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<document id="4AE18068366B0459D87D07D6A89E8962" ID-DOI="10.5281/zenodo.187516" ID-GBIF-Dataset="aa61272c-ae6b-4bc6-89f8-19b912abd2eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="187516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461071548377" checkinUser="plazi" docAuthor="De, Willem H." docDate="2009" docId="953D87D5FFC0FF90FF3BE8E4ACB6FDF2" docLanguage="en" docName="zt02092p020.pdf" docOrigin="Zootaxa 2092" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Testudinella clypeata Muller 1786" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="6904FFADFFCAFF9CFFACE942AF65FFA9" masterDocTitle="A review of the marine and brackish-water species of Testudinella (Rotifera: Monogononta, Testudinellidae), with the description of two new species" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1698584922083" updateUser="plazi">
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<mods:title id="0CF9F3F4649FCD4CCCA8B1EEF2B85A07">A review of the marine and brackish-water species of Testudinella (Rotifera: Monogononta, Testudinellidae), with the description of two new species</mods:title>
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<mods:namePart id="3EF413B3F9CC6E980C95CF2221EB2004">De, Willem H.</mods:namePart>
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<mods:typeOfResource id="490B33EC0E7D9C47DECD93590FE5A69C">text</mods:typeOfResource>
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<mods:date id="74F47A5D1A04B222542387BB4076BDD9">2009</mods:date>
<mods:detail id="95388E07D8585C5E36B27B16D20F87C3" type="volume">
<mods:number id="80434F6EBCC9F958805B1CA8B0C5C361">2092</mods:number>
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<mods:classification id="2747C71FC841D90FE2CC0630695B6DEA">journal article</mods:classification>
<mods:identifier id="4705F85DF28412B7EC1541ABFE68F887" type="DOI">10.5281/zenodo.187516</mods:identifier>
<mods:identifier id="DB6A459A80E883379CEC9A7D9022DE9F" type="GBIF-Dataset">aa61272c-ae6b-4bc6-89f8-19b912abd2eb</mods:identifier>
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<treatment id="953D87D5FFC0FF90FF3BE8E4ACB6FDF2" ID-DOI="http://doi.org/10.5281/zenodo.6215591" ID-GBIF-Taxon="119616006" ID-Zenodo-Dep="6215591" LSID="urn:lsid:plazi:treatment:953D87D5FFC0FF90FF3BE8E4ACB6FDF2" httpUri="http://treatment.plazi.org/id/953D87D5FFC0FF90FF3BE8E4ACB6FDF2" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="558E6548FFC0FF96FF3BE8E4AE7AFE4A" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="1D2B36C3FFC0FF96FF3BE8E4AD2EFE69" blockId="10.[151,587,422,483]" box="[151,587,422,448]" pageId="10" pageNumber="11">
<heading id="466381AFFFC0FF96FF3BE8E4AD2EFE69" bold="true" box="[151,587,422,448]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="DA944D40FFC0FF96FF3BE8E4AD2EFE69" ID-CoL="55JM9" authority="Muller, 1786" authorityName="Muller" authorityYear="1786" box="[151,587,422,448]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FF3BE8E4AD2EFE69" bold="true" box="[151,587,422,448]" pageId="10" pageNumber="11">
<emphasis id="2FE0EAD1FFC0FF96FF3BE8E4AEF1FE69" bold="true" box="[151,404,422,448]" italics="true" pageId="10" pageNumber="11">Testudinella clypeata</emphasis>
(Müller, 1786)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="1D2B36C3FFC0FF96FF3BE88BAE7AFE4A" blockId="10.[151,587,422,483]" box="[151,287,457,483]" pageId="10" pageNumber="11">
(
<figureCitation id="85AF2A46FFC0FF96FF33E88BAE73FE4A" box="[159,278,457,483]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="10.[151,262,1413,1437]" captionStartId-1="11.[151,255,1057,1081]" captionStartId-2="11.[151,257,1970,1994]" captionTargetBox-0="[158,1427,783,1373]" captionTargetBox-1="[249,1337,310,1035]" captionTargetBox-2="[476,1110,1135,1969]" captionTargetId-0="figure@10.[151,1436,772,1406]" captionTargetId-1="figure@11.[249,1338,310,1051]" captionTargetId-2="figure@11.[476,1111,1135,2042]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionText-0="FIGURE 8. Testudinella clypeata. A – C. Loricae, ventral view. D – F. Loricae, cross-sectional view. G. Foot pseudosegments. Scale bars: A – F: 50 µm, G: 25 µm." captionText-1="FIGURE 9. Testudinella clypeata, variation in shape of specimen. A, B. Contracted. C. Swimming. Scale bar: 50 µm." captionText-2="FIGURE 10. Testudinella clypeata, scanning electron microscope photographs of complete set of trophi. A. Frontal view. B. Caudal view. Scale bar: 10 µm. fo: fulcrum opening, sc: sub-ventral chamber." httpUri-0="https://zenodo.org/record/187524/files/figure.png" httpUri-1="https://zenodo.org/record/187525/files/figure.png" httpUri-2="https://zenodo.org/record/187526/files/figure.png" pageId="10" pageNumber="11">Figs 8–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="558E6548FFC0FF96FF3BEB54ADD8F823" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="1D2B36C3FFC0FF96FF3BEB54AE84FD24" blockId="10.[151,1436,534,717]" pageId="10" pageNumber="11">
<taxonomicName id="DA944D40FFC0FF96FF3BEB54AE36FD84" box="[151,339,534,557]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="subSpecies" species="clypeata" subSpecies="typica">
