treatments-xml/data/9C/34/DD/9C34DD3CC009FF96FF77FCD9436E4FB3.xml

<document id="7F44226FB5332E0CF21C2690A0FD498D" ID-CLB-Dataset="46187" ID-DOI="10.11646/zootaxa.3780.1.2" ID-GBIF-Dataset="b38919fb-f89d-42c8-8395-f6b726986951" ID-ISSN="1175-5326" ID-Zenodo-Dep="225615" ID-ZooBank="810ED246-F468-4081-A3F0-54F23A6BC895" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460399085331" checkinUser="plazi" docAuthor="Rojas-Runjaic, Fernando J. M., Delgado, J. Amanda &amp; Guayasamin, Juan M." docDate="2014" docId="9C34DD3CC009FF96FF77FCD9436E4FB3" docLanguage="en" docName="zt03780p050.pdf" docOrigin="Zootaxa 3780 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristimantis munozi Rojas-Runjaic, Delgado &amp; Guayasamin, 2014, sp. nov." docType="treatment" docVersion="10" masterDocId="600DA544C00BFF9CFFE0FFBF41554A5C" masterDocTitle="A new rainfrog of the Pristimantis myersi Group (Amphibia, Craugastoridae) from Volcán Pichincha, Ecuador" updateTime="1698299006133" updateUser="ExternalLinkService">
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<mods:title id="B7FEAA62EB92ADCB445B4A876DFF52FB">A new rainfrog of the Pristimantis myersi Group (Amphibia, Craugastoridae) from Volcán Pichincha, Ecuador</mods:title>
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<mods:namePart id="393F0541DEBBED285362B0494818AFDD">Rojas-Runjaic, Fernando J. M.</mods:namePart>
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<mods:namePart id="E8BB1ADF8402A7200E7B82FB3715D48A">Delgado, J. Amanda</mods:namePart>
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<mods:namePart id="E01AC710AA8B36BCDE13D08B6B8F4043">Guayasamin, Juan M.</mods:namePart>
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<mods:date id="880797A89A8FA2077927F77DEFC998DE">2014</mods:date>
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<treatment id="9C34DD3CC009FF96FF77FCD9436E4FB3" ID-DOI="http://doi.org/10.5281/zenodo.5664296" ID-GBIF-Taxon="119419016" ID-Zenodo-Dep="5664296" LSID="urn:lsid:plazi:treatment:9C34DD3CC009FF96FF77FCD9436E4FB3" httpUri="http://treatment.plazi.org/id/9C34DD3CC009FF96FF77FCD9436E4FB3" lastPageId="10" pageId="2">
<subSubSection id="5C873FA1C009FF9EFF77FCD940AD4E51" pageId="2" type="nomenclature">
<paragraph id="14226C2AC009FF9EFF77FCD940BA49DC" blockId="2.[151,495,870,930]" box="[151,495,870,896]" pageId="2">
<heading id="4F6ADB46C009FF9EFF77FCD940BA49DC" bold="true" box="[151,495,870,896]" fontSize="11" level="1" pageId="2" reason="1">
<emphasis id="26E9B038C009FF9EFF77FCD940BA49DC" bold="true" box="[151,495,870,896]" pageId="2">
<taxonomicName id="D39D17A9C009FF9EFF77FCD940DF49DC" box="[151,394,870,896]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" phylum="Chordata" rank="species" species="munozi" status="sp. nov.">
<emphasis id="26E9B038C009FF9EFF77FCD940DF49DC" bold="true" box="[151,394,870,896]" italics="true" pageId="2">Pristimantis munozi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DDA0D43C009FF9EFE72FCD940BA49DC" box="[402,495,870,896]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="14226C2AC009FF9EFF77FC3640DA49FE" blockId="2.[151,495,870,930]" box="[151,399,905,930]" pageId="2">
(
<figureCitation id="8CA670AFC009FF9EFF7EFC36407349FE" box="[158,294,905,930]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1800,1822]" captionStartId-1="5.[151,250,1524,1546]" captionStartId-2="6.[151,250,1701,1723]" captionTargetBox-0="[151,1436,517,1778]" captionTargetBox-1="[151,1436,301,1502]" captionTargetBox-2="[151,1436,481,1677]" captionTargetId-0="figure@4.[151,1436,517,1778]" captionTargetId-1="figure@5.[151,1436,301,1503]" captionTargetId-2="figure@6.[151,1436,481,1679]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." captionText-1="FIGURE 2. Pristimantis munozi sp. nov. Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante." captionText-2="FIGURE 3. Pristimantis munozi sp. nov. (holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a, b, e) &amp; F. J. M. Rojas- Runjaic (c, d)." httpUri-0="https://zenodo.org/record/225616/files/figure.png" httpUri-1="https://zenodo.org/record/225617/files/figure.png" httpUri-2="https://zenodo.org/record/225618/files/figure.png" pageId="2">Figures 1–3</figureCitation>
