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<document id="1ED35C1C9DB18A5AF8323C869A72504D" ID-DOI="10.1111/j.1096-3642.2012.00826.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5408089" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630348471941" checkinUser="marcus" docAuthor="Han, Hongxiang, Galsworthy, Anthony C. &amp; Xue, Dayong" docDate="2012" docId="0389DF41FF81A91BFEB2A909FDB6FB93" docLanguage="en" docName="j.1096-3642.2012.00826.x.pdf" docOrigin="Zoological Journal of the Linnean Society 165 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00826.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Comibaena bellula Han &amp; Galsworthy &amp; Xue 2012, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="747" masterDocId="FFB0A739FF96A903FF93AC20FFF5FFC1" masterDocTitle="The Comibaenini of China (Geometridae: Geometrinae), with a review of the tribe" masterLastPageNumber="772" masterPageNumber="723" pageNumber="746" updateTime="1699240510822" updateUser="plazi">
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<mods:title id="1261D06D8DEA76A92108CE430C075308">The Comibaenini of China (Geometridae: Geometrinae), with a review of the tribe</mods:title>
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<mods:namePart id="9A95744335B60612638A2C52DABE519D">Han, Hongxiang</mods:namePart>
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<mods:namePart id="43A21ED9782A6C223B814CB4779A23D8">Galsworthy, Anthony C.</mods:namePart>
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<mods:namePart id="14BEB7B30B54EF2058374464A895C976">Xue, Dayong</mods:namePart>
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<mods:title id="5B4665CA8A2609249DD72F37F626330F">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="3494032F2BECFD0F8331E02267A9BB67">2012</mods:date>
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<mods:number id="8F43A6EA41C48005900422DBBBB4AEA1">2012-07-25</mods:number>
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<mods:number id="D958F0EF7E4288A78BAAB0EE6766F725">165</mods:number>
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<mods:identifier id="65078421A26A7387F7CE062A2878CB43" type="DOI">10.1111/j.1096-3642.2012.00826.x</mods:identifier>
<mods:identifier id="27129323E5A5DE85EA022983BFC3654E" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="897C628D8411F0CC49CE486A38AA4809" type="Zenodo-Dep">5408089</mods:identifier>
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<treatment id="0389DF41FF81A91BFEB2A909FDB6FB93" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93" httpUri="http://treatment.plazi.org/id/0389DF41FF81A91BFEB2A909FDB6FB93" lastPageId="24" lastPageNumber="747" pageId="23" pageNumber="746">
<subSubSection id="C33A3DDCFF81A914FEB2A909FD7BFAFF" box="[289,654,1321,1342]" pageId="23" pageNumber="746" type="nomenclature">
<paragraph id="8B9F6E57FF81A914FEB2A909FD7BFAFF" blockId="23.[289,654,1321,1375]" box="[289,654,1321,1342]" pageId="23" pageNumber="746">
<heading id="D0D7D93BFF81A914FEB2A909FD7BFAFF" box="[289,654,1321,1342]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="746" reason="4">
<taxonomicName id="4C2015D4FF81A914FEB2A909FDEBFAFC" authority="Han &amp; Galsworthy &amp; Xue, 2012" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[289,542,1321,1342]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="bellula" status="sp. nov.">COMIBAENA BELLULA</taxonomicName>
<emphasis id="B954B245FF81A914FDBAA90CFD7BFAFF" bold="true" box="[553,654,1321,1342]" pageId="23" pageNumber="746">
<taxonomicNameLabel id="A2670F3EFF81A914FDBAA90CFD7BFAFF" box="[553,654,1321,1342]" pageId="23" pageNumber="746" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C33A3DDCFF81A91BFEC2A969FDB6FB93" lastPageId="24" lastPageNumber="747" pageId="23" pageNumber="746" type="description">
<paragraph id="8B9F6E57FF81A914FEC2A969FDABFA9E" blockId="23.[289,654,1321,1375]" box="[337,606,1353,1375]" pageId="23" pageNumber="746">
(