<emphasis id="2FE0EAD1FFC0FF96FF3BEB54AE36FD84" box="[151,339,534,557]" italics="true" pageId="10" pageNumber="11">T. clypeata typica</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FE3DEB54AD7FFD84" author="Voigt" box="[401,538,534,557]" pageId="10" pageNumber="20" refString="Voigt, M. (1957) Rotatoria. Die Radertiere Mitteleuropas. Gebruder Borntraeger, Berlin-Nikolassee, I. Textband, 508 pp., II. Tafelband, 115 pls." type="book" year="1957">Voigt (1957)</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FD89EB54ADF2FD84" ID-CoL="7C3D9" box="[549,663,534,557]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="elliptica">
<emphasis id="2FE0EAD1FFC0FF96FD89EB54ADF2FD84" box="[549,663,534,557]" italics="true" pageId="10" pageNumber="11">T. elliptica</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FD79EB54ACD1FD84" author="Jersabek" box="[725,948,534,557]" pageId="10" pageNumber="20" refString="Jersabek, C. D., Segers, H. &amp; Morris, P. J. (2003) An illustrated online catalog of the Rotifera in the Academy of Natural Sciences of Philadelphia (version 1.0: 2003 - April- 8). [WWW database] URL, http: // rotifer. acnatsci. org / rotifer. php" type="book chapter" year="2003">
Jersabek
<emphasis id="2FE0EAD1FFC0FF96FC94EB54AC0BFD84" box="[824,878,534,557]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2003
</bibRefCitation>
, ANSP Catalog Number 1828)? non
<taxonomicName id="DA944D40FFC0FF96FF72EB74AE30FDE4" ID-CoL="55JM9" box="[222,341,566,589]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FF72EB74AE30FDE4" box="[222,341,566,589]" italics="true" pageId="10" pageNumber="11">T. clypeata</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FE36EB74AD30FDE4" author="Rousselet" box="[410,597,566,589]" pageId="10" pageNumber="20" refString="Rousselet, C. F. (1898) Notes on some little-known species of Pterodina. The Journal of the Quekett Microscopical Club, 2 nd Ser., 7, 24 - 30, pls 3 - 5." type="journal article" year="1898">Rousselet (1898)</bibRefCitation>
,
<bibRefCitation id="79054B32FFC0FF96FDCFEB74ACCEFDE4" author="Dieffenbach" box="[611,939,566,589]" pageId="10" pageNumber="19" refString="Dieffenbach, H. &amp; Sachse, R. (1912) Rotatoria. In: Brauer, A. (Ed.), Die Susswasserfauna Deutschlands, Heft 14: Rotatoria, Gastrotricha. Verlag von Gustav Fischer, Jena, pp. 1 - 239." type="journal article" year="1912">Dieffenbach &amp; Sachse (1912)</bibRefCitation>
and
<bibRefCitation id="79054B32FFC0FF96FC48EB74AB8AFDE4" author="Bartos" box="[996,1263,566,589]" pageId="10" pageNumber="19" refString="Bartos, E. (1951) Czechoslovakian species of genera Testudinella and Pompholyx. (Rot.). Sbornik Klubu p r irodov e deckeho v Brne, 29, 10 - 20. (In Czechoslovak)." type="journal article" year="1951">
Bartoš (1951,
<figureCitation id="85AF2A46FFC0FF96FB2EEB74ABACFDE4" box="[1154,1225,566,589]" captionStart="FIGURE 7" captionStartId="9.[151,255,1747,1771]" captionTargetBox="[186,1399,700,1746]" captionTargetId="figure@9.[186,1401,700,1834]" captionTargetPageId="9" captionText="FIGURE 7. Testudinella obscura, scanning electron microscope photographs of trophi. A. Complete set, frontal view. B. Detail left manubrium. C. Complete set, caudal view. D. Complete set, lateral view. Scale bars: 10 µm. a: alula, dc: dorsal chamber, fo: fulcrum opening, mc: median chamber, sc: sub-ventral chamber, vc: ventral chamber." httpUri="https://zenodo.org/record/187523/files/figure.png" pageId="10" pageNumber="11">Figs 7</figureCitation>
,
<figureCitation id="85AF2A46FFC0FF96FB74EB74AB83FDE4" box="[1240,1254,566,589]" captionStart="FIGURE 8" captionStartId="10.[151,262,1413,1437]" captionTargetBox="[158,1427,783,1373]" captionTargetId="figure@10.[151,1436,772,1406]" captionTargetPageId="10" captionText="FIGURE 8. Testudinella clypeata. A – C. Loricae, ventral view. D – F. Loricae, cross-sectional view. G. Foot pseudosegments. Scale bars: A – F: 50 µm, G: 25 µm." httpUri="https://zenodo.org/record/187524/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
)
</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FB52EB74AA05FDE4" ID-CoL="4PWGR" box="[1278,1376,566,589]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="crassa">
<emphasis id="2FE0EAD1FFC0FF96FB52EB74AA05FDE4" box="[1278,1376,566,589]" italics="true" pageId="10" pageNumber="11">T. crassa</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FF6BEB14AE0FFDC4" author="Harring" box="[199,362,598,621]" pageId="10" pageNumber="20" refString="Harring, H. K. (1913) Synopsis of the Rotatoria. Smithsonian Institution United States National Museum Bulletin, 81, 226 pp." type="book" year="1913">Harring (1913)</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FED9EB14AE8FFDC4" ID-CoL="55JM9" box="[373,490,598,621]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FED9EB14AE8FFDC4" box="[373,490,598,621]" italics="true" pageId="10" pageNumber="11">T. clypeata</emphasis>
</taxonomicName>
<taxonomicName id="DA944D40FFC0FF96FE5DEB14AD01FDC4" box="[497,612,598,621]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="variety" species="clypeata" variety="crassa">
var.
