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<mods:title id="B9A901F9FE4E1064E06DDE3D17E1A928">Songs in the genus Uromenus (Orthoptera: Tettigoniidae: Ephippigerini): A review with new information about some species</mods:title>
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<mods:affiliation id="F15336CC975E30B1DC9CA449B51EBA43">Nagy Fuvaros u. 6, 1084 Budapest, Hungary; saksup @ gmail. com; https: // orcid. org / 0000 - 0001 - 8008 - 749 X</mods:affiliation>
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<mods:affiliation id="971EE5C07AE8688D2DC6B73E4139FB63">Department of Systematic Zoology and Ecology, ELTE, Eötvös Loránd University, Budapest, 1117, Hungary; gergely. szovenyi @ ttk. elte. hu; https: // orcid. org / 0000 - 0001 - 9632 - 4066</mods:affiliation>
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<mods:namePart id="9A4D4CCCE5907A752F2DACF67C3AE143">Chobanov, Dragan P.</mods:namePart>
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<mods:affiliation id="D1B21FC6FEBB151ABAFFB5C90A5D7264">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Boulevard, 1000 Sofia, Bulgaria. dchobanov @ gmail. com; https: // orcid. org / 0000 - 0002 - 1642 - 0363</mods:affiliation>
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<heading id="331F81F67134FFE906B4F8F91554F889" bold="true" box="[151,579,1846,1873]" fontSize="11" level="1" pageId="12" pageNumber="105" reason="1">
<taxonomicName id="AFE84D197134FFE906B4F8F91554F889" ID-CoL="7DVMK" authority="Nadig, 1976" authorityName="Nadig" authorityYear="1976" box="[151,579,1846,1873]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="105" phylum="Arthropoda" rank="species" species="choumarae">
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<emphasis id="5A9CEA887134FFE906B4F8F916B0F888" bold="true" box="[151,423,1846,1873]" italics="true" pageId="12" pageNumber="105">Uromenus choumarae</emphasis>
<bibRefCitation id="0C794B6B7134FFE9078DF8F91554F889" author="Nadig, A." box="[430,579,1846,1872]" pageId="12" pageNumber="105" pagination="329 - 348" refId="ref10346" refString="Nadig, A. (1976) Beitrage zur Kenntnis der Orthopteren Marokkos: I. Die marokkanischen Arten der Gattung Ephippigerida I. Bolivar, 1903 und die ' Gruppe cockerelli ' der Gattung Uromenus (Steropleurus) I. Bolivar, 1878. Revue Suisse de Zoologie, 83, 329 - 348. https: // doi. org / 10.5962 / bhl. part. 91443" type="journal article" year="1976">Nadig, 1976</bibRefCitation>
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<paragraph id="6857369A7134FFE906B4F8B1163AF838" blockId="12.[151,1436,1918,2017]" pageId="12" pageNumber="105">
<materialsCitation id="D8803CC77134FFE906B4F8B1163FF838" ID-GBIF-Occurrence="3320086309" collectingDate="2013-06-10" collectorName="D. Chobanov. The &amp; In" country="Morocco" elevation="1940" location="Tizi'n Test" pageId="12" pageNumber="105" specimenCount="1">
<collectingCountry id="10FF760A7134FFE906B4F8B1163FF841" box="[151,296,1918,1944]" name="Morocco" pageId="12" pageNumber="105">MOROCCO</collectingCountry>
, south of
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pass, 30.85885 N, 008.37760 W,
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<quantity id="AF109B7F7134FFE905A4F8B114CBF840" box="[903,988,1918,1945]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="12" pageNumber="105" unit="m" value="1940.0">1940 m</quantity>
a.s.l.
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,
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<collectingDate id="0C12E9B27134FFE90206F8B113B7F841" box="[1061,1184,1918,1944]" pageId="12" pageNumber="105" value="2013-06-10">10 vi 2013</collectingDate>
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, leg.