<figureCitation id="131B72D2FF81A914FECAA969FE46FA9F" box="[345,435,1353,1374]" captionStart="Figures 1–26" captionStartId="6.[144,234,1656,1675]" captionTargetBox="[148,1414,202,1609]" captionTargetId="figure-0@6.[144,1425,195,1626]" captionTargetPageId="6" captionText="Figures 1–26. Comibaenini moths. 1, Comibaena quadrinotata; 2, Comibaena fuscidorsata; 3, Comibaena cassidara; 4, Comibaena attenuata; 5, Comibaena biplaga; 6, Comibaena apicipicta; 7, Comibaena striataria; 8, Comibaena delineata; 9, Comibaena dubernardi; 10, Comibaena ornataria; 11–15, Comibaena pictipennis; 11, from Darjeeling; 12, syntype of Comibaena pictipennis superornataria; 13, from Tengchong, Yunnan; 14, from Baoshan, Yunnan; 15, from Nanchuan, Sichuan; 16, Comibaena parornataria sp. nov., holotype; 17, Comibaena bellula sp. nov., holotype; 18, Comibaena decora sp. nov., holotype; 19, Comibaena auromaculata sp. nov., holotype; 20, Comibaena tibetensis sp. nov., holotype; 21, Comibaena tancrei; 22, Comibaena sheni sp. nov., holotype; 23, Comibaena argentataria; 24, Comibaena signifera subargentaria; 25, Comibaena takasago; 26, Comibaena flavicans. Scale bar = 13 mm." figureDoi="http://doi.org/10.5281/zenodo.5408091" httpUri="https://zenodo.org/record/5408091/files/figure.png" pageId="23" pageNumber="746">FIGS 17</figureCitation>
,
<figureCitation id="131B72D2FF81A914FE50A969FE14FA9F" box="[451,481,1353,1374]" captionStart="Figures 67–78" captionStartId="9.[164,254,1685,1704]" captionTargetBox="[165,1445,201,1646]" captionTargetId="figure-0@9.[164,1445,193,1655]" captionTargetPageId="9" captionText="Figures 67–78. Male genitalia. 67–69, Comibaena pictipennis. 67, from Dahaoping, Tengchong, Yunnan; 68, from Nyingchi, Tibet; 69, from Qingchengshan, Sichuan; 70, Comibaena parornataria; 71, Comibaena bellula; 72, Comibaena decora; 73, Comibaena auromaculata; 74, Comibaena tibetensis; 75, Comibaena tancrei; 76, Comibaena sheni; 77, Comibaena argentataria; 78, Comibaena signifera subargentaria. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5408098" httpUri="https://zenodo.org/record/5408098/files/figure.png" pageId="23" pageNumber="746">71</figureCitation>
,
<figureCitation id="131B72D2FF81A914FE63A969FDE9FA9F" box="[496,540,1353,1374]" captionStart="Figures 108–131" captionStartId="13.[164,254,1693,1712]" captionTargetBox="[188,1424,196,1663]" captionTargetId="figure-0@13.[177,1432,193,1663]" captionTargetPageId="13" captionText="Figures 108–131. Aedeagus. 108, Comibaena quadrinotata; 109, Comibaena fuscidorsata; 110, Comibaena cassidara; 111, Comibaena attenuata; 112, Comibaena biplaga; 113, Comibaena apicipicta; 114, Comibaena striataria; 115, Comibaena delineata; 116, Comibaena dubernardi; 117, Comibaena ornataria; 118–119, Comibaena pictipennis; 118, from Tengchong, Yunnan; 119, from Nyingchi; 120, Comibaena parornataria; 121, Comibaena bellula; 122, Comibaena decora; 123, Comibaena auromaculata; 124, Comibaena tibetensis; 125, Comibaena tancrei; 126, Comibaena sheni; 127, Comibaena argentataria; 128, Comibaena signifera subargentaria; 129, Comibaena takasago; 130, Comibaena flavicans; 131, Comibaena nigromacularia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5408106" httpUri="https://zenodo.org/record/5408106/files/figure.png" pageId="23" pageNumber="746">121</figureCitation>
,
<figureCitation id="131B72D2FF81A914FDB8A969FDA3FA9E" box="[555,598,1353,1375]" captionStart="Figures 176–187" captionStartId="16.[144,234,1712,1731]" captionTargetBox="[144,1415,211,1673]" captionTargetId="figure-0@16.[144,1425,193,1682]" captionTargetPageId="16" captionText="Figures 176–187. Female genitalia. 176, Comibaena quadrinotata; 177, Comibaena cassidara; 178, Comibaena attenuata; 179, Comibaena biplaga; 180, Comibaena striataria; 181, Comibaena delineata; 182, Comibaena dubernardi; 183, Comibaena ornataria; 184, Comibaena pictipennis; 185, Comibaena parornataria; 186, Comibaena bellula; 187, Comibaena tancrei. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5408114" httpUri="https://zenodo.org/record/5408114/files/figure.png" pageId="23" pageNumber="746">186</figureCitation>
)
</paragraph>
<paragraph id="8B9F6E57FF81A914FF37A950FEAFF935" blockId="23.[164,779,1391,1903]" pageId="23" pageNumber="746">
<emphasis id="B954B245FF81A914FF37A950FED9FA44" box="[164,300,1392,1413]" italics="true" pageId="23" pageNumber="746">Description:</emphasis>
Antenna in male bipectinate, outer rami longer than inner rami; about terminal one-quarter simple in male; simple filiform in female. Frons and labial palpus black with a few white scales. Vertex black. Tegula pale yellowish-green. Dorsal side of thorax green. A white patch present between metathorax and the first abdominal segment. Abdomen with anterior three segments green, with white dorsal line and white intersegments, other segments possibly whitish, but perhaps discoloured in specimens examined. Hind tibia in male dilated, with hairpencil, terminal extension longer than half length of the first tarsus.