<emphasis id="2FE0EAD1FFC0FF96FDB3EB14AD01FDC4" box="[543,612,598,621]" italics="true" pageId="10" pageNumber="11">crassa</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FD08EB14AC2FFDC4" author="Remane" box="[676,842,598,621]" pageId="10" pageNumber="20" refString="Remane, A. (1929) Rotatoria. In: Grimpe, G. (Ed.), Die Tierwelt der Nord- und Ostsee, [L. 16, T. 7. e.], Leipzig: Akademische Verlagsgesellschaft m. b. H., pp. 1 - 156." type="journal article" year="1929">Remane (1929)</bibRefCitation>
and
<bibRefCitation id="79054B32FFC0FF96FCD2EB14AB6DFDC4" author="Voigt" box="[894,1032,598,621]" pageId="10" pageNumber="20" refString="Voigt, M. (1957) Rotatoria. Die Radertiere Mitteleuropas. Gebruder Borntraeger, Berlin-Nikolassee, I. Textband, 508 pp., II. Tafelband, 115 pls." type="book" year="1957">Voigt (1957)</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FBBFEB14ABB0FDC4" box="[1043,1237,598,621]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="subSpecies" species="clypeata" subSpecies="crassa">
<emphasis id="2FE0EAD1FFC0FF96FBBFEB14ABB0FDC4" box="[1043,1237,598,621]" italics="true" pageId="10" pageNumber="11">T. clypeata crassa</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FABFEB14AE6BFD24" author="Wiszniewski" pageId="10" pageNumber="20" refString="Wiszniewski, J. (1954) Materiaux relatifs a la nomenclature et a la bibliographie des rotiferes. Polskie Archiwum Hydrobiologii, 2, 1 - 249." type="journal article" year="1954">Wiszniewski (1954)</bibRefCitation>
and
<bibRefCitation id="79054B32FFC0FF96FEEEEB34AE84FD24" author="Berzins" box="[322,481,630,653]" pageId="10" pageNumber="19" refString="Berzins, B. (1960) Rotatoria VI, order Monogononta. Conseil International pour l'Exploration de la Mer: Zooplankton, Sheet 89, 4 pp." type="book" year="1960">Bērziņš (1960)</bibRefCitation>
</paragraph>
<paragraph id="1D2B36C3FFC0FF96FF3BEBD4AE36FD64" blockId="10.[151,1436,534,717]" pageId="10" pageNumber="11">
non
<taxonomicName id="DA944D40FFC0FF96FF69EBD4AE5DFD04" ID-CoL="55JM9" box="[197,312,662,685]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FF69EBD4AE5DFD04" box="[197,312,662,685]" italics="true" pageId="10" pageNumber="11">T. clypeata</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FEDAEBD4AD16FD04" author="Bartos" box="[374,627,662,685]" pageId="10" pageNumber="19" refString="Bartos, E. (1951) Czechoslovakian species of genera Testudinella and Pompholyx. (Rot.). Sbornik Klubu p r irodov e deckeho v Brne, 29, 10 - 20. (In Czechoslovak)." type="journal article" year="1951">
Bartoš (1951,
<figureCitation id="85AF2A46FFC0FF96FDA0EBD4AD2AFD04" box="[524,591,662,685]" captionStart="FIGURE 3" captionStartId="3.[151,255,1512,1536]" captionTargetBox="[151,1434,387,1490]" captionTargetId="figure@3.[151,1436,387,1562]" captionTargetPageId="3" captionText="FIGURE 3. Testudinella bicorniculata sp. nov., scanning electron microscope photographs of trophi. A. Complete set, frontal view. B. Complete set, caudal view. C. Complete set, lateral view. D. Detail proximal fulcrum opening and basal apophyses. E. Detail left ramus with arched scleropili. Scale bars 10 µm. a: alula, as: arched scleropili, ba: basal apophyses, dc: dorsal chamber, fo: fulcrum opening, mc: median chamber, sc: sub-ventral chamber, vc: ventral chamber" httpUri="https://zenodo.org/record/187519/files/figure.png" pageId="10" pageNumber="11">Figs 3</figureCitation>
,
