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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1075.75799" ID-GBIF-Dataset="b4a20b60-f573-4728-99f8-37d184182426" ID-PMC="PMC8671713" ID-Pensoft-Pub="1313-2970-1075-137" ID-Pensoft-UUID="C2FE64BE1F215D1C8311C7C8B064E6D1" ID-PubMed="35046753" ID-ZooBank="72BC3720940B448A95225CF226FDB5C6" ModsDocID="1313-2970-1075-137" checkinTime="1638937112639" checkinUser="pensoft" docAuthor="Cai 1, Yu-Jie, Tang 1, Liang &amp; Schillhammer 2, Harald" docDate="2021" docId="A3E046B440715ADAA433A2B432BD1590" docLanguage="en" docName="ZooKeys 1075: 137-174" docOrigin="ZooKeys 1075" docPubDate="2021-12-07" docSource="http://dx.doi.org/10.3897/zookeys.1075.75799" docTitle="Hesperosoma (Paramichrotus) parvioculatum Cai 1 &amp; Tang 1 &amp; Schillhammer 2 2021, sp. nov." docType="treatment" docUuid="616E3B86-58DA-4D44-BEBD-C30DE0FEB289" docUuidSource="ZooBank" docVersion="4" id="C2FE64BE1F215D1C8311C7C8B064E6D1" lastPageNumber="137" masterDocId="C2FE64BE1F215D1C8311C7C8B064E6D1" masterDocTitle="A review of the genus Hesperosoma Scheerpeltz (Coleoptera, Staphylinidae, Staphylininae) of China" masterLastPageNumber="174" masterPageNumber="137" pageNumber="137" updateTime="1668151200414" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the genus Hesperosoma Scheerpeltz (Coleoptera, Staphylinidae, Staphylininae) of China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cai 1, Yu-Jie</mods:namePart>
<mods:affiliation>College of Science, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tang 1, Liang</mods:namePart>
<mods:affiliation>College of Science, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schillhammer 2, Harald</mods:namePart>
<mods:affiliation>College of Science, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1075</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>137</mods:start>
<mods:end>174</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1075.75799</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1075.75799</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1075-137</mods:identifier>
<mods:identifier type="ZooBank">72BC3720940B448A95225CF226FDB5C6</mods:identifier>
<mods:identifier type="Pensoft-UUID">C2FE64BE1F215D1C8311C7C8B064E6D1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="191254524" LSID="urn:lsid:zoobank.org:act:616E3B86-58DA-4D44-BEBD-C30DE0FEB289" httpUri="http://treatment.plazi.org/id/A3E046B440715ADAA433A2B432BD1590" lastPageNumber="137" pageId="0" pageNumber="137">
<subSubSection pageId="0" pageNumber="137" type="nomenclature">
<paragraph pageId="0" pageNumber="137">
<taxonomicName LSID="http://zoobank.org/616E3B86-58DA-4D44-BEBD-C30DE0FEB289" authority="Cai 1 &amp; Tang 1 &amp; Schillhammer 2, 2021" authorityName="Cai 1 &amp; Tang 1 &amp; Schillhammer 2" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Hesperosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesperosoma (Paramichrotus) parvioculatum" order="Coleoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="parvioculatum" status="sp. nov." subGenus="Paramichrotus">Hesperosoma (Paramichrotus) parvioculatum</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">
<figureCitation captionStart="Figures 9095" captionStartId="F18" captionText="Figures 90 - 95. Hesperosoma parvioculatum sp. nov. 90 - 91 habitus 92 - 94 aedeagus, lateral (92) and ventral (93) views, paramere (94) 95 female abdominal tergite X. Scale bars: 2 mm (90 - 91), 0.2 mm (92 - 95)." figureDoi="10.3897/zookeys.1075.75799.figures90-95" httpUri="https://binary.pensoft.net/fig/620021" pageId="0" pageNumber="137">Figures 90-95</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="materials_examined">
<paragraph pageId="0" pageNumber="137">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="137">
<materialsCitation collectingDate="2014-05-26" collectorName="Yan" country="China" county="Yanling County" elevation="1620" latitude="26.302776" location="Nanfengmian" longLatPrecision="20" longitude="114.003334" specimenCount="♂" stateProvince="Hunan Prov." typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="137">
