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<document id="5E614324B83FE2721B372B0CEC977143" ID-DOI="10.11646/zootaxa.3626.1.5" ID-GBIF-Dataset="135b7553-b75c-4020-b1a6-c689a05b7e66" ID-ISSN="1175-5326" ID-Zenodo-Dep="218625" ID-ZooBank="58AD6857-8CDD-4423-88D0-619CD8D793AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460581053656" checkinUser="plazi" docAuthor="Joshi, Jahnavi &amp; Edgecombe, Gregory D." docDate="2013" docId="4F7A87F2FFC0FFA9FF0BFF28FDB3F871" docLanguage="en" docName="zt03626p145.pdf" docOrigin="Zootaxa 3626 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Digitipes periyarensis Joshi &amp; Edgecombe, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="116" masterDocId="B343FF8AFFD1FFB8FF9CFFBFFFE3FFFC" masterDocTitle="Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity" masterLastPageNumber="145" masterPageNumber="99" pageNumber="116" updateTime="1698324300789" updateUser="plazi">
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<mods:title id="3AB7D0888BCEB73DF2B48684F6DEE712">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<treatment id="4F7A87F2FFC0FFA9FF0BFF28FDB3F871" ID-DOI="http://doi.org/10.5281/zenodo.3508271" ID-GBIF-Taxon="119563399" ID-Zenodo-Dep="3508271" LSID="urn:lsid:plazi:treatment:4F7A87F2FFC0FFA9FF0BFF28FDB3F871" httpUri="http://treatment.plazi.org/id/4F7A87F2FFC0FFA9FF0BFF28FDB3F871" lastPageNumber="116" pageId="17" pageNumber="116">
<subSubSection id="8FC9656FFFC0FFA9FF0BFF28FCF5FEE4" pageId="17" pageNumber="116" type="nomenclature">
<paragraph id="C76C36E4FFC0FFA9FF0BFF28FE06FF4D" blockId="17.[151,485,151,210]" box="[151,485,151,177]" pageId="17" pageNumber="116">
<heading id="9C248188FFC0FFA9FF0BFF28FE06FF4D" bold="true" box="[151,485,151,177]" fontSize="11" level="1" pageId="17" pageNumber="116" reason="1">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFF28FE06FF4D" bold="true" box="[151,485,151,177]" pageId="17" pageNumber="116">
<taxonomicName id="00D34D67FFC0FFA9FF0BFF28FE7EFF4D" box="[151,413,151,177]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="species" species="periyarensis" status="sp. nov.">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFF28FE7EFF4D" bold="true" box="[151,413,151,177]" italics="true" pageId="17" pageNumber="116">Digitipes periyarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFC0FFA9FE3FFF28FE06FF4D" box="[419,485,151,177]" pageId="17" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF0BFF05FEC4FF2E" blockId="17.[151,485,151,210]" box="[151,295,186,210]" pageId="17" pageNumber="116">
(
<figureCitation id="5FE82A61FFC0FFA9FF03FF05FEFCFF2E" box="[159,287,186,210]" captionStart-0="FIGURES 57 62" captionStart-1="FIGURES 63 67" captionStartId-0="18.[151,264,1777,1800]" captionStartId-1="19.[151,264,1159,1182]" captionTargetBox-0="[151,1435,219,1729]" captionTargetBox-1="[151,1435,193,1137]" captionTargetId-0="figure@18.[151,1436,219,1731]" captionTargetId-1="figure@19.[151,1436,193,1138]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." captionText-1="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri-0="https://zenodo.org/record/218635/files/figure.png" httpUri-1="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Figs 5767</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF0BFEBCFCF5FEE4" blockId="17.[151,790,258,280]" box="[151,790,258,280]" pageId="17" pageNumber="116">
<taxonomicName id="00D34D67FFC0FFA9FF0BFEBCFF17FEE4" box="[151,244,259,280]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFEBCFF17FEE4" box="[151,244,259,280]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
putative species 4. Joshi and Karanth, 2012: figs 2, 3.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF0BFEFAFBCEFE7D" pageId="17" pageNumber="116" type="materials_examined">
<paragraph id="C76C36E4FFC0FFA9FF0BFEFAFBCEFE7D" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFEFAFEBAFEA2" bold="true" box="[151,345,325,350]" pageId="17" pageNumber="116">