<figureCitation id="85AF2A46FFC0FF96FDF6EBD4AD0CFD04" box="[602,617,662,685]" captionStart="FIGURE 4" captionStartId="5.[151,256,1291,1315]" captionTargetBox="[485,1090,406,1245]" captionTargetId="figure@5.[476,1111,390,1284]" captionTargetPageId="5" captionText="FIGURE 4. Testudinella elongata sp. nov. A. Lorica, dorsal view. B. Lorica, ventral view. C. Lorica, cross-sectional view. D. Foot pseudosegments. Scale bars A – C: 50 µm, D: 25 µm." httpUri="https://zenodo.org/record/187520/files/figure.png" pageId="10" pageNumber="11">4</figureCitation>
)
</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FDD1EBD4AC22FD04" ID-CoL="55JMB" box="[637,839,662,685]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="subSpecies" species="clypeata" subSpecies="dentata">
<emphasis id="2FE0EAD1FFC0FF96FDD1EBD4AC22FD04" box="[637,839,662,685]" italics="true" pageId="10" pageNumber="11">T. clypeata dentata</emphasis>
</taxonomicName>
after
<bibRefCitation id="79054B32FFC0FF96FC28EBD4AB39FD04" author="Wiszniewski" box="[900,1116,662,685]" pageId="10" pageNumber="20" refString="Wiszniewski, J. (1954) Materiaux relatifs a la nomenclature et a la bibliographie des rotiferes. Polskie Archiwum Hydrobiologii, 2, 1 - 249." type="journal article" year="1954">Wiszniewski (1954)</bibRefCitation>
,
<taxonomicName id="DA944D40FFC0FF96FBCAEBD4ABBFFD04" ID-CoL="55JM9" box="[1126,1242,662,685]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FBCAEBD4ABBFFD04" box="[1126,1242,662,685]" italics="true" pageId="10" pageNumber="11">T. clypeata</emphasis>
</taxonomicName>
f.
<taxonomicName id="DA944D40FFC0FF96FB5AEBD4AE36FD64" ID-CoL="55JN6" authority="Koste (1978)" authorityName="Koste" authorityYear="1978" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="pseudoclypeata">
<emphasis id="2FE0EAD1FFC0FF96FB5AEBD4AAF9FD04" box="[1270,1436,662,685]" italics="true" pageId="10" pageNumber="11">pseudoclypeata</emphasis>
<bibRefCitation id="79054B32FFC0FF96FF6BEBF4AE36FD64" author="Koste" box="[199,339,694,717]" pageId="10" pageNumber="20" refString="Koste, W. (1978) Rotatoria. Die Radertiere Mitteleuropas begrundet von Max Voigt. Monogononta. Gebruder Borntraeger, Berlin, Stuttgart, I. Textband, 673 pp., II. Tafelband, 234 pls." type="book" year="1978">Koste (1978)</bibRefCitation>
</taxonomicName>
</paragraph>
<caption id="49EB664BFFC0FF96FF3BECC7ADA9FA16" httpUri="https://zenodo.org/record/187524/files/figure.png" pageId="10" pageNumber="11" targetBox="[158,1427,783,1373]" targetPageId="10">
<paragraph id="1D2B36C3FFC0FF96FF3BECC7ADA9FA16" blockId="10.[151,1437,1413,1471]" pageId="10" pageNumber="11">
<emphasis id="2FE0EAD1FFC0FF96FF3BECC7AE40FA34" bold="true" box="[151,293,1413,1437]" pageId="10" pageNumber="11">FIGURE 8.</emphasis>
<taxonomicName id="DA944D40FFC0FF96FE82ECC7AD49FA35" ID-CoL="55JM9" box="[302,556,1413,1436]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="10" pageNumber="11" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC0FF96FE82ECC7AD49FA35" box="[302,556,1413,1436]" italics="true" pageId="10" pageNumber="11">Testudinella clypeata</emphasis>
</taxonomicName>
.
<emphasis id="2FE0EAD1FFC0FF96FD91ECC7AD19FA34" bold="true" box="[573,636,1413,1437]" pageId="10" pageNumber="11">A–C.</emphasis>
Loricae, ventral view.
<emphasis id="2FE0EAD1FFC0FF96FC38ECC7ACABFA34" bold="true" box="[916,974,1413,1437]" pageId="10" pageNumber="11">D–F.</emphasis>
Loricae, cross-sectional view.
<emphasis id="2FE0EAD1FFC0FF96FAE8ECC7AA3FFA34" bold="true" box="[1348,1370,1413,1437]" pageId="10" pageNumber="11">G.</emphasis>
Foot pseudosegments. Scale bars: A–F: 50 µm, G: 25 µm.