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male stridulatory file carries about 92 teeth (
<figureCitation id="F0D32A1F7134FFE90495F86D15EDF864" box="[694,762,1954,1981]" captionStart="FIGURE 7" captionStartId="9.[151,260,1961,1987]" captionTargetBox="[154,1433,1292,1934]" captionTargetId="figure-59@9.[151,1436,1289,1937]" captionTargetPageId="9" captionText="FIGURE 7. Male stridulatory files in Uromenus. A—U. elegans (Crete), B—U. dyrrhachiacus, C—U. finoti, D—U. galvagnii, E—U. robustus, F—U. tobboganensis, G—U. melillae, H—U. innocentii, I—U. choumarae." figureDoi="http://doi.org/10.5281/zenodo.5027572" httpUri="https://zenodo.org/record/5027572/files/figure.png" pageId="12" pageNumber="105">Fig. 7</figureCitation>
;
<tableCitation id="256A03217134FFE90525F86D1459F865" box="[774,846,1954,1980]" captionStart="TABLE 2" captionStartId="16.[151,239,151,175]" captionTargetPageId="16" captionText="TABLE 2. Measurements of stridulatory files of the Uromenus species studied here" httpUri="http://table.plazi.org/id/3C9766127128FFF506B4FF581303FF76" pageId="12" pageNumber="105" tableUuid="3C9766127128FFF506B4FF581303FF76">Tab. 2</tableCitation>
).
<collectorName id="C51D534C7134FFE90541F86C146EF864" box="[866,889,1955,1981]" pageId="12" pageNumber="105">In</collectorName>
the middle of the file the inter-tooth intervals are about
<quantity id="AF109B7F7134FFE906FFF808163FF838" box="[220,296,1991,2017]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="12" pageNumber="105" unit="mm" value="77.0">77 µm</quantity>
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.
</paragraph>
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<emphasis id="5A9CEA887135FFE806E4FF581676FF68" bold="true" box="[199,353,151,177]" pageId="13" pageNumber="106">Morphology.</emphasis>
The species was described under (sub-)genus
<taxonomicName id="AFE84D197135FFE80553FF5814E9FF68" authorityName="Bolivar" authorityYear="1878" box="[880,1022,151,177]" class="Insecta" family="Tettigoniidae" genus="Steropleurus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="5A9CEA887135FFE80553FF5814E9FF68" box="[880,1022,151,177]" italics="true" pageId="13" pageNumber="106">Steropleurus</emphasis>
</taxonomicName>
and classified together with
<emphasis id="5A9CEA887135FFE80365FF58124CFF68" box="[1350,1371,151,177]" italics="true" pageId="13" pageNumber="106">S.</emphasis>
(now
<taxonomicName id="AFE84D197135FFE806B4FF74161CFF0D" authorityName="Bolivar" authorityYear="1878" box="[151,267,187,212]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="5A9CEA887135FFE806B4FF74161CFF0D" box="[151,267,187,212]" italics="true" pageId="13" pageNumber="106">Uromenus</emphasis>
</taxonomicName>
)
<emphasis id="5A9CEA887135FFE8073EFF74169DFF0C" box="[285,394,187,213]" italics="true" pageId="13" pageNumber="106">cockerelli</emphasis>
<bibRefCitation id="0C794B6B7135FFE807B0FF741539FF0C" author="Uvarov, B. P." box="[403,558,187,213]" pageId="13" pageNumber="106" pagination="210 - 214" refId="ref11070" refString="Uvarov, B. P. (1931) Orthoptera collected by Prof. T. D. A. Cockerell in Morocco. Buletin de la Societe des Sciences naturelles du Maroc, 10, 210 - 214. [1930]" type="journal article" year="1931">
Uvarov,
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in the
<emphasis id="5A9CEA887135FFE804A2FF7415E2FF0C" box="[641,757,187,213]" italics="true" pageId="13" pageNumber="106">Cockerelli</emphasis>
Species group (
<bibRefCitation id="0C794B6B7135FFE8058DFF74132EFF0C" author="Nadig, A." box="[942,1081,186,213]" pageId="13" pageNumber="106" pagination="109 - 183" refId="ref10461" refString="Nadig, A. (1995) Die marokkanischen Arten der Gattung Steropleurus Bolivar, 1878 (Orthoptera Ephippigeridae). Atti dell'Accademia Roveretana degli Agiati, Atti anni accademici, 244 [1994] 4 B, 109 - 183." type="journal article" year="1995">Nadig 1995</bibRefCitation>
).