<emphasis italics="true" pageId="0" pageNumber="137">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="China">China</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="137">
<collectingRegion country="China" name="Hunan">Hunan Prov.</collectingRegion>
</emphasis>
<specimenCount></specimenCount>
; glued on a card with labels as follows: &quot;
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Hunan">Hunan</collectingRegion>
,
<collectingCounty>Yanling County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:A3E046B440715ADAA433A2B432BD1590:DEC2A1B5CFD30F45840FD721D4ED528C" country="China" county="Yanling County" latitude="26.302776" longLatPrecision="20" longitude="114.003334" name="Nanfengmian" stateProvince="Hunan Prov.">Nanfengmian</location>
;
<geoCoordinate degrees="26" direction="north" minutes="18" orientation="latitude" precision="15" seconds="10" value="26.302776">26°18'10&quot;N</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="00" orientation="longitude" precision="15" seconds="12" value="114.003334">114°00'12&quot;E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">1620 m</elevation>
</quantity>
;
<collectingDate value="2014-05-26">26 May 2014</collectingDate>
; Peng, Shen, Yu &amp;
<collectorName>Yan</collectorName>
leg.; mixed forest, leaf litter, wood sifted&quot; &quot;
<typeStatus>Holotype</typeStatus>
/
<taxonomicName authorityName="Cai 1 &amp; Tang 1 &amp; Schillhammer 2" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Hesperosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesperosoma (Paramichrotus) parvioculatum" order="Coleoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="parvioculatum" subGenus="Paramichrotus">Hesperosoma (Paramichrotus) parvioculatum</taxonomicName>
/ Cai, Tang &amp; Schillhammer&quot; [red handwritten label]; SHNU
</materialsCitation>
.
<materialsCitation collectingDate="2020-05-10" collectorName="Li, Wang" country="China" county="Yizhang County" elevation="1100" latitude="24.9275" location="Mang Mt." longLatPrecision="21" longitude="112.991104" specimenCount="1" specimenCount-male="1" stateProvince="Hunan Prov." typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="137">
<emphasis italics="true" pageId="0" pageNumber="137">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="China">China</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="137">
<collectingRegion country="China" name="Hunan">Hunan Prov.</collectingRegion>
</emphasis>
<specimenCount type="male">1♂</specimenCount>
;
<collectingCounty>Yizhang County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:A3E046B440715ADAA433A2B432BD1590:23271A1D8BBB07B27A1DBB2E7A3169EE" country="China" county="Yizhang County" latitude="24.9275" longLatPrecision="21" longitude="112.991104" name="Mang Mt." stateProvince="Hunan Prov.">Mang Mt.</location>
;
<geoCoordinate degrees="24" direction="north" minutes="55" orientation="latitude" precision="15" seconds="39" value="24.9275">24°55'39&quot;N</geoCoordinate>
,
<geoCoordinate degrees="112" direction="east" minutes="59" orientation="longitude" precision="15" seconds="28" value="112.991104">112°59'28&quot;E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000-1200 m</elevation>
</quantity>
;
<collectingDate value="2020-05-10">10 May 2020</collectingDate>
;
<collectorName>Li</collectorName>
&amp;
<collectorName>Wang</collectorName>
leg.; FIT; NMW
</materialsCitation>
<materialsCitation collectingDate="2020-07-17" collectorName="Li &amp; Wang" country="China" county="Yizhang County" elevation="1100" latitude="24.9275" location="Mang Mt." longLatPrecision="21" longitude="112.991104" specimenCount="1" specimenCount-female="1" stateProvince="Hunan Prov." typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
; same collection data as for the preceding; but
<collectingDate value="2020-07-17">17 Jul 2020</collectingDate>
; SHNU
</materialsCitation>
. -