</paragraph>
<paragraph id="8B9F6E57FF81A914FF2FAADDFB0CFCBC" blockId="23.[164,779,1391,1903]" lastBlockId="23.[826,1441,198,1413]" pageId="23" pageNumber="746">
Forewing length:
<specimenCount id="9D26A5DEFF81A914FE0AAB20FE46F8D3" box="[409,435,1792,1810]" pageId="23" pageNumber="746" type="male">♂</specimenCount>
<specimenCount id="9D26A5DEFF81A914FE27AB20FE32F8D3" box="[436,455,1792,1810]" pageId="23" pageNumber="746" type="female">♀</specimenCount>
14.5–
<quantity id="4CD8C3B2FF81A914FD84AADDFD92F8D3" box="[535,615,1789,1811]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="746" unit="mm" value="16.0">16 mm</quantity>
. Wings clear green, with diffuse, white, wavy transverse stripes in all but marginal areas. Forewing costa with a wide grass-green band, slightly tapering towards apex; anteromedial line white, slender, forming a tooth on lower margin of cell; postmedial line close to outer margin, white, slightly curved outwards, folded inwards on CuA
<subScript id="17A46C12FF81A914FBD7AD0CFBB9FEF9" attach="left" box="[1092,1100,300,312]" fontSize="5" pageId="23" pageNumber="746">2</subScript>
, then merging with the dark reddish-brown subtornal patch, becoming red or reddish-brown inside the patch; area outside postmedial line from costa to M
<subScript id="17A46C12FF81A914FBCFADA7FB91FE52" attach="left" box="[1116,1124,391,403]" fontSize="5" pageId="23" pageNumber="746">3</subScript>
paler and appearing as an indistinct white patch, white submarginal line visible; terminal line a series of black dots between veins; fringes pale green, terminal half much paler. Hindwing inner margin elongate; apex with a green patch, bordered by a dark brown to black submarginal line, Rs brown inside the patch; submarginal line very close to outer margin, forming a tooth on the end of vein M
<subScript id="17A46C12FF81A914FC02AE5CFC6CFD49" attach="left" box="[913,921,636,648]" fontSize="5" pageId="23" pageNumber="746">3</subScript>
, another bigger tooth on anal fold, background colour visible inside; a pink patch present on tornal angle below the larger tooth; black terminal line linear above M
<subScript id="17A46C12FF81A914FB85AEF8FBEBFD25" attach="left" box="[1046,1054,728,740]" fontSize="5" pageId="23" pageNumber="746">2</subScript>
, and appearing as a series of dots below it; fringes pinkish, basal part darker, middle part much paler, appearing as two rings, much darker on vein ends of Rs and M
<subScript id="17A46C12FF81A914FBC8AF14FB96FC81" attach="left" box="[1115,1123,820,832]" fontSize="5" pageId="23" pageNumber="746">1</subScript>
. Discal spots on both wings small black dots. Underside much paler than upperside, streaks on upperside discernible.
</paragraph>
<paragraph id="8B9F6E57FF81A914FCC0AFA6FB28FC18" blockId="23.[826,1441,198,1413]" pageId="23" pageNumber="746">Third sternite in the male with a pair of setal patches; eighth sternite with posterior margin sclerotized, slightly protruding at middle.</paragraph>
<paragraph id="8B9F6E57FF81A914FCC0AFC1FC61FB0F" blockId="23.[826,1441,198,1413]" pageId="23" pageNumber="746">Male genitalia. Uncus well divided. Socii broad basally, tapering and hooked apically, with a small blunt process at middle. Costal lobe with terminal part strongly sclerotized, spinose, and pointed, bearing a pointed tooth at middle of the ventral margin. Vinculum widely bifurcate, with the middle part flat, lateral branches tiny and blunt. Aedeagus slender.</paragraph>
<paragraph id="8B9F6E57FF81A914FCC0A8F6FB87FA44" blockId="23.[826,1441,198,1413]" pageId="23" pageNumber="746">
Female genitalia. Sterigma strongly sclerotized, posterior margin of lamella postvaginalis with a small rounded concavity; lamella antevaginalis wrinkled, posterior margin sinuous, anterior margin with two tiny protrusions extending dorsally at middle (dark and indistinct in
<figureCitation id="131B72D2FF81A914FB95A94FFB90FA44" box="[1030,1125,1391,1413]" captionStart="Figures 176–187" captionStartId="16.[144,234,1712,1731]" captionTargetBox="[144,1415,211,1673]" captionTargetId="figure-0@16.[144,1425,193,1682]" captionTargetPageId="16" captionText="Figures 176–187. Female genitalia. 176, Comibaena quadrinotata; 177, Comibaena cassidara; 178, Comibaena attenuata; 179, Comibaena biplaga; 180, Comibaena striataria; 181, Comibaena delineata; 182, Comibaena dubernardi; 183, Comibaena ornataria; 184, Comibaena pictipennis; 185, Comibaena parornataria; 186, Comibaena bellula; 187, Comibaena tancrei. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5408114" httpUri="https://zenodo.org/record/5408114/files/figure.png" pageId="23" pageNumber="746">Fig. 186</figureCitation>
).