<taxonomicName id="AFE84D197135FFE8026DFF7413E6FF0C" authorityName="Nadig" authorityYear="1976" box="[1102,1265,187,213]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="choumarae">
<emphasis id="5A9CEA887135FFE8026DFF7413E6FF0C" box="[1102,1265,187,213]" italics="true" pageId="13" pageNumber="106">U. choumarae</emphasis>
</taxonomicName>
is typical with very long incurved male cerci with longer, basally very wide basal part (before the tooth) and much shorter and narrower apical part bearing a small sharp tooth at the tip (
<figureCitation id="F0D32A1F7135FFE80522FECC147DFEC4" box="[769,874,259,285]" captionStart="FIGURE 10" captionStartId="14.[151,260,1927,1953]" captionTargetBox="[173,1414,378,1902]" captionTargetId="figure-109@14.[173,1414,378,1903]" captionTargetPageId="14" captionText="FIGURE 10. Cerci of the Moroccan species of Uromenus studied here and of U. dyrrhachiacus. A—U. finoti, B—U. galvagnii, C—U. robustus, D—U. tobboganensis, E—U. melillae, F—U. innocentii, G—U. choumarae, H—U. dyrrhachiacus" figureDoi="http://doi.org/10.5281/zenodo.5027580" httpUri="https://zenodo.org/record/5027580/files/figure.png" pageId="13" pageNumber="106">Fig. 10G</figureCitation>
<figureCitation id="F0D32A1F7135FFE80549FECC1484FEC4" box="[874,915,259,285]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1871,1895]" captionStartId-1="4.[151,250,1977,2001]" captionTargetBox-0="[151,1436,561,1848]" captionTargetBox-1="[163,1428,327,1941]" captionTargetId-0="figure-106@2.[151,1436,561,1848]" captionTargetId-1="figure-107@4.[151,1436,309,1954]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. General appearance and habitats of Uromenus species. A—U. elegans (Italy, Lazio), BD—U. elegans (Crete; photos B and C taken by Ionuţ Iorgu), E, F—U. dyrrhachiacus, G—U. agarenus, HJ—U. finoti, K, L—U. robustus, M, N—U. tobboganensis, O—U. maroccanus, P, Q—U. melillae, R—U. innocentii, S, T—U. choumarae." captionText-1="FIGURE 2. Oscillograms of male calling songs of the Uromenus species studied here, 10 second sections." figureDoi-0="http://doi.org/10.5281/zenodo.5027546" figureDoi-1="http://doi.org/10.5281/zenodo.5027550" httpUri-0="https://zenodo.org/record/5027546/files/figure.png" httpUri-1="https://zenodo.org/record/5027550/files/figure.png" pageId="13" pageNumber="106">12</figureCitation>
). Male 10th tergum, epiproct and titillators are shown on
<figureCitation id="F0D32A1F7135FFE80729FEE81667FE98" box="[266,368,295,321]" captionStart="FIGURE 10" captionStartId="14.[151,260,1927,1953]" captionTargetBox="[173,1414,378,1902]" captionTargetId="figure-109@14.[173,1414,378,1903]" captionTargetPageId="14" captionText="FIGURE 10. Cerci of the Moroccan species of Uromenus studied here and of U. dyrrhachiacus. A—U. finoti, B—U. galvagnii, C—U. robustus, D—U. tobboganensis, E—U. melillae, F—U. innocentii, G—U. choumarae, H—U. dyrrhachiacus" figureDoi="http://doi.org/10.5281/zenodo.5027580" httpUri="https://zenodo.org/record/5027580/files/figure.png" pageId="13" pageNumber="106">Fig. 10G</figureCitation>
<figureCitation id="F0D32A1F7135FFE80753FEE81680FE98" box="[368,407,295,321]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1871,1895]" captionStartId-1="4.[151,250,1977,2001]" captionTargetBox-0="[151,1436,561,1848]" captionTargetBox-1="[163,1428,327,1941]" captionTargetId-0="figure-106@2.[151,1436,561,1848]" captionTargetId-1="figure-107@4.[151,1436,309,1954]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. General appearance and habitats of Uromenus species. A—U. elegans (Italy, Lazio), BD—U. elegans (Crete; photos B and C taken by Ionuţ Iorgu), E, F—U. dyrrhachiacus, G—U. agarenus, HJ—U. finoti, K, L—U. robustus, M, N—U. tobboganensis, O—U. maroccanus, P, Q—U. melillae, R—U. innocentii, S, T—U. choumarae." captionText-1="FIGURE 2. Oscillograms of male calling songs of the Uromenus species studied here, 10 second sections." figureDoi-0="http://doi.org/10.5281/zenodo.5027546" figureDoi-1="http://doi.org/10.5281/zenodo.5027550" httpUri-0="https://zenodo.org/record/5027546/files/figure.png" httpUri-1="https://zenodo.org/record/5027550/files/figure.png" pageId="13" pageNumber="106">12</figureCitation>
and