</paragraph>
</caption>
<paragraph id="1D2B36C3FFC0FF96FF3BECAFADD8F823" blockId="10.[151,1437,1517,2007]" pageId="10" pageNumber="11">
This species is easily recognized by the elliptical lorica (length/width ratio 1.30–1.55, average 1.45), which is often slightly angular postero-laterally (
<figureCitation id="85AF2A46FFC0FF96FD2CEF56ADA9F987" box="[640,716,1556,1582]" captionStart="FIGURE 8" captionStartId="10.[151,262,1413,1437]" captionTargetBox="[158,1427,783,1373]" captionTargetId="figure@10.[151,1436,772,1406]" captionTargetPageId="10" captionText="FIGURE 8. Testudinella clypeata. A – C. Loricae, ventral view. D – F. Loricae, cross-sectional view. G. Foot pseudosegments. Scale bars: A – F: 50 µm, G: 25 µm." httpUri="https://zenodo.org/record/187524/files/figure.png" pageId="10" pageNumber="11">Figs 8</figureCitation>
A, 9C) apparently depending on the state of contraction or fixation (
<figureCitation id="85AF2A46FFC0FF96FEABEF78AE31F9FD" box="[263,340,1594,1620]" captionStart="FIGURE 9" captionStartId="11.[151,255,1057,1081]" captionTargetBox="[249,1337,310,1035]" captionTargetId="figure@11.[249,1338,310,1051]" captionTargetPageId="11" captionText="FIGURE 9. Testudinella clypeata, variation in shape of specimen. A, B. Contracted. C. Swimming. Scale bar: 50 µm." httpUri="https://zenodo.org/record/187525/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
). The dorsal anterior margin is convex, and the ventral anterior margin shows a broad Vshaped sinus; a dorsal transversal fold is often present at the end of the lorica. In cross-sectional view (
<figureCitation id="85AF2A46FFC0FF96FAC0EF23AFC1F90B" captionStart="FIGURE 8" captionStartId="10.[151,262,1413,1437]" captionTargetBox="[158,1427,783,1373]" captionTargetId="figure@10.[151,1436,772,1406]" captionTargetPageId="10" captionText="FIGURE 8. Testudinella clypeata. A – C. Loricae, ventral view. D – F. Loricae, cross-sectional view. G. Foot pseudosegments. Scale bars: A – F: 50 µm, G: 25 µm." httpUri="https://zenodo.org/record/187524/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
D–F) the dorsal margin appears convex, the lateral edges are broadly rounded and separated by a shallow sulcus from a convex median part ventrally. The characteristic foot-opening is a wide inverted U-shaped slit, situated ventral and sub-terminal. The foot (
<figureCitation id="85AF2A46FFC0FF96FD03EF97AD93F946" box="[687,758,1749,1775]" captionStart="FIGURE 8" captionStartId="10.[151,262,1413,1437]" captionTargetBox="[158,1427,783,1373]" captionTargetId="figure@10.[151,1436,772,1406]" captionTargetPageId="10" captionText="FIGURE 8. Testudinella clypeata. A – C. Loricae, ventral view. D – F. Loricae, cross-sectional view. G. Foot pseudosegments. Scale bars: A – F: 50 µm, G: 25 µm." httpUri="https://zenodo.org/record/187524/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
G) is composed of a long and wrinkled proximal part, a clearly offset slender distal pseudosegment and a less defined short penultimate pseudosegment. The dorsal and lateral antennae are situated somewhat posterior to the anterior third of the lorica: ratio lorica length/ position of antenna(e) relative to antero-dorsal margin varying from 2.70–2.85 (average 2.75) and 2.35–2.75 (average 2.58) respectively. Two red eyespots.
</paragraph>
</subSubSection>
<subSubSection id="558E6548FFC0FF90FF6AEED4ACB6FDF2" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="1D2B36C3FFC0FF97FF6AEED4AD15FF56" blockId="10.[151,1437,1517,2007]" lastBlockId="11.[151,1436,152,255]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
The trophi (
<figureCitation id="85AF2A46FFC0FF96FEFBEED4AECBF819" box="[343,430,1942,1968]" captionStart="FIGURE 10" captionStartId="11.[151,257,1970,1994]" captionTargetBox="[476,1110,1135,1969]" captionTargetId="figure@11.[476,1111,1135,2042]" captionTargetPageId="11" captionText="FIGURE 10. Testudinella clypeata, scanning electron microscope photographs of complete set of trophi. A. Frontal view. B. Caudal view. Scale bar: 10 µm. fo: fulcrum opening, sc: sub-ventral chamber." httpUri="https://zenodo.org/record/187526/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
and
<bibRefCitation id="79054B32FFC0FF96FE41EED4ADC0F819" author="De" box="[493,677,1942,1968]" pageId="10" pageNumber="19" refString="De Smet, W. H. (2005) Study of the trophi of Testudinella Bory de St. Vincent and Pompholyx Gosse (Rotifera: Testudinellidae) by scanning electron microscopy. Hydrobiologia, 546, 203 - 211." type="journal article" year="2005">