</paragraph>
<paragraph id="8B9F6E57FF81A91BFCA9A9ECFEF6FEF6" blockId="23.[826,1441,1483,1903]" lastBlockId="24.[144,759,198,801]" lastPageId="24" lastPageNumber="747" pageId="23" pageNumber="746">
<emphasis id="B954B245FF81A914FCA9A9ECFB56FA21" box="[826,1187,1483,1505]" italics="true" pageId="23" pageNumber="746">
Diagnosis:
<taxonomicName id="4C2015D4FF81A914FC50A9EBFB56FA21" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[963,1187,1483,1504]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="bellula">Comibaena bellula</taxonomicName>
</emphasis>
is quite similar to
<taxonomicName id="4C2015D4FF81A914FCA9A9CAFBC7FA3E" authorityName=": Prout" authorityYear="1912" box="[826,1074,1514,1535]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="ornataria">
<emphasis id="B954B245FF81A914FCA9A9CAFBC7FA3E" box="[826,1074,1514,1535]" italics="true" pageId="23" pageNumber="746">Comibaena ornataria</emphasis>
</taxonomicName>
in wing pattern, but the submarginal line of the hindwing forms a tooth at the middle, which is absent in
<taxonomicName id="4C2015D4FF81A914FBE1AA07FA93F9FD" authorityName=": Prout" authorityYear="1912" box="[1138,1382,1575,1596]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="ornataria">
<emphasis id="B954B245FF81A914FBE1AA07FA93F9FD" box="[1138,1382,1575,1596]" italics="true" pageId="23" pageNumber="746">Comibaena ornataria</emphasis>
</taxonomicName>
, and the tornal patch is larger.
<taxonomicName id="4C2015D4FF81A914FBE7AA65FAB9F99B" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[1140,1356,1605,1626]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B954B245FF81A914FBE7AA65FAB9F99B" box="[1140,1356,1605,1626]" italics="true" pageId="23" pageNumber="746">Comibaena bellula</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C2015D4FF81A914FC27AA44FB4FF9BB" authorityName=": Prout" authorityYear="1933" box="[948,1210,1636,1658]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B954B245FF81A914FC27AA44FB4FF9BB" box="[948,1210,1636,1658]" italics="true" pageId="23" pageNumber="746">Comibaena pictipennis</emphasis>
</taxonomicName>
, but the apical and tornal patches, and the middle tooth on the hindwing are much smaller.
<taxonomicName id="4C2015D4FF81A914FBA4AA81FAEFF977" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[1079,1306,1697,1718]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B954B245FF81A914FBA4AA81FAEFF977" box="[1079,1306,1697,1718]" italics="true" pageId="23" pageNumber="746">Comibaena bellula</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C2015D4FF81A914FCA9AAE0FB95F914" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[826,1120,1728,1749]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="parornataria">
<emphasis id="B954B245FF81A914FCA9AAE0FB95F914" box="[826,1120,1728,1749]" italics="true" pageId="23" pageNumber="746">Comibaena parornataria</emphasis>
</taxonomicName>
in the similar hindwing apical patch, but differs in the tornal patch and terminal line on the forewing, and the middle tooth and tornal patch on the hindwing. In the male genitalia,
<taxonomicName id="4C2015D4FF81A914FCA9AB1AFBE5F88E" authorityName="Han &amp; Galsworthy &amp; Xue" authorityYear="2012" box="[826,1040,1850,1871]" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B954B245FF81A914FCA9AB1AFBE5F88E" box="[826,1040,1850,1871]" italics="true" pageId="23" pageNumber="746">Comibaena bellula</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C2015D4FF81A914FB4EAB1AFC87F8AF" authorityName=": Prout" authorityYear="1912" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="746" phylum="Arthropoda" rank="species" species="ornataria">
<emphasis id="B954B245FF81A914FB4EAB1AFC87F8AF" italics="true" pageId="23" pageNumber="746">Comibaena ornataria</emphasis>
</taxonomicName>
in having the middle part of the vinculum shallowly concave and only very tiny lateral branches. In the female genitalia, the posterior margin of the sterigma is more deeply concave than in
<taxonomicName id="4C2015D4FF8EA91BFDE5AD23FF09FEF6" authorityName=": Prout" authorityYear="1912" class="Insecta" family="Geometridae" genus="Comibaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="747" phylum="Arthropoda" rank="species" species="ornataria">
<emphasis id="B954B245FF8EA91BFDE5AD23FF09FEF6" italics="true" pageId="24" pageNumber="747">Comibaena ornataria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F6E57FF8EA91BFF3BAD60FDC6FCE1" blockId="24.[144,759,198,801]" pageId="24" pageNumber="747">
