arm in the foreground is the first right one). Scale bar: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3888837" figureDoi-1="http://doi.org/10.5281/zenodo.3888839" httpUri-0="https://zenodo.org/record/3888837/files/figure.png" httpUri-1="https://zenodo.org/record/3888839/files/figure.png" pageId="34" pageNumber="35">19–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD38FFD393D79745FE27F88A" box="[189,496,1884,1910]" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="8B803687BD38FFD393D79745FE27F88A" blockId="34.[189,496,1884,1910]" box="[189,496,1884,1910]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB2F11A9BD38FFD393D79745FE27F88A" box="[189,496,1884,1910]" pageId="34" pageNumber="35">
<treatmentCitation id="0A9E1096BD38FFD393D79745FE3BF88A" box="[189,492,1884,1910]" pageId="34" pageNumber="35">
<taxonomicName id="4C3F4D04BD38FFD393D79745FE3BF88A" authority="Leach, 1817: 137" authorityName="Leach" authorityPageNumber="137" authorityYear="1817" box="[189,492,1884,1910]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="34" pageNumber="35" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD38FFD393D79745FEC2F88A" box="[189,277,1884,1910]" italics="true" pageId="34" pageNumber="35">Sepiola</emphasis>
Leach, 1817: 137
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD38FFD393D797B9FD01F817" pageId="34" pageNumber="35" type="type_taxon">
<paragraph id="8B803687BD38FFD393D797B9FE8FF845" blockId="34.[189,344,1951,1978]" box="[189,344,1951,1978]" pageId="34" pageNumber="35">
<heading id="D0C881EBBD38FFD393D797B9FE8FF845" bold="true" box="[189,344,1951,1978]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B94BEA95BD38FFD393D797B9FE8FF845" bold="true" box="[189,344,1951,1978]" pageId="34" pageNumber="35">
<typeStatus id="54848825BD38FFD393D797B9FF2DF846" box="[189,250,1952,1978]" pageId="34" pageNumber="35">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD38FFD393D797C8FD01F817" blockId="34.[189,726,2001,2027]" box="[189,726,2001,2027]" pageId="34" pageNumber="35">
<taxonomicName id="4C3F4D04BD38FFD393D797C8FDFFF817" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[189,552,2001,2027]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="rondeletii">
<emphasis id="B94BEA95BD38FFD393D797C8FE5AF817" box="[189,397,2001,2027]" italics="true" pageId="34" pageNumber="35">Sepiola rondeletii</emphasis>
Leach, 1817
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD39FFD293D79112FE57FD56" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8B803687BD39FFD293D79112FEE1FED9" blockId="35.[189,310,267,293]" box="[189,310,267,293]" pageId="35" pageNumber="36">
<heading id="D0C881EBBD39FFD293D79112FEE1FED9" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B94BEA95BD39FFD293D79112FEE1FED9" bold="true" box="[189,310,267,293]" pageId="35" pageNumber="36">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD39FFD293D79125FE57FD56" blockId="35.[189,1399,316,682]" pageId="35" pageNumber="36">
<taxonomicName id="4C3F4D04BD39FFD293D79125FEEBFEAA" authorityName="Leach" authorityYear="1817" box="[189,316,316,342]" class="Cephalopoda" family="Sepiolidae" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="subFamily" subFamily="Sepiolinae">Sepiolinae</taxonomicName>
with fins rounded; their length about half mantle length. Suckers biseriate on arms I to III and at least proximally on arm IV. Tentacle club suckers in 4 to 8 longitudinal rows. Mantle-head occipital band narrow (not reaching over the ocular globes). A pair of kidney-shaped photophores on ventral surface of ink-sac. Gladius present, reduced. Ventral mantle margin slightly sinuate, without any deep funnel indentation. Hectocotylus (male left arm I) tripartite: basal part with three suckers, two ventral and one dorsal; copulatory apparatus with transverse formation of two ventral and two dorsal modified suckerless pedicels, all four of them lengthened and fused with each other throughout their length (few additional dorsal modified pedicels may be present mesially to transverse formation); distal part with some enlarged suckers in dorsal row. Female bursa copulatrix roughly ear-shaped, devoid of cover.