<figureCitation id="F0D32A1F7135FFE807EDFEE81523FE98" box="[462,564,295,321]" captionStart="FIGURE 11" captionStartId="15.[151,260,1942,1968]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure-19@15.[278,1310,179,1918]" captionTargetPageId="15" captionText="FIGURE 11. Titillators of U. dyrrhachiacus and of the Moroccan species of Uromenus studied here. A—U. dyrrhachiacus, B—U. finoti, C—U. galvagnii, D—U. robustus, E—U. tobboganensis, F—U. melillae, G—U. innocentii, H—U. choumarae. Scale 2 mm except for U. dyrrhachiacus - 0.9 mm." figureDoi="http://doi.org/10.5281/zenodo.5027603" httpUri="https://zenodo.org/record/5027603/files/figure.png" pageId="13" pageNumber="106">Fig. 11H</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="3C9766127135FFE806B4F8F51340F845" ID-DOI="http://doi.org/10.5281/zenodo.5027578" ID-Zenodo-Dep="5027578" httpUri="https://zenodo.org/record/5027578/files/figure.png" pageId="13" pageNumber="106" startId="13.[151,260,1850,1876]" targetBox="[158,1425,354,1810]" targetPageId="13">
<paragraph id="6857369A7135FFE806B4F8F51340F845" blockId="13.[151,1436,1850,1948]" pageId="13" pageNumber="106">
<emphasis id="5A9CEA887135FFE806B4F8F51632F88D" bold="true" box="[151,293,1850,1876]" pageId="13" pageNumber="106">FIGURE 9.</emphasis>
Female tegmina in
<emphasis id="5A9CEA887135FFE80436F8F41587F88D" box="[533,656,1851,1876]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE80436F8F4159BF88D" authorityName="Bolivar" authorityYear="1878" box="[533,652,1851,1876]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="genus">Uromenus</taxonomicName>
.
</emphasis>
A—
<taxonomicName id="AFE84D197135FFE804EFF8F414B1F88D" authority="(Crete)" baseAuthorityName="Crete" baseAuthorityYear="1853" box="[716,934,1850,1876]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="5A9CEA887135FFE804EFF8F4145FF88D" box="[716,840,1850,1876]" italics="true" pageId="13" pageNumber="106">U. elegans</emphasis>
(Crete)
</taxonomicName>
, B—
<emphasis id="5A9CEA887135FFE805C7F8F413A4F88D" box="[996,1203,1850,1876]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE805C7F8F413B8F88D" baseAuthorityName="Karny" baseAuthorityYear="1918" box="[996,1199,1850,1876]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="dyrrhachiacus">U. dyrrhachiacus</taxonomicName>
,
</emphasis>
C
<emphasis id="5A9CEA887135FFE802F2F8F41282F88D" box="[1233,1429,1850,1876]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE802CCF8F41282F88D" baseAuthorityName="Serville" baseAuthorityYear="1838" box="[1263,1429,1850,1876]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="rugosicollis">U. rugosicollis</taxonomicName>
</emphasis>
, D—
<emphasis id="5A9CEA887135FFE806E4F890164EF8A1" box="[199,345,1887,1912]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE806E4F8901642F8A1" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1882" box="[199,341,1887,1912]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="agarenus">U. agarenus</taxonomicName>
,
</emphasis>
E—
<emphasis id="5A9CEA887135FFE807ADF89016E4F8A1" box="[398,499,1886,1912]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE807ADF89016F8F8A1" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1882" box="[398,495,1886,1912]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="finoti">U. finoti</taxonomicName>
,
</emphasis>
F
<emphasis id="5A9CEA887135FFE80429F89015FEF8A1" box="[522,745,1886,1912]" italics="true" pageId="13" pageNumber="106">
<taxonomicName id="AFE84D197135FFE8040BF89015FEF8A1" authorityName="Nadig" authorityYear="1994" box="[552,745,1886,1912]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="tobboganensis">U. tobboganensis</taxonomicName>
</emphasis>
and G—
<taxonomicName id="AFE84D197135FFE80572F89014F3F8A1" baseAuthorityName="E.Bonnet &amp; Finot" baseAuthorityYear="1885" box="[849,996,1887,1912]" class="Insecta" family="Tettigoniidae" genus="Uromenus" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="106" phylum="Arthropoda" rank="species" species="innocentii">
<emphasis id="5A9CEA887135FFE80572F89014F3F8A1" box="[849,996,1887,1912]" italics="true" pageId="13" pageNumber="106">U. innocentii</emphasis>
</taxonomicName>
. Note the long stridulatory vein on the upper side of the right tegmen crossing the mirror-like translucent tegmen membrane.