<materialsCitation id="3B48640AFF8EA91BFF3BAD60FD06FEB4" collectingDate="2009-06-20" collectionCode="IZCAS" collectorName="Qi Feng" country="China" elevation="3260" location="Mountain Vivarium" municipality="Lijiang" pageId="24" pageNumber="747" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="549BD0F5FF8EA91BFF3BAD60FEE4FE97" box="[168,273,320,342]" pageId="24" pageNumber="747" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D26A5DEFF8EA91BFEB6AD63FECBFE94" box="[293,318,323,341]" pageId="24" pageNumber="747" type="male">♂</specimenCount>
,
<collectingCountry id="F3372EC7FF8EA91BFEC7AD60FE68FE97" box="[340,413,320,342]" name="China" pageId="24" pageNumber="747">China</collectingCountry>
:
<collectingRegion id="49E4A0B5FF8EA91BFE3CAD61FDFEFE97" box="[431,523,321,342]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BFBF42DFF8EA91BFD8CAD60FD86FE97" box="[543,627,320,342]" pageId="24" pageNumber="747">Lijiang</collectingMunicipality>
,
<location id="8EFF388CFF8EA91BFD14AD60FF0BFEB4" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFD14AD60FF0BFEB4" country="China" municipality="Lijiang" name="Mountain Vivarium" pageId="24" pageNumber="747" stateProvince="Yunnan">Mountain Vivarium</location>
,
<quantity id="4CD8C3B2FF8EA91BFE98AD7FFE94FEB5" box="[267,353,351,373]" metricMagnitude="3" metricUnit="m" metricValue="3.26" pageId="24" pageNumber="747" unit="m" value="3260.0">
<elevation id="000D8964FF8EA91BFE98AD7FFE94FEB5" box="[267,353,351,373]" metricMagnitude="3" metricUnit="m" metricValue="3.26" pageId="24" pageNumber="747" unit="m" value="3260.0">3260 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFEFCAD7FFE13FEB4" box="[367,486,351,373]" pageId="24" pageNumber="747" value="2009-06-20">
<collectingDate id="EFDAB17FFF8EA91BFEFCAD7FFE13FEB4" box="[367,486,351,373]" pageId="24" pageNumber="747" value="2009-06-20">20.vi.2009</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFDBFAD7FFD7CFEB5" box="[556,649,351,373]" pageId="24" pageNumber="747">Qi Feng</collectorName>
(
<collectionCode id="ED31F692FF8EA91BFD09AD7FFD1EFEB4" box="[666,747,351,373]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="24" pageNumber="747">IZCAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B48640AFF8EA91BFF03AD5EFDB6FE10" collectingDate="2009-06-20" collectionCode="IZCAS" collectorName="Qi Feng &amp; Han Hongxiang &amp; Xue Dayong &amp; Yang Chao" country="China" elevation="3260" location="Mountain Vivarium" municipality="Lijiang" pageId="24" pageNumber="747" specimenCount="8" specimenCount-male="8" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="549BD0F5FF8EA91BFF03AD5EFEF6FE52" box="[144,259,382,403]" pageId="24" pageNumber="747" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED31F692FF8EA91BFE84AD5EFE9DFE55" box="[279,360,382,404]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="24" pageNumber="747">IZCAS</collectionCode>
):
<collectingCountry id="F3372EC7FF8EA91BFEECAD5DFE3DFE52" box="[383,456,381,403]" name="China" pageId="24" pageNumber="747">China</collectingCountry>
:
<specimenCount id="9D26A5DEFF8EA91BFE4BAD5EFE0AFE53" box="[472,511,382,403]" pageId="24" pageNumber="747" type="male">8♂</specimenCount>
, same data as holotype, 15–18,
<date id="FF9E4897FF8EA91BFEBEADBCFE51FE73" box="[301,420,412,434]" pageId="24" pageNumber="747" value="2009-06-20">
<collectingDate id="EFDAB17FFF8EA91BFEBEADBCFE51FE73" box="[301,420,412,434]" pageId="24" pageNumber="747" value="2009-06-20">20.vi.2009</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFE62ADBCFDA0FE73" box="[497,597,412,434]" pageId="24" pageNumber="747">Qi Feng</collectorName>
,
<collectorName id="26D50B81FF8EA91BFDF5ADBDFF31FE10" pageId="24" pageNumber="747">Han Hongxiang</collectorName>
,
<collectorName id="26D50B81FF8EA91BFF49AD9BFE9AFE11" box="[218,367,443,464]" pageId="24" pageNumber="747">Xue Dayong</collectorName>
and
<collectorName id="26D50B81FF8EA91BFE2FAD9BFDB6FE10" box="[444,579,443,465]" pageId="24" pageNumber="747">Yang Chao</collectorName>
</materialsCitation>
;
<materialsCitation id="3B48640AFF8EA91BFDCAAD9BFE7AFDCC" collectingDate="2009-06-19" collectorName="Yang Chao &amp; Qi Feng &amp; Xue Dayong" country="China" elevation="3097" location="Wenhai" municipality="Lijiang" pageId="24" pageNumber="747" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFDCAAD9BFD75FE0E" box="[601,640,443,464]" pageId="24" pageNumber="747" type="male">3♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFD04AD9BFD06FE11" box="[663,755,443,464]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BFBF42DFF8EA91BFF03ADF9FF11FE2E" box="[144,228,473,495]" pageId="24" pageNumber="747">Lijiang</collectingMunicipality>
,
<location id="8EFF388CFF8EA91BFF63ADF9FEBCFE2E" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFF63ADF9FEBCFE2E" box="[240,329,473,495]" country="China" municipality="Lijiang" name="Wenhai" pageId="24" pageNumber="747" stateProvince="Yunnan">Wenhai</location>