De
<bibRefCitation id="79054B32FFC0FF96FDB5EED4ADC0F819" author="Smet" box="[537,677,1942,1968]" pageId="10" pageNumber="19" refString="De Smet, W. H. (2005) Study of the trophi of Testudinella Bory de St. Vincent and Pompholyx Gosse (Rotifera: Testudinellidae) by scanning electron microscopy. Hydrobiologia, 546, 203 - 211." type="journal article" year="2005">Smet (2005</bibRefCitation>
</bibRefCitation>
)) are characterized by 14–16/14–16 (left/right) unci teeth. The three major teeth show a slender and distinctly offset head. The manubria are composed of three large superimposed chambers, the dorsal, median and ventral, and a strongly reduced sub-ventral chamber (
<figureCitation id="85AF2A46FFC1FF97FAC7E9DAAFD2FF71" captionStart="FIGURE 10" captionStartId="11.[151,257,1970,1994]" captionTargetBox="[476,1110,1135,1969]" captionTargetId="figure@11.[476,1111,1135,2042]" captionTargetPageId="11" captionText="FIGURE 10. Testudinella clypeata, scanning electron microscope photographs of complete set of trophi. A. Frontal view. B. Caudal view. Scale bar: 10 µm. fo: fulcrum opening, sc: sub-ventral chamber." httpUri="https://zenodo.org/record/187526/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
B: sc). About 28–30/25–26 arched rami scleropili are apparent in caudal view. The fulcrum shows a distinct proximal opening frontally (
<figureCitation id="85AF2A46FFC1FF97FE4EE9A7AD5FFF56" box="[482,570,229,255]" captionStart="FIGURE 10" captionStartId="11.[151,257,1970,1994]" captionTargetBox="[476,1110,1135,1969]" captionTargetId="figure@11.[476,1111,1135,2042]" captionTargetPageId="11" captionText="FIGURE 10. Testudinella clypeata, scanning electron microscope photographs of complete set of trophi. A. Frontal view. B. Caudal view. Scale bar: 10 µm. fo: fulcrum opening, sc: sub-ventral chamber." httpUri="https://zenodo.org/record/187526/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
: fo).
</paragraph>
<caption id="49EB664BFFC1FF97FF3BED63AAE1FB90" box="[151,1412,1057,1081]" httpUri="https://zenodo.org/record/187525/files/figure.png" pageId="11" pageNumber="12" targetBox="[249,1337,310,1035]" targetPageId="11">
<paragraph id="1D2B36C3FFC1FF97FF3BED63AAE1FB90" blockId="11.[151,1412,1057,1081]" box="[151,1412,1057,1081]" pageId="11" pageNumber="12">
<emphasis id="2FE0EAD1FFC1FF97FF3BED63AE7FFB90" bold="true" box="[151,282,1057,1081]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
<taxonomicName id="DA944D40FFC1FF97FE8CED60AD67FB90" box="[288,514,1058,1081]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="11" pageNumber="12" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC1FF97FE8CED60AD67FB90" box="[288,514,1058,1081]" italics="true" pageId="11" pageNumber="12">Testudinella clypeata</emphasis>
</taxonomicName>
, variation in shape of specimen.
<emphasis id="2FE0EAD1FFC1FF97FCCEED63ACFEFB90" bold="true" box="[866,923,1057,1081]" pageId="11" pageNumber="12">A, B.</emphasis>
Contracted.
<emphasis id="2FE0EAD1FFC1FF97FB88ED63AB5BFB90" bold="true" box="[1060,1086,1057,1081]" pageId="11" pageNumber="12">C.</emphasis>
Swimming. Scale bar: 50 µm.
</paragraph>
</caption>
<caption id="49EB664BFFC1FF97FF3BEEF0AB54F842" httpUri="https://zenodo.org/record/187526/files/figure.png" pageId="11" pageNumber="12" targetBox="[476,1110,1135,1969]" targetPageId="11">
<paragraph id="1D2B36C3FFC1FF97FF3BEEF0AB54F842" blockId="11.[151,1436,1970,2028]" pageId="11" pageNumber="12">
<emphasis id="2FE0EAD1FFC1FF97FF3BEEF0AE4FF863" bold="true" box="[151,298,1970,1994]" pageId="11" pageNumber="12">FIGURE 10.</emphasis>
<taxonomicName id="DA944D40FFC1FF97FE9EEEF0AD78F860" box="[306,541,1970,1993]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="11" pageNumber="12" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC1FF97FE9EEEF0AD78F860" box="[306,541,1970,1993]" italics="true" pageId="11" pageNumber="12">Testudinella clypeata</emphasis>
</taxonomicName>
, scanning electron microscope photographs of complete set of trophi.
<emphasis id="2FE0EAD1FFC1FF97FA87EEF0AA20F863" bold="true" box="[1323,1349,1970,1994]" pageId="11" pageNumber="12">A.</emphasis>
Frontal view.
<emphasis id="2FE0EAD1FFC1FF97FF7AEE96AF8BF845" bold="true" box="[214,238,2004,2028]" pageId="11" pageNumber="12">B.</emphasis>
Caudal view. Scale bar: 10 µm. fo: fulcrum opening, sc: sub-ventral chamber.