</paragraph>
</subSubSection>
<subSubSection id="C325650CBD39FFD293D792CDFF38FAFA" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8B803687BD39FFD293D792CDFE5EFD12" blockId="35.[189,393,724,750]" box="[189,393,724,750]" pageId="35" pageNumber="36">
<heading id="D0C881EBBD39FFD293D792CDFE5EFD12" bold="true" box="[189,393,724,750]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B94BEA95BD39FFD293D792CDFE5EFD12" bold="true" box="[189,393,724,750]" pageId="35" pageNumber="36">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD39FFD293D7931CFC18FC97" blockId="35.[189,1399,773,875]" pageId="35" pageNumber="36">
<taxonomicName id="4C3F4D04BD39FFD293D7931CFDF9FCE3" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[189,558,773,799]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="rondeletii">
<emphasis id="B94BEA95BD39FFD293D7931CFE58FCE3" box="[189,399,773,799]" italics="true" pageId="35" pageNumber="36">Sepiola rondeletii</emphasis>
Leach, 1817
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD29156931CFCE4FCE3" authority="Naef, 1912" authorityName="Naef" authorityYear="1912" box="[572,819,773,799]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="affinis">
<emphasis id="B94BEA95BD39FFD29156931CFD74FCE3" box="[572,675,773,799]" italics="true" pageId="35" pageNumber="36">S. affinis</emphasis>
Naef, 1912
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD2902B931CFB4FFCE3" authority="d'Orbigny, 1842" authorityName="d'Orbigny" authorityYear="1842" box="[833,1176,773,799]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="atlantica">
<emphasis id="B94BEA95BD39FFD2902B931CFC1FFCE3" box="[833,968,773,799]" italics="true" pageId="35" pageNumber="36">S. atlantica</emphasis>
d’Orbigny, 1842
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD297CC931CFE5AFCBA" authority="Bello &amp; Salman, 2015" authorityName="Bello &amp; Salman" authorityYear="2015" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="boletzkyi">
<emphasis id="B94BEA95BD39FFD297CC931CFAFAFCE3" box="[1190,1325,773,799]" italics="true" pageId="35" pageNumber="36">S. boletzkyi</emphasis>
Bello &amp; Salman, 2015
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD292F59332FD2EFCB9" authority="Bello, 2013" authorityName="Bello" authorityYear="2013" box="[415,761,811,837]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="bursadhaesa">
<emphasis id="B94BEA95BD39FFD292F59332FD8EFCB9" box="[415,601,811,837]" italics="true" pageId="35" pageNumber="36">S. bursadhaesa</emphasis>
Bello, 2013
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD290669332FB9FFCB9" authority="Naef, 1912" authorityName="Naef" authorityYear="1912" box="[780,1096,811,837]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="intermedia">
<emphasis id="B94BEA95BD39FFD290669332FC78FCB9" box="[780,943,811,837]" italics="true" pageId="35" pageNumber="36">S. intermedia</emphasis>
Naef, 1912
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD297369332FAA6FCB9" authority="Naef, 1912" authorityName="Naef" authorityYear="1912" box="[1116,1393,811,837]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="robusta">
<emphasis id="B94BEA95BD39FFD297369332FB0EFCB9" box="[1116,1241,811,837]" italics="true" pageId="35" pageNumber="36">S. robusta</emphasis>
Naef, 1912
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD293D79348FDC5FC97" authority="Levy, 1912" authorityName="Levy" authorityYear="1912" box="[189,530,849,875]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="steenstrupiana">
<emphasis id="B94BEA95BD39FFD293D79348FE50FC96" box="[189,391,849,875]" italics="true" pageId="35" pageNumber="36">S. steenstrupiana</emphasis>
Levy, 1912
</taxonomicName>
and
<taxonomicName id="4C3F4D04BD39FFD291219348FC1DFC97" authority="de Heij &amp; Goud, 2010" authorityName="de Heij &amp; Goud" authorityYear="2010" box="[587,970,848,875]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="tridens">
<emphasis id="B94BEA95BD39FFD291219348FD6EFC97" box="[587,697,849,875]" italics="true" pageId="35" pageNumber="36">S. tridens</emphasis>
de Heij &amp; Goud, 2010
</taxonomicName>
.
</paragraph>
<paragraph id="8B803687BD39FFD293D7938CFEF8FC53" blockId="35.[189,303,917,943]" box="[189,303,917,943]" pageId="35" pageNumber="36">
<heading id="D0C881EBBD39FFD293D7938CFEF8FC53" bold="true" box="[189,303,917,943]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B94BEA95BD39FFD293D7938CFEF8FC53" bold="true" box="[189,303,917,943]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B803687BD39FFD293D793DFFB64FB61" blockId="35.[189,1398,966,1181]" pageId="35" pageNumber="36">
The revised diagnosis of the genus
<taxonomicName id="4C3F4D04BD39FFD2911F93DFFD18FC1C" authorityName="Leach" authorityYear="1817" box="[629,719,966,992]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD39FFD2911F93DFFD18FC1C" box="[629,719,966,992]" italics="true" pageId="35" pageNumber="36">Sepiola</emphasis>
</taxonomicName>
reported above was formulated to comply with the rearrangement of
<taxonomicName id="4C3F4D04BD39FFD292EB93F2FDD7FBF9" authorityName="Leach" authorityYear="1817" box="[385,512,1003,1029]" class="Cephalopoda" family="Sepiolidae" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="subFamily" subFamily="Sepiolinae">Sepiolinae</taxonomicName>