<materialsCitation collectingDate="2013-07-08" collectorName="Dai, Peng, Xie" country="China" county="Wufeng County" elevation="1160" latitude="30.085835" location="Houhe Natural Reserve" longLatPrecision="20" longitude="110.55139" specimenCount="1" specimenCount-male="1" stateProvince="Hubei Prov." typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="137">
<collectingRegion country="China" name="Hubei">Hubei Prov.</collectingRegion>
</emphasis>
<specimenCount type="male">1♂</specimenCount>
,
<collectingCounty>Wufeng County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:A3E046B440715ADAA433A2B432BD1590:1BFBDD0797EB6B91B4E126450D319F50" country="China" county="Wufeng County" latitude="30.085835" longLatPrecision="20" longitude="110.55139" name="Houhe Natural Reserve" stateProvince="Hubei Prov.">Houhe Natural Reserve</location>
;
<geoCoordinate degrees="30" direction="north" minutes="05" orientation="latitude" precision="15" seconds="09" value="30.085835">30°05'09&quot;N</geoCoordinate>
,
<geoCoordinate degrees="110" direction="east" minutes="33" orientation="longitude" precision="15" seconds="05" value="110.55139">110°33'05&quot;E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">1160 m</elevation>
</quantity>
;
<collectingDate value="2013-07-08">08 Jul 2013</collectingDate>
;
<collectorName>Dai</collectorName>
,
<collectorName>Peng</collectorName>
&amp;
<collectorName>Xie</collectorName>
leg.; along path in a mixed forest; bamboo; leaf litter; sifted; SHNU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">Description.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Measurements of male</emphasis>
: BL: 10.61-13.33 mm, FL: 6.35-6.73 mm. HL: 1.58-1.79 mm, HW: 2.57-2.79 mm, EYL: 0.62-0.71 mm, TL: 0.86-0.96 mm, PL: 2.15-2.38 mm, PW: 1.95-2.11 mm, EL: 2.85-3.04 mm, EW: 2.83-3.10 mm. HW/HL: 1.48-1.63, TL/EYL: 1.35-1.39, PL/PW: 1.08-1.13, EL/EW: 0.98-1.01.
</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Measurements of female</emphasis>
: BL: 13.79 mm, FL: 6.82 mm. HL: 1.83 mm, HW: 2.384 mm, EYL: 0.71 mm, TL: 0.92 mm, PL: 2.42 mm, PW: 2.07 mm, EL: 3.01 mm, EW: 3.13 mm. HW/HL: 1.30, TL/EYL: 1.30, PL/PW: 1.17, EL/EW: 0.96.
</paragraph>
<paragraph pageId="0" pageNumber="137">Head, pronotum and scutellum black; elytra red, with a large black patch with faint bluish hue, nearly occupying posterior two thirds, continuing on to hypomeron, but not reaching ventro-lateral margin, posterior margin and suture narrowly yellowish-red; abdomen with segments III-V reddish, VI black with anterior margin narrowly reddish, VII black with posterior margin narrowly yellowish, VIII with proximal half yellowish and distal half black, IX dark brown, X black, narrowly, obscurely reddish at base; antennae with segments 1-7 black, base and apex of segment 1, 2 and base of segment 3 reddish, segments 8-11 creamy white; mandibles dark reddish-brown; maxillary palpi reddish, labial palpi with segments 1 and 3 reddish, segment 2 black brown; legs reddish-yellow, femora reddish-brown.</paragraph>
<paragraph pageId="0" pageNumber="137">Head 1.48-1.63 times as long as wide, rounded trapezoid, tempora narrowed behind eyes; dorsal surface coarsely and very densely punctate, punctures contiguous, including clypeus; antennae rather short, antennae with segments 4-7 weakly oblong, segments 8 and 10 about as long as wide.</paragraph>
<paragraph pageId="0" pageNumber="137">Pronotum 1.08-1.13 times as long as wide, widest at about level of large antero-lateral seta, narrowed towards base in weak concave arc, surface with dense and coarse punctation similar to that on head, with narrow impunctate mid-line, ground pubescence slightly more obvious than that on head; scutellum uniformly, densely punctate, space between punctures with very fine wavy microsculpture.</paragraph>
<paragraph pageId="0" pageNumber="137">Elytra as long as wide, along sides distinctly longer than pronotum, with slightly uneven surface, densely punctate and pubescent, with distinct depression between shoulders, scutellum and apical margin of elytra; pubescence yellow, long and dense along suture and posterior elytral margin.</paragraph>