</paragraph>
</caption>
<paragraph id="1D2B36C3FFC6FF90FF6AE9DAAFA0FF71" blockId="12.[151,1437,152,603]" pageId="12" pageNumber="13">Measurements. Lorica length 150–170 µm, lorica width 100–130 µm, distal foot pseudosegment 18–22 µm.</paragraph>
<paragraph id="1D2B36C3FFC6FF90FF6AE9A7ACB6FDF2" blockId="12.[151,1437,152,603]" pageId="12" pageNumber="13">
According to
<bibRefCitation id="79054B32FFC6FF90FEC2E9A7ADDAFF56" box="[366,703,229,255]" pageId="12" pageNumber="19" refString="De Ridder, M. &amp; Segers, H. (1997) Monogonont Rotifera recorded in the world literature (except Africa) from 1960 to 1992. Studiedocumenten van het K. B. I. N., 88, 481 pp." type="book">De Ridder &amp; Segers (1997)</bibRefCitation>
,
<taxonomicName id="DA944D40FFC6FF90FD62E9A7AC2AFF57" box="[718,847,229,254]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="12" pageNumber="13" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC6FF90FD62E9A7AC2AFF57" box="[718,847,229,254]" italics="true" pageId="12" pageNumber="13">T. clypeata</emphasis>
</taxonomicName>
is a widespread species occurring in the marine littoral and brackish waters all over the Palaearctic region. The species has apparently often been confused with
<taxonomicName id="DA944D40FFC6FF90FF78E870AE37FEE2" box="[212,338,306,331]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="12" pageNumber="13" phylum="Rotifera" rank="species" species="elliptica">
<emphasis id="2FE0EAD1FFC6FF90FF78E870AE37FEE2" box="[212,338,306,331]" italics="true" pageId="12" pageNumber="13">T. elliptica</emphasis>
</taxonomicName>
, however, the reliable illustrated records by
<bibRefCitation id="79054B32FFC6FF90FCCAE870ABE3FEE5" box="[870,1158,306,332]" pageId="12" pageNumber="19" refString="De Manuel, J., Pretus, J. LL. &amp; Jaume, D. (1992) Rotifers from the Balearic archipelago. Hydrobiologia, 239, 33 - 41." type="journal article">De Manuel et al. (1992)</bibRefCitation>
, De Maeseneer (1980),
<bibRefCitation id="79054B32FFC6FF90FF3BE81BAE16FEDA" author="De" box="[151,371,345,371]" pageId="12" pageNumber="19" refString="De Ridder, M. (1957 a) Enkele raderdiertjes van het Zwin. Mededelingen Belgische Nationale Vereniging der Leraren in de Biologie, 3, 6 pp." type="book" year="1957" yearSuffix="a">De Ridder (1957a</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FE2DE81BAEF6FEDA" author="De" box="[385,403,345,371]" pageId="12" pageNumber="19" refString="De Ridder, M. (1957 b) Onderzoekingen over brakwaterrotatorien. II. Het Zwin te Knokke. Natuurwetenschappelijk Tijdschrift, 39, 109 - 126, pl. 1." type="journal article" year="1957" yearSuffix="b">b</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FE0CE81BAEBAFEDA" author="Ridder" box="[416,479,345,371]" pageId="12" pageNumber="19" refString="De Ridder, M. (1967) Quelques rotiferes de Majorque. Les Naturalistes Belges, 48, 409 - 418." type="journal article" year="1967">1967</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FE41E81BAD4BFEDA" author="Ridder" box="[493,558,345,371]" pageId="12" pageNumber="19" refString="De Ridder, M. (1972) Rotatoria. In: Tuxen, S. L. (Ed.), The Zoology of Iceland, Vol. II, P. 13, Ejnar Munksgaard: Copenhagen and Reykjavik, 106 pp, 11 pls." type="book" year="1972">1972</bibRefCitation>
),
<bibRefCitation id="79054B32FFC6FF90FDEFE81BAD90FEDA" author="Gillard" box="[579,757,345,371]" lastPageId="12" lastPageNumber="20" pageId="12" pageNumber="19" refString="Gillard, A. (1947) Het geslacht Testudinella Bory de St Vincent (Rotat.) in Belgie. Natuurwetenschappelijk Tijdschrift, 29, 153 - 158, pl. 2." type="journal article" year="1947">Gillard (1947)</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FCAEE81BAB4BFEDA" author="Hudson" box="[770,1070,345,371]" pageId="12" pageNumber="20" refString="Hudson, C. T. &amp; Gosse, P. H. (1886) The Rotifera; or wheel-animalcules, both British and foreign. Longmans, Green, and Co., London, Vol. I, VI + 128 pp., 15 pls, Vol. II, 144 pp., pls 16 - 30." type="book chapter" year="1886">Hudson &amp; Gosse (1886)</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FB97E81BAB9CFEDA" author="Seehaus" box="[1083,1273,345,371]" pageId="12" pageNumber="20" refString="Seehaus, W. (1930) Zur Morphologie der Radertiergattung Testudinella Bory de St. Vincent (= Pterodina Ehrenberg). Zeitschrift fur wissenschaftliche Zoologie, 137, 175 - 273." type="journal article" year="1930">Seehaus (1930)</bibRefCitation>
, von
<bibRefCitation id="79054B32FFC6FF90FA91E81BAF88FE33" author="Hofsten" pageId="12" pageNumber="20" refString="von Hofsten, N. (1911 - 1912) Marine, litorale Rotatorien der skandinavischen Westkuste. Zoologiska Bidrag, Uppsala, 1, 163 - 228." type="journal article" year="1911">Hofsten (1911)</bibRefCitation>
,
<bibRefCitation id="79054B32FFC6FF90FF57E8C2AEDCFE33" author="Wulfert" box="[251,441,384,410]" pageId="12" pageNumber="20" refString="Wulfert, K. (1942) Uber die Meeres- und Brackwasser-Rotatorien in der Umgebung von Rovigno d'Istria. Thalassia - Istituto Italo-Germanico di Biologia Marina di Rovigno d'Istria - Venezia, 4, 3 - 26, pl. 1 - 6." type="journal article" year="1942">Wulfert (1942)</bibRefCitation>
and personal observations confirm this general distribution. The reports from freshwater habitats are probably misidentifications concerning
<taxonomicName id="DA944D40FFC6FF90FC26E8E5AB6FFE69" box="[906,1034,423,448]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="12" pageNumber="13" phylum="Rotifera" rank="species" species="elliptica">
<emphasis id="2FE0EAD1FFC6FF90FC26E8E5AB6FFE69" box="[906,1034,423,448]" italics="true" pageId="12" pageNumber="13">T. elliptica</emphasis>
</taxonomicName>
.