</paragraph>
</caption>
<subSubSection id="20F265117136FFEB06E4FF591600FEBD" pageId="14" pageNumber="107" type="biology_ecology">
<paragraph id="6857369A7136FFEB06E4FF591600FEBD" blockId="14.[151,1437,150,356]" pageId="14" pageNumber="107">
<emphasis id="5A9CEA887136FFEB06E4FF59163BFF69" bold="true" box="[199,300,150,176]" pageId="14" pageNumber="107">Ecology.</emphasis>
The species was collected close to its
<typeStatus id="B75388387136FFEB04D7FF581433FF68" box="[756,804,151,177]" pageId="14" pageNumber="107">type</typeStatus>
locality (Tizin Test pass) where it was common. The nymphs and young adults kept on bushes of
<taxonomicName id="AFE84D197136FFEB04BAFF7414A0FF0D" box="[665,951,186,213]" class="Magnoliopsida" family="Fabaceae" genus="Argyrocytisus" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="107" phylum="Tracheophyta" rank="species" species="battandieri">
<emphasis id="5A9CEA887136FFEB04BAFF7414A0FF0D" box="[665,951,186,213]" italics="true" pageId="14" pageNumber="107">Argyrocytisus battandieri</emphasis>
</taxonomicName>
and on
<taxonomicName id="AFE84D197136FFEB0234FF7513BFFF0C" box="[1047,1192,186,213]" class="Mammalia" family="Tenrecidae" genus="Echinops" kingdom="Animalia" order="Afrosoricida" pageId="14" pageNumber="107" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="5A9CEA887136FFEB0234FF751368FF0D" box="[1047,1151,186,212]" italics="true" pageId="14" pageNumber="107">Echinops</emphasis>
sp.
</taxonomicName>
at the steep southern slopes of the southwestern High Atlas Mountains (
<figureCitation id="F0D32A1F7136FFEB04E7FF11140FFF21" box="[708,792,222,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1871,1895]" captionTargetBox="[151,1436,561,1848]" captionTargetId="figure-106@2.[151,1436,561,1848]" captionTargetPageId="2" captionText="FIGURE 1. General appearance and habitats of Uromenus species. A—U. elegans (Italy, Lazio), BD—U. elegans (Crete; photos B and C taken by Ionuţ Iorgu), E, F—U. dyrrhachiacus, G—U. agarenus, HJ—U. finoti, K, L—U. robustus, M, N—U. tobboganensis, O—U. maroccanus, P, Q—U. melillae, R—U. innocentii, S, T—U. choumarae." figureDoi="http://doi.org/10.5281/zenodo.5027546" httpUri="https://zenodo.org/record/5027546/files/figure.png" pageId="14" pageNumber="107">Fig. 1T</figureCitation>
). The high altitude of its locality with south exposition defines cooler Köppen climate
<typeStatus id="B75388387136FFEB07FAFECC151EFEC4" box="[473,521,259,285]" pageId="14" pageNumber="107">type</typeStatus>
CSb (Warm-summer mediterranean) towards DSb (Warm-summer mediterranean continental)(
<bibRefCitation id="0C794B6B7136FFEB0701FEE916E3FE99" author="Verner, D. &amp; Treguer, D. &amp; Redwood, J. &amp; Christensen, J. &amp; McDonnell, R. &amp; Elbert, C. &amp; Konishi, Y. &amp; Belghazi, S." box="[290,500,294,321]" pageId="14" pageNumber="107" refId="ref11113" refString="Verner, D., Treguer, D., Redwood, J., Christensen, J., McDonnell, R., Elbert, C., Konishi, Y. &amp; Belghazi, S. (2018) Climate Variability, Drought, and Drought Management in Morocco's Agricultural Sector. World Bank Group, Washington, D. C. Available from: http: // documents. worldbank. org / curated / en / 353801538414553978 / Climate-Variability-Drought-and-Drought- Management-in-Moroccos-Agricultural-Sector (accessed 8 June 2021) https: // doi. org / 10.1596 / 30603" type="url" year="2018">
Verner
<emphasis id="5A9CEA887136FFEB075BFEE816BEFE99" box="[376,425,294,320]" italics="true" pageId="14" pageNumber="107">et al</emphasis>
. 2018
</bibRefCitation>
). The species is possibly overlooked and may have broad range in the southwestern High Atlas.
</paragraph>
</subSubSection>
</treatment>
</document>