according to the present results. It dismisses several species previously included in
<taxonomicName id="4C3F4D04BD39FFD2923E9408FE79FBD7" authorityName="Leach" authorityYear="1817" box="[340,430,1041,1067]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD39FFD2923E9408FE79FBD7" box="[340,430,1041,1067]" italics="true" pageId="35" pageNumber="36">Sepiola</emphasis>
</taxonomicName>
, namely
<taxonomicName id="4C3F4D04BD39FFD2914D940BFD19FBD7" authorityName="Jatta" authorityYear="1896" box="[551,718,1042,1068]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="aurantiaca">
<emphasis id="B94BEA95BD39FFD2914D940BFD19FBD7" box="[551,718,1042,1068]" italics="true" pageId="35" pageNumber="36">S. aurantiaca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD2918B940BFCACFBD7" authorityName="Sasaki" authorityYear="1918" box="[737,891,1041,1068]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="birostrata">
<emphasis id="B94BEA95BD39FFD2918B940BFCACFBD7" box="[737,891,1041,1068]" italics="true" pageId="35" pageNumber="36">S. birostrata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD290E5940BFBCAFBD7" authorityName="Adam" authorityYear="1984" box="[911,1053,1041,1068]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="knudseni">
<emphasis id="B94BEA95BD39FFD290E5940BFBCAFBD7" box="[911,1053,1041,1068]" italics="true" pageId="35" pageNumber="36">S. knudseni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD2975B940BFB65FBD7" authorityName="Naef" authorityYear="1912" box="[1073,1202,1041,1068]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="ligulata">
<emphasis id="B94BEA95BD39FFD2975B940BFB65FBD7" box="[1073,1202,1041,1068]" italics="true" pageId="35" pageNumber="36">S. ligulata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D04BD39FFD297AF940BFAE8FBD7" authorityName="Grimpe" authorityYear="1921" box="[1221,1343,1041,1068]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="pfefferi">
<emphasis id="B94BEA95BD39FFD297AF940BFAE8FBD7" box="[1221,1343,1041,1068]" italics="true" pageId="35" pageNumber="36">S. pfefferi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D04BD39FFD293D7942EFE8FFBAD" authorityName="in Voss" authorityYear="1963" box="[189,344,1079,1105]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="trirostrata">
<emphasis id="B94BEA95BD39FFD293D7942EFE8FFBAD" box="[189,344,1079,1105]" italics="true" pageId="35" pageNumber="36">S. trirostrata</emphasis>
</taxonomicName>
, which are better placed in different genera. The revised genus
<taxonomicName id="4C3F4D04BD39FFD29737942EFB60FBAD" authorityName="Leach" authorityYear="1817" box="[1117,1207,1079,1105]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD39FFD29737942EFB60FBAD" box="[1117,1207,1079,1105]" italics="true" pageId="35" pageNumber="36">Sepiola</emphasis>
</taxonomicName>
includes all the species in the
<taxonomicName id="4C3F4D04BD39FFD2920D9444FDE3FB8B" authorityName="d'Orbigny" authorityYear="1842" box="[359,564,1117,1143]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="atlantica">
<emphasis id="B94BEA95BD39FFD2920D9444FDE3FB8B" box="[359,564,1117,1143]" italics="true" pageId="35" pageNumber="36">Sepiola atlantica</emphasis>
</taxonomicName>
-group
<emphasis id="B94BEA95BD39FFD291E19447FD19FB8B" box="[651,718,1118,1143]" italics="true" pageId="35" pageNumber="36">sensu</emphasis>
<bibRefCitation id="EFAE4B76BD39FFD291BC9444FC83FB8B" author="Naef A." box="[726,852,1117,1143]" pageId="35" pageNumber="36" pagination="1 - 863" refId="ref28169" refString="Naef A. 1923. Die Cephalopoden. Fauna und Flora des Golfes von Neapel 35 (1): 1 - 863. [English translation by Mercado A. 1972; Smithsonian Institution, Washington D. C.: 1 - 917]" type="journal article" year="1923">Naef 1923</bibRefCitation>
(see also Bello 2013; Bello &amp; Salman 2015). Hence, the
<taxonomicName id="4C3F4D04BD39FFD2922F949AFE67FB61" authorityName="d'Orbigny" authorityYear="1842" box="[325,432,1155,1181]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="atlantica">
<emphasis id="B94BEA95BD39FFD2922F949AFE67FB61" box="[325,432,1155,1181]" italics="true" pageId="35" pageNumber="36">atlantica</emphasis>
</taxonomicName>
-group coincides with the genus
<taxonomicName id="4C3F4D04BD39FFD2905D949AFC46FB61" authorityName="Leach" authorityYear="1817" box="[823,913,1155,1181]" class="Cephalopoda" family="Sepiolidae" genus="Sepiola" kingdom="Animalia" order="Sepiolida" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
<emphasis id="B94BEA95BD39FFD2905D949AFC46FB61" box="[823,913,1155,1181]" italics="true" pageId="35" pageNumber="36">Sepiola</emphasis>
</taxonomicName>
as revised in this paper.
</paragraph>
<paragraph id="8B803687BD39FFD293D794DFFF38FAFA" blockId="35.[189,1398,1222,1286]" pageId="35" pageNumber="36">The overall distribution of these species ranges from the north-eastern Atlantic Ocean to the Mediterranean Sea.</paragraph>
</subSubSection>
</treatment>
</document>
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