<bibRefCitation id="79054B32FFC6FF90FBB6E8E4AA01FE69" box="[1050,1380,422,448]" pageId="12" pageNumber="19" refString="De Ridder, M. &amp; Segers, H. (1997) Monogonont Rotifera recorded in the world literature (except Africa) from 1960 to 1992. Studiedocumenten van het K. B. I. N., 88, 481 pp." type="book">De Ridder &amp; Segers (1997)</bibRefCitation>
also report the species from the Nearctic region (Cap Breton Island, Nova
<collectingCountry id="65837653FFC6FF90FC65E88FAB70FE4E" box="[969,1045,461,487]" name="United Kingdom" pageId="12" pageNumber="13">Scotia</collectingCountry>
,
<collectingCountry id="65837653FFC6FF90FB8DE88FAB18FE4E" box="[1057,1149,461,487]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
), incorrectly referring to
<bibRefCitation id="79054B32FFC6FF90FF3BE8B6AE89FDA7" author="Chengalath" box="[151,492,500,526]" pageId="12" pageNumber="19" refString="Chengalath, R. &amp; Koste, W. (1988) Composition of littoral rotifer communities of Cape Breton Island, Nova Scotia, Canada. Verhandlungen der Internationalen Vereinigung fur Theoretische und Angewandte Limnologie, 23, 2019 - 2027." type="journal article" year="1988">Chengalath &amp; Koste (1988)</bibRefCitation>
. However, the specimens mentioned sub
<taxonomicName id="DA944D40FFC6FF90FC49E8B6AB0DFDA4" box="[997,1128,500,525]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="12" pageNumber="13" phylum="Rotifera" rank="species" species="elliptica">
<emphasis id="2FE0EAD1FFC6FF90FC49E8B6AB0DFDA4" box="[997,1128,500,525]" italics="true" pageId="12" pageNumber="13">T. elliptica</emphasis>
</taxonomicName>
by
<bibRefCitation id="79054B32FFC6FF90FB3BE8B6AAF2FDA7" author="Jersabek" box="[1175,1431,500,526]" pageId="12" pageNumber="20" refString="Jersabek, C. D., Segers, H. &amp; Morris, P. J. (2003) An illustrated online catalog of the Rotifera in the Academy of Natural Sciences of Philadelphia (version 1.0: 2003 - April- 8). [WWW database] URL, http: // rotifer. acnatsci. org / rotifer. php" type="book chapter" year="2003">Jersabek et al. (2003</bibRefCitation>
; ANSP Catalog Number 1828) and originating from a brackish pool near Atlantic City, New
<collectingCountry id="65837653FFC6FF90FB72EB58AA4CFD9D" box="[1246,1321,538,564]" name="Jersey" pageId="12" pageNumber="13">Jersey</collectingCountry>
,
<collectingCountry id="65837653FFC6FF90FA99EB58AA15FD9D" box="[1333,1392,538,564]" name="United States of America" pageId="12" pageNumber="13">USA</collectingCountry>
are distinctly
<taxonomicName id="DA944D40FFC6FF90FEA7EB03AEEEFDF3" box="[267,395,577,602]" class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" order="Flosculariaceae" pageId="12" pageNumber="13" phylum="Rotifera" rank="species" species="clypeata">
<emphasis id="2FE0EAD1FFC6FF90FEA7EB03AEEEFDF3" box="[267,395,577,602]" italics="true" pageId="12" pageNumber="13">T. clypeata</emphasis>
</taxonomicName>
, illustrating the species’ presence in the Nearctic.
</paragraph>
</subSubSection>
</treatment>
</document>
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