, 
<tableCitation id="591F5991C009FF9EFED3FC3640D149FE" box="[307,388,905,930]" captionStart="TABLE 1" captionStartId="8.[151,239,476,499]" captionTargetBox="[161,1115,558,1336]" captionTargetPageId="8" captionText="TABLE 1. Morphometric variation in adult males (n = 8) of Pristimantis munozi. X? ± SD: mean ± standard deviation; Min – Max: minimum – maximum." httpUri="http://table.plazi.org/id/40E23CA2C003FF94FF77FE6340AC484F" pageId="2" tableUuid="40E23CA2C003FF94FF77FE6340AC484F">Table 1</tableCitation>
)
</paragraph>
<paragraph id="14226C2AC009FF9EFF77FC6E40AD4E51" blockId="2.[151,513,977,1038]" pageId="2">English name: Muñoz’ Rainfrog Spanish name: Cutín de Muñoz</paragraph>
</subSubSection>
<subSubSection id="5C873FA1C009FF9EFF77FB8242274EBA" pageId="2" type="materials_examined">
<paragraph id="14226C2AC009FF9EFF77FB8240644EC2" blockId="2.[151,1436,1085,2010]" pageId="2">
<emphasis id="26E9B038C009FF9EFF77FB82405E4E0A" bold="true" box="[151,267,1085,1110]" pageId="2">
<typeStatus id="CB26D288C009FF9EFF77FB8240534E0A" box="[151,262,1085,1110]" pageId="2" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male, 
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1782 (field number ANF 1129), from Reserva Verdecocha, Province of Pichincha, 
<collectingCountry id="6C8A2CBAC009FF9EFF77FBDE41A14E26" box="[151,244,1121,1146]" name="Ecuador" pageId="2">Ecuador</collectingCountry>
(
<geoCoordinate id="71A90AEDC009FF9EFEE3FBDE402A4E26" box="[259,383,1121,1146]" direction="south" orientation="latitude" pageId="2" precision="1" value="-0.096361116">0°5'46.9&quot;S</geoCoordinate>
, 
<geoCoordinate id="71A90AEDC009FF9EFE6BFBDE43794E26" box="[395,556,1121,1146]" direction="west" orientation="longitude" pageId="2" precision="1" value="-78.60425">78°36'15.3&quot;W</geoCoordinate>
; elevation 
<quantity id="D365C1CFC009FF9EFD45FBDE43A84E26" box="[677,765,1121,1146]" metricMagnitude="3" metricUnit="m" metricValue="2.851" pageId="2" unit="m" value="2851.0">2851 m</quantity>
), collected on 21 of 
<date id="60234AEAC009FF9EFC01FBDE45044E26" box="[993,1105,1121,1146]" pageId="2" value="2012-07">July 2012</date>
, by F.J.M. Rojas-Runjaic and J.A. Delgado.
</paragraph>
<paragraph id="14226C2AC009FF9EFF27FB1642274EBA" blockId="2.[151,1436,1085,2010]" pageId="2">
<emphasis id="26E9B038C009FF9EFF27FB16402E4E9E" bold="true" box="[199,379,1193,1218]" pageId="2">Paratopotypes.</emphasis>
Seven adult males, 
<collectionCode id="728CF4EFC009FF9EFD83FB1543E94E9D" box="[611,700,1194,1217]" pageId="2">MZUTI</collectionCode>
<date id="60234AEAC009FF9EFD25FB1542134E9E" box="[709,838,1194,1218]" pageId="2" value="1777" valueMax="1781">1777–1781</date>
, 
<date id="60234AEAC009FF9EFCB3FB1542844E9E" box="[851,977,1194,1218]" pageId="2" value="1783" valueMax="1784">1783–1784</date>
(field numbers ANF 
<date id="60234AEAC009FF9EFB2BFB15441C4E9E" box="[1227,1353,1194,1218]" pageId="2" value="1124" valueMax="1128">1124–1128</date>
, 
<date id="60234AEAC009FF9EFAB7FB15419B4EBA" pageId="2" value="1130" valueMax="1131">1130– 1131</date>
respectively), with the same collection data as the 
<typeStatus id="CB26D288C009FF9EFCE8FB72423B4EBA" box="[776,878,1229,1254]" pageId="2" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="5C873FA1C009FF9FFF27FB4E435F4D58" lastPageId="3" pageId="2" type="diagnosis">
<paragraph id="14226C2AC009FF9FFF27FB4E447D4A89" blockId="2.[151,1436,1085,2010]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" pageId="2">
<emphasis id="26E9B038C009FF9EFF27FB4E40144F56" bold="true" box="