<typeStatus id="18688846FFC0FFA9FF0BFEFAFF31FEA1" box="[151,210,325,349]" pageId="17" pageNumber="116">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="18688846FFC0FFA9FEFCFEFAFE2BFEA1" box="[352,456,325,349]" pageId="17" pageNumber="116" type="holotype">Holotype</typeStatus>
CES
<date id="B36D1024FFC0FFA9FD9FFEFAFDB9FEA1" box="[515,602,325,349]" pageId="17" pageNumber="116" value="37-09-10">091037</date>
, female, from Periyar, Kerala,
<collectingCountry id="BFC47674FFC0FFA9FC28FEFAFC13FEA1" box="[948,1008,325,349]" name="India" pageId="17" pageNumber="116">India</collectingCountry>
, 9°58402 N 77°34902 E, leg. J. Joshi,
<date id="B36D1024FFC0FFA9FF0BFED6FF0CFE7D" box="[151,239,361,385]" pageId="17" pageNumber="116" value="2009-09">ix.2009</date>
.
<typeStatus id="18688846FFC0FFA9FF65FED5FEB8FE7E" box="[249,347,362,386]" pageId="17" pageNumber="116" type="paratype">Paratype</typeStatus>
CES
<date id="B36D1024FFC0FFA9FE09FED6FE0FFE7D" box="[405,492,361,385]" pageId="17" pageNumber="116" value="38-09-10">091038</date>
, female, from
<typeStatus id="18688846FFC0FFA9FD13FED5FD5CFE7E" box="[655,703,362,386]" pageId="17" pageNumber="116">type</typeStatus>
locality, same collection details.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FE32FDB7FE59" box="[201,596,397,421]" pageId="17" pageNumber="116" type="etymology">
<paragraph id="C76C36E4FFC0FFA9FF55FE32FDB7FE59" blockId="17.[151,1437,325,1933]" box="[201,596,397,421]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FE32FEB3FE59" bold="true" box="[201,336,397,421]" pageId="17" pageNumber="116">Etymology.</emphasis>
For the Periyar region.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FE0FFB46FDA5" pageId="17" pageNumber="116" type="diagnosis">
<paragraph id="C76C36E4FFC0FFA9FF55FE0FFB46FDA5" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FE0FFEA0FE35" bold="true" box="[201,323,432,457]" pageId="17" pageNumber="116">Diagnosis.</emphasis>
Cephalic plate slightly longer than wide. Basal three antennal articles glabrous dorsally. Forcipular tooth plates longer than wide. Paramedian sutures nearly complete on most sternites. All legs lacking tarsal, tibial and femoral spurs. Femur of ultimate leg with pair of longitudinal grooves on dorsal surface, the more medial of them contiguous with a longitudinal groove on the prefemur and tibia; tarsus of ultimate legs markedly more slender than short, robust tibia and femur. Coxopleural process short; pores relatively sparse.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FDDAFB2FF9D9" pageId="17" pageNumber="116" type="description">
<paragraph id="C76C36E4FFC0FFA9FF55FDDAFE8AFD39" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FDDAFEB8FD82" bold="true" box="[201,347,613,638]" pageId="17" pageNumber="116">Description.</emphasis>
Length to
<quantity id="002B9B01FFC0FFA9FE78FDDAFDDFFD81" box="[484,572,613,637]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="116" unit="mm" value="58.0">58 mm</quantity>
. 17 antennal articles; basal three articles glabrous dorsally, two glabrous ventrally. Anterior longitudinal median furrow at most ca 20% length of cephalic plate, considerably shorter in
<typeStatus id="18688846FFC0FFA9FF0BFD12FF1AFD39" box="[151,249,685,709]" pageId="17" pageNumber="116" type="holotype">holotype</typeStatus>
(
<figureCitation id="5FE82A61FFC0FFA9FE94FD12FEBFFD39" box="[264,348,685,709]" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FD6EFA92FCF1" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">Cephalic plate and T1 orange; anterior tergites mixed blue and yellow-brown; tergites in posterior half of trunk mostly light brown with blue margins; antennae and legs 120 pale yellow apart from pale blue tarsi on leg 20.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FCA6FF0EFC61" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Forcipular coxosternal tooth plates longer than wide, markedly so in