,
<quantity id="4CD8C3B2FF8EA91BFEC4ADF9FE59FE2E" box="[343,428,473,495]" metricMagnitude="3" metricUnit="m" metricValue="3.097" pageId="24" pageNumber="747" unit="m" value="3097.0">
<elevation id="000D8964FF8EA91BFEC4ADF9FE59FE2E" box="[343,428,473,495]" metricMagnitude="3" metricUnit="m" metricValue="3.097" pageId="24" pageNumber="747" unit="m" value="3097.0">3097 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFE29ADF9FDC4FE2E" box="[442,561,473,495]" pageId="24" pageNumber="747" value="2009-06-19">
<collectingDate id="EFDAB17FFF8EA91BFE29ADF9FDC4FE2E" box="[442,561,473,495]" pageId="24" pageNumber="747" value="2009-06-19">19.vi.2009</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFDE0ADFAFD07FE2E" box="[627,754,473,495]" pageId="24" pageNumber="747">Yang Chao</collectorName>
,
<collectorName id="26D50B81FF8EA91BFF03ADD8FF04FDCC" box="[144,241,504,526]" pageId="24" pageNumber="747">Qi Feng</collectorName>
,
<collectorName id="26D50B81FF8EA91BFF6CADD8FE7AFDCC" box="[255,399,504,525]" pageId="24" pageNumber="747">Xue Dayong</collectorName>
</materialsCitation>
, IZCAS slide no. Geom-01059;
<materialsCitation id="3B48640AFF8EA91BFF03AE37FDC9FD8A" collectingDate="2009-06-23" collectorName="Yang Chao &amp; Han Hongxiang" country="China" elevation="3296" location="Ganheba" municipality="Lijiang" pageId="24" pageNumber="747" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFF03AE37FF43FDEA" box="[144,182,535,556]" pageId="24" pageNumber="747" type="male">3♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFF5AAE37FED0FDED" box="[201,293,535,556]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BFBF42DFF8EA91BFEA6AE36FE7CFDED" box="[309,393,534,556]" pageId="24" pageNumber="747">Lijiang</collectingMunicipality>
,
<location id="8EFF388CFF8EA91BFE09AE36FDF6FDED" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFE09AE36FDF6FDED" box="[410,515,534,556]" country="China" municipality="Lijiang" name="Ganheba" pageId="24" pageNumber="747" stateProvince="Yunnan">Ganheba</location>
,
<quantity id="4CD8C3B2FF8EA91BFD87AE36FD9CFDED" box="[532,617,534,556]" metricMagnitude="3" metricUnit="m" metricValue="3.296" pageId="24" pageNumber="747" unit="m" value="3296.0">
<elevation id="000D8964FF8EA91BFD87AE36FD9CFDED" box="[532,617,534,556]" metricMagnitude="3" metricUnit="m" metricValue="3.296" pageId="24" pageNumber="747" unit="m" value="3296.0">3296 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFDE8AE36FD07FDED" box="[635,754,534,556]" pageId="24" pageNumber="747" value="2009-06-23">
<collectingDate id="EFDAB17FFF8EA91BFDE8AE36FD07FDED" box="[635,754,534,556]" pageId="24" pageNumber="747" value="2009-06-23">23.vi.2009</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFF41AE15FEA9FD8A" box="[210,348,565,587]" pageId="24" pageNumber="747">Yang Chao</collectorName>
,
<collectorName id="26D50B81FF8EA91BFEE5AE15FDC9FD8A" box="[374,572,565,587]" pageId="24" pageNumber="747">Han Hongxiang</collectorName>
</materialsCitation>
;
<materialsCitation id="3B48640AFF8EA91BFDC6AE15FF38FD49" collectingDate="2009-06-21" collectorName="Yang Chao" country="China" elevation="2680" location="Yushuizhai" municipality="Lijiang" pageId="24" pageNumber="747" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFDC6AE15FD89FD8B" box="[597,636,565,586]" pageId="24" pageNumber="747" type="male">2♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFD04AE15FD06FD8B" box="[663,755,565,586]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BFBF42DFF8EA91BFF03AE74FF11FDAB" box="[144,228,596,618]" pageId="24" pageNumber="747">Lijiang</collectingMunicipality>
,
<location id="8EFF388CFF8EA91BFF65AE74FE8DFDAB" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFF65AE74FE8DFDAB" box="[246,376,596,618]" country="China" municipality="Lijiang" name="Yushuizhai" pageId="24" pageNumber="747" stateProvince="Yunnan">Yushuizhai</location>
,
<quantity id="4CD8C3B2FF8EA91BFE1FAE74FE17FDA8" box="[396,482,596,618]" metricMagnitude="3" metricUnit="m" metricValue="2.68" pageId="24" pageNumber="747" unit="m" value="2680.0">
<elevation id="000D8964FF8EA91BFE1FAE74FE17FDA8" box="[396,482,596,618]" metricMagnitude="3" metricUnit="m" metricValue="2.68" pageId="24" pageNumber="747" unit="m" value="2680.0">2680 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFE65AE74FD9BFDAB" box="[502,622,596,618]" pageId="24" pageNumber="747" value="2009-06-21">
<collectingDate id="EFDAB17FFF8EA91BFE65AE74FD9BFDAB" box="[502,622,596,618]" pageId="24" pageNumber="747" value="2009-06-21">21.vi.2009</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFD2DAE74FF38FD49" pageId="24" pageNumber="747">Yang Chao</collectorName>