<paragraph pageId="0" pageNumber="137">Abdominal tergites III-V with transverse basal depression and pair of short oblique basal carinae, punctation very sparse and rather coarse basally and laterally, gradually becoming finer towards apical margin, tergites VI-VIII entirely with fine and uniform punctation and pubescence, colour of pubescence corresponding with colour of integument underneath; legs long and slender.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Male.</emphasis>
Protarsomeres 1-4 moderately dilated, heart-shaped; sternite VII with the usual setose groove and patch of long yellow setae on median portion, posterior margin broadly emarginate at middle; sternite VIII with posterior margin emarginate at middle; aedeagus (Figs
<figureCitation captionStart="Figures 9095" captionStartId="F18" captionText="Figures 90 - 95. Hesperosoma parvioculatum sp. nov. 90 - 91 habitus 92 - 94 aedeagus, lateral (92) and ventral (93) views, paramere (94) 95 female abdominal tergite X. Scale bars: 2 mm (90 - 91), 0.2 mm (92 - 95)." figureDoi="10.3897/zookeys.1075.75799.figures90-95" httpUri="https://binary.pensoft.net/fig/620021" pageId="0" pageNumber="137">92-94</figureCitation>
) with apex of median lobe broad, anterior margin with gibbosities at middle (ventral view); paramere (Fig.
<figureCitation captionStart="Figures 9095" captionStartId="F18" captionText="Figures 90 - 95. Hesperosoma parvioculatum sp. nov. 90 - 91 habitus 92 - 94 aedeagus, lateral (92) and ventral (93) views, paramere (94) 95 female abdominal tergite X. Scale bars: 2 mm (90 - 91), 0.2 mm (92 - 95)." figureDoi="10.3897/zookeys.1075.75799.figures90-95" httpUri="https://binary.pensoft.net/fig/620021" pageId="0" pageNumber="137">94</figureCitation>
) with narrow apical portion, shorter than median lobe.
</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Female.</emphasis>
Tergite X (Fig.
<figureCitation captionStart="Figures 9095" captionStartId="F18" captionText="Figures 90 - 95. Hesperosoma parvioculatum sp. nov. 90 - 91 habitus 92 - 94 aedeagus, lateral (92) and ventral (93) views, paramere (94) 95 female abdominal tergite X. Scale bars: 2 mm (90 - 91), 0.2 mm (92 - 95)." figureDoi="10.3897/zookeys.1075.75799.figures90-95" httpUri="https://binary.pensoft.net/fig/620021" pageId="0" pageNumber="137">95</figureCitation>
) slightly asymmetrical with short apex.
</paragraph>
<caption doi="10.3897/zookeys.1075.75799.figures90-95" httpUri="https://binary.pensoft.net/fig/620021" pageId="0" pageNumber="137" start="Figures 9095" startId="F18">
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Figures 90-95.</emphasis>
<taxonomicName authorityName="Cai 1 &amp; Tang 1 &amp; Schillhammer 2" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Hesperosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hesperosoma parvioculatum" order="Coleoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="parvioculatum">
<emphasis italics="true" pageId="0" pageNumber="137">Hesperosoma parvioculatum</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="137">90-91</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="137">92-94</emphasis>
aedeagus, lateral (
<emphasis bold="true" pageId="0" pageNumber="137">92</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="137">93</emphasis>
) views, paramere (
<emphasis bold="true" pageId="0" pageNumber="137">94</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="137">95</emphasis>
female abdominal tergite X. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="137">90-91</emphasis>
), 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="137">92-95</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="etymology">
<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="137">The specific epithet refers to the small eyes of the new species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="distribution">
<paragraph pageId="0" pageNumber="137">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="137">China (Hubei and Hunan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="diagnosis">