<typeStatus id="18688846FFC0FFA9FC46FCA6FBDFFCCD" box="[986,1084,793,817]" pageId="17" pageNumber="116" type="holotype">holotype</typeStatus>
(
<figureCitation id="5FE82A61FFC0FFA9FBD0FCA6FB31FCCD" box="[1100,1234,793,817]" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Figs 58, 59</figureCitation>
); four main teeth, not arranged in groups, the outer tooth smaller than the inner three; base of tooth plates defined by relative acute oblique sutures diverging at 105°125°. Trochanteroprefemoral process with indistinctly defined teeth along inner margin.
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FC16FA9DFC3D" blockId="17.[151,1437,325,1933]" box="[201,1406,937,962]" pageId="17" pageNumber="116">Second maxillary claw with slender accessory spurs. Article 2 of telopodite bearing a slender spine distally.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FC72FDE2FBAD" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Tergites with paramedian sutures complete from TT4 or 5, short suture confined to anterior edge of TT23. Tergites marginate from 5 or 6. Tergites smooth, lacking median ridge or keels (
<figureCitation id="5FE82A61FFC0FFA9FBF8FC4EFB5FFBF5" box="[1124,1212,1009,1033]" captionStart="FIGURES 63 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 66</figureCitation>
) apart from weak, incomplete median keel on tergite of ultimate leg-bearing segment; lateral part of posterior tergites with low, irregularly anastomosing ridges.
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FBE2FD3DFB65" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">Sternites with paramedian sutures nearly complete in most segments, especially in anterior part of trunk, those in posterior few segments ca 80% length of sternite.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FB1AFADCFAF9" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Tergite of ultimate leg-bearing segment with lateral margins gently converging posteriorly, posterolateral margins mostly straight, converging to a broad, evenly rounded posteromedial margin (
<figureCitation id="5FE82A61FFC0FFA9FB31FB76FAE5FB1D" box="[1197,1286,1225,1249]" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 61</figureCitation>
). Sternite of ultimate leg-bearing segment with sides converging posteriorly, posterior margin gently concave (
<figureCitation id="5FE82A61FFC0FFA9FB42FB52FAD0FAF9" box="[1246,1331,1261,1285]" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FAAEFB2FF9D9" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Coxopleural process short, barely inflected from posterolateral margin of coxopleuron in ventral view (
<figureCitation id="5FE82A61FFC0FFA9FAF3FAAEFF55FAB1" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 62</figureCitation>
), with two apical spines, lacking lateral spine(s). Pore field terminating strongly beneath dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFC0FFA9FEAFFAE6FE6BFA8D" box="[307,392,1369,1393]" captionStart="FIGURES 57 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 60</figureCitation>
); pores relative sparse; non-porose area on coxopleural process short. Ultimate leg prefemur with width at distal end nearly 40% its length, of nearly equal width along its length (
<figureCitation id="5FE82A61FFC0FFA9FBFBFAC2FB5EFA69" box="[1127,1213,1405,1429]" captionStart="FIGURES 63 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 64</figureCitation>
). Prefemoral spines robust, with a short, curved apex (
<figureCitation id="5FE82A61FFC0FFA9FD87FA1EFD93FA45" box="[539,624,1441,1465]" captionStart="FIGURES 63 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 63</figureCitation>
): VL 3, VM 2(3), DM 2. Femur with width at its distal end 4050% its length (
<figureCitation id="5FE82A61FFC0FFA9FF70FA7AFEA3FA21" box="[236,320,1477,1501]" captionStart="FIGURES 63 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 64</figureCitation>