</materialsCitation>
;
<materialsCitation id="3B48640AFF8EA91BFF4AAE52FE14FD66" collectingDate="1962-08-04" collectorName="Song Shimei" country="China" location="Lijiang Yulongshan" pageId="24" pageNumber="747" specimenCount="5" specimenCount-male="5" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFF4AAE52FEF5FD46" box="[217,256,626,648]" pageId="24" pageNumber="747" type="male">5♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFE9DAE53FE9FFD49" box="[270,362,627,648]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<location id="8EFF388CFF8EA91BFEE5AE52FDA3FD49" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFEE5AE52FDA3FD49" box="[374,598,626,648]" country="China" name="Lijiang Yulongshan" pageId="24" pageNumber="747" stateProvince="Yunnan">Lijiang Yulongshan</location>
, 9, 22, 23.vii.,
<date id="FF9E4897FF8EA91BFF03AEB1FEFCFD66" box="[144,265,657,679]" pageId="24" pageNumber="747" value="1962-08-04">
<collectingDate id="EFDAB17FFF8EA91BFF03AEB1FEFCFD66" box="[144,265,657,679]" pageId="24" pageNumber="747" value="1962-08-04">4.viii.1962</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFEDEAEB1FE14FD66" box="[333,481,657,679]" pageId="24" pageNumber="747">Song Shimei</collectorName>
</materialsCitation>
, IZCAS slide no. Geom- 00204;
<materialsCitation id="3B48640AFF8EA91BFF7EAE90FE2FFD25" collectingDate="1984-07-14" collectorName="Liu Dajun" country="China" elevation="3200" location="Lijiang Yulongshan" pageId="24" pageNumber="747" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFF7EAE90FEE6FD05" box="[237,275,688,709]" pageId="24" pageNumber="747" type="male">1♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFEB8AE90FE72FD04" box="[299,391,688,709]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<location id="8EFF388CFF8EA91BFE0FAE8FFD73FD04" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFE0FAE8FFD73FD04" box="[412,646,687,709]" country="China" name="Lijiang Yulongshan" pageId="24" pageNumber="747" stateProvince="Yunnan">Lijiang Yulongshan</location>
,
<quantity id="4CD8C3B2FF8EA91BFD0FAE90FD07FD04" box="[668,754,688,709]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="24" pageNumber="747" unit="m" value="3200.0">
<elevation id="000D8964FF8EA91BFD0FAE90FD07FD04" box="[668,754,688,709]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="24" pageNumber="747" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFF03AEEEFEFAFD25" box="[144,271,718,740]" pageId="24" pageNumber="747" value="1984-07-14">
<collectingDate id="EFDAB17FFF8EA91BFF03AEEEFEFAFD25" box="[144,271,718,740]" pageId="24" pageNumber="747" value="1984-07-14">14.vii.1984</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFECFAEEEFE2FFD25" box="[348,474,718,740]" pageId="24" pageNumber="747">Liu Dajun</collectorName>
</materialsCitation>
;
<materialsCitation id="3B48640AFF8EA91BFE7FAEEEFF3AFCE0" collectingDate="1984-07-11" collectingDateMax="1984-07-20" collectingDateMin="1984-07-11" collectorName="Liu Dajun &amp; Chen Yixin" country="China" elevation="2825" location="Lijiang Yulongshan" pageId="24" pageNumber="747" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D26A5DEFF8EA91BFE7FAEEEFDE7FD22" box="[492,530,718,739]" pageId="24" pageNumber="747" type="male">2♂</specimenCount>
<specimenCount id="9D26A5DEFF8EA91BFDB3AEEEFDB7FD22" box="[544,578,718,739]" pageId="24" pageNumber="747" type="female">2♀</specimenCount>
, same locality,
<quantity id="4CD8C3B2FF8EA91BFF03AECCFEDFFCC3" box="[144,298,748,770]" metricMagnitude="3" metricUnit="m" metricValue="2.825" metricValueMax="2.85" metricValueMin="2.8" pageId="24" pageNumber="747" unit="m" value="2825.0" valueMax="2850.0" valueMin="2800.0">
<elevation id="000D8964FF8EA91BFF03AECCFEDFFCC3" box="[144,298,748,770]" metricMagnitude="3" metricUnit="m" metricValue="2.825" metricValueMax="2.85" metricValueMin="2.8" pageId="24" pageNumber="747" unit="m" value="2825.0" valueMax="2850.0" valueMin="2800.0">2800–2850 m</elevation>
</quantity>
,
<date id="FF9E4897FF8EA91BFEA4AECDFE28FCC2" box="[311,477,749,771]" pageId="24" pageNumber="747" value="1984-07-11" valueMax="1984-07-20" valueMin="1984-07-11">
<collectingDate id="EFDAB17FFF8EA91BFEA4AECDFE28FCC2" box="[311,477,749,771]" pageId="24" pageNumber="747" value="1984-07-11" valueMax="1984-07-20" valueMin="1984-07-11">11–20.vii.1984</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFDB3AECDFD60FCC2" box="[544,661,749,771]" pageId="24" pageNumber="747">Liu Dajun</collectorName>
&amp;
<collectorName id="26D50B81FF8EA91BFD29AECDFF3AFCE0" pageId="24" pageNumber="747">Chen Yixin</collectorName>
</materialsCitation>
, IZCAS slide no. Geom-00203.