<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="137">
Externally,
<taxonomicName lsidName="H. parvioculatum" pageId="0" pageNumber="137" rank="species" species="parvioculatum">
<emphasis italics="true" pageId="0" pageNumber="137">H. parvioculatum</emphasis>
</taxonomicName>
sp. nov. is virtually identical to
<taxonomicName lsidName="H. nigricolle" pageId="0" pageNumber="137" rank="species" species="nigricolle">
<emphasis italics="true" pageId="0" pageNumber="137">H. nigricolle</emphasis>
</taxonomicName>
from Myanmar, but differs mainly, aside from the distribution, by the shape of the aedeagus: paramere with narrow apical portion (paramere with broad apical portion, distinctly bilobed in
<taxonomicName lsidName="H. nigricolle" pageId="0" pageNumber="137" rank="species" species="nigricolle">
<emphasis italics="true" pageId="0" pageNumber="137">H. nigricolle</emphasis>
</taxonomicName>
); the wider head with HW/HL about 1.48-1.63 (1.35 in
<taxonomicName lsidName="H. nigricolle" pageId="0" pageNumber="137" rank="species" species="nigricolle">
<emphasis italics="true" pageId="0" pageNumber="137">H. nigricolle</emphasis>
</taxonomicName>
); posterior angles of the head being rounded (slightly prominent in
<taxonomicName lsidName="H. nigricolle" pageId="0" pageNumber="137" rank="species" species="nigricolle">
<emphasis italics="true" pageId="0" pageNumber="137">H. nigricolle</emphasis>
</taxonomicName>
); legs reddish-yellow, femora reddish-brown (legs reddish to yellowish-red, distal halves of femora black in
<taxonomicName lsidName="H. nigricolle" pageId="0" pageNumber="137" rank="species" species="nigricolle">
<emphasis italics="true" pageId="0" pageNumber="137">H. nigricolle</emphasis>
</taxonomicName>
). In appearance, it is also similar to
<taxonomicName lsidName="H. klapperichi" pageId="0" pageNumber="137" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="137">H. klapperichi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="H. alexpuchneri" pageId="0" pageNumber="137" rank="species" species="alexpuchneri">
<emphasis italics="true" pageId="0" pageNumber="137">H. alexpuchneri</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="H. guizhouense" pageId="0" pageNumber="137" rank="species" species="guizhouense">
<emphasis italics="true" pageId="0" pageNumber="137">H. guizhouense</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="H. yunnanense" pageId="0" pageNumber="137" rank="species" species="yunnanense">
<emphasis italics="true" pageId="0" pageNumber="137">H. yunnanense</emphasis>
</taxonomicName>
, but can be distinguished from
<taxonomicName lsidName="H. klapperichi" pageId="0" pageNumber="137" rank="species" species="klapperichi">
<emphasis italics="true" pageId="0" pageNumber="137">H. klapperichi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="H. alexpuchneri" pageId="0" pageNumber="137" rank="species" species="alexpuchneri">
<emphasis italics="true" pageId="0" pageNumber="137">H. alexpuchneri</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="H. guizhouense" pageId="0" pageNumber="137" rank="species" species="guizhouense">
<emphasis italics="true" pageId="0" pageNumber="137">H. guizhouense</emphasis>
</taxonomicName>
by the larger black elytral patch, occupying more than half of the elytral disc. From
<taxonomicName lsidName="H. yunnanense" pageId="0" pageNumber="137" rank="species" species="yunnanense">
<emphasis italics="true" pageId="0" pageNumber="137">H. yunnanense</emphasis>
</taxonomicName>
, it is distinguished by the smaller eyes with TL/EYL about 1.35-1.39 (1.18-1.29 in
<taxonomicName lsidName="H. yunnanense" pageId="0" pageNumber="137" rank="species" species="yunnanense">
<emphasis italics="true" pageId="0" pageNumber="137">H. yunnanense</emphasis>
</taxonomicName>
) and the wider head with HW/HL about 1.48-1.63 (1.26-1.40 in
<taxonomicName lsidName="H. yunnanense" pageId="0" pageNumber="137" rank="species" species="yunnanense">
<emphasis italics="true" pageId="0" pageNumber="137">H. yunnanense</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>