). Dorsal, medial and lateral surfaces of prefemur and femur rugose, with large, shallow pits; similar pits adjacent to longitudinal groove on dorsomedial side of tibia (
<figureCitation id="5FE82A61FFC0FFA9FCEEFA56FC24F9FD" box="[882,967,1513,1537]" captionStart="FIGURES 63 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 65</figureCitation>
). Ultimate leg tarsus 1 2.5 times length of tarsus 2; tarsus 1 23.3 times longer than pretarsus; pretarsus with pair of short accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55F98EFDB3F871" pageId="17" pageNumber="116" type="discussion">
<paragraph id="C76C36E4FFC0FFA9FF55F98EFDC9F926" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55F98EFEADF9B6" bold="true" box="[201,334,1585,1610]" pageId="17" pageNumber="116">Discussion.</emphasis>
This species is the most morphologically distinct among Indian
<taxonomicName id="00D34D67FFC0FFA9FBBEF98DFB6AF9B5" box="[1058,1161,1586,1609]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FBBEF98DFB6AF9B5" box="[1058,1161,1586,1609]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
, readily identified by its complete absence of tarsal spurs, the long paramedian sutures on the sternites (nearly complete versus not more than 35% the length of the sternites in other Indian
<taxonomicName id="00D34D67FFC0FFA9FD48F9C5FCD8F96D" box="[724,827,1658,1681]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FD48F9C5FCD8F96D" box="[724,827,1658,1681]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
), and by the robust, dorsally-grooved prefemur, femur and tibia of the ultimate legs. Complete paramedian sutures on the sternites are known elsewhere in
<taxonomicName id="00D34D67FFC0FFA9FB6CF921FAB4F949" box="[1264,1367,1694,1717]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FB6CF921FAB4F949" box="[1264,1367,1694,1717]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
in the
<typeStatus id="18688846FFC0FFA9FF0BF97DFF24F926" box="[151,199,1730,1754]" pageId="17" pageNumber="116">type</typeStatus>
species,
<taxonomicName id="00D34D67FFC0FFA9FEB1F97DFDC9F926" authority="Attems." authorityName="Attems." box="[301,554,1729,1754]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="species" species="verdascens">
<emphasis id="F5A7EAF6FFC0FFA9FEB1F97DFE29F925" box="[301,458,1729,1753]" italics="true" pageId="17" pageNumber="116">D. verdascens</emphasis>
Attems.
</taxonomicName>
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55F95AFDB3F871" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
The two known specimens are both females so no data are at hand with respect to the condition of the femoral process in males. Despite its distinctive character states, its membership in
<taxonomicName id="00D34D67FFC0FFA9FC74F8B5FBACF8DD" box="[1000,1103,1802,1825]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FC74F8B5FBACF8DD" box="[1000,1103,1802,1825]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
is indicated by the support at the nodes in the phylogeny that unite it with Group A of Joshi and Karanth (2012) (posterior probability 0.94 for it grouping with
<taxonomicName id="00D34D67FFC0FFA9FEA0F8EDFE11F895" box="[316,498,1874,1897]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFC0FFA9FEA0F8EDFE11F895" box="[316,498,1874,1897]" italics="true" pageId="17" pageNumber="116">D. coonoorensis</emphasis>
</taxonomicName>
and
<taxonomicName id="00D34D67FFC0FFA9FDB5F8EDFD7FF895" box="[553,668,1873,1897]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFC0FFA9FDB5F8EDFD7FF895" box="[553,668,1873,1897]" italics="true" pageId="17" pageNumber="116">D. indicus</emphasis>
</taxonomicName>
) and that clade as a whole grouping with Groups B and C (posterior probability 1, bootstrap support 100%).
</paragraph>
</subSubSection>
</treatment>
</document>