</paragraph>
<paragraph id="8B9F6E57FF8EA91BFF03AF72FEB1FC67" blockId="24.[144,759,850,934]" pageId="24" pageNumber="747">
<emphasis id="B954B245FF8EA91BFF03AF72FE0BFCA6" box="[144,510,850,871]" italics="true" pageId="24" pageNumber="747">Additional materials examined:</emphasis>
<materialsCitation id="3B48640AFF8EA91BFD9AAF72FECAFC67" collectingDate="1984-07-17" collectionCode="IZCAS" collectorName="Liu Dajun" country="China" elevation="2800" location="Lijiang Yulongshan" pageId="24" pageNumber="747" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan">
<collectingCountry id="F3372EC7FF8EA91BFD9AAF72FDA7FCA9" box="[521,594,850,872]" name="China" pageId="24" pageNumber="747">China</collectingCountry>
:
<specimenCount id="9D26A5DEFF8EA91BFDF3AF73FD73FCA6" box="[608,646,851,872]" pageId="24" pageNumber="747" type="male">2♂</specimenCount>
,
<collectingRegion id="49E4A0B5FF8EA91BFD04AF73FD06FCA9" box="[663,755,851,872]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
:
<location id="8EFF388CFF8EA91BFF03AF51FE84FC46" LSID="urn:lsid:plazi:treatment:0389DF41FF81A91BFEB2A909FDB6FB93:8EFF388CFF8EA91BFF03AF51FE84FC46" box="[144,369,881,903]" country="China" name="Lijiang Yulongshan" pageId="24" pageNumber="747" stateProvince="Yunnan">Lijiang Yulongshan</location>
,
<quantity id="4CD8C3B2FF8EA91BFEEDAF51FE21FC47" box="[382,468,881,902]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="24" pageNumber="747" unit="m" value="2800.0">
<elevation id="000D8964FF8EA91BFEEDAF51FE21FC47" box="[382,468,881,902]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="24" pageNumber="747" unit="m" value="2800.0">2800 m</elevation>
</quantity>
, 15,
<date id="FF9E4897FF8EA91BFD9FAF51FD7EFC46" box="[524,651,881,903]" pageId="24" pageNumber="747" value="1984-07-17">
<collectingDate id="EFDAB17FFF8EA91BFD9FAF51FD7EFC46" box="[524,651,881,903]" pageId="24" pageNumber="747" value="1984-07-17">17.vii.1984</collectingDate>
</date>
, coll.
<collectorName id="26D50B81FF8EA91BFD5CAF51FF23FC67" pageId="24" pageNumber="747">Liu Dajun</collectorName>
(
<collectionCode id="ED31F692FF8EA91BFF74AFB0FECDFC67" box="[231,312,912,934]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="24" pageNumber="747">IZCAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F6E57FF8EA91BFF03AFF7FE1EFC2C" blockId="24.[144,491,983,1005]" box="[144,491,983,1005]" pageId="24" pageNumber="747">
<emphasis id="B954B245FF8EA91BFF03AFF7FED4FC2D" box="[144,289,983,1004]" italics="true" pageId="24" pageNumber="747">Distribution:</emphasis>
<collectingCountry id="F3372EC7FF8EA91BFEB8AFF7FE85FC2C" box="[299,368,983,1005]" name="China" pageId="24" pageNumber="747">China</collectingCountry>
(
<collectingRegion id="49E4A0B5FF8EA91BFE11AFF7FE2AFC2C" box="[386,479,983,1005]" country="China" name="Yunnan" pageId="24" pageNumber="747">Yunnan</collectingRegion>
).
</paragraph>
<paragraph id="8B9F6E57FF8EA91BFF03A83EFDB6FB93" blockId="24.[144,758,1054,1107]" pageId="24" pageNumber="747">
<emphasis id="B954B245FF8EA91BFF03A83EFEF8FBF2" box="[144,269,1054,1075]" italics="true" pageId="24" pageNumber="747">Etymology:</emphasis>
The specific name is from the Latin
<emphasis id="B954B245FF8EA91BFD5EA83EFF3CFB93" italics="true" pageId="24" pageNumber="747">bellulus</emphasis>
, which means small and pretty.
</paragraph>
</subSubSection>
</treatment>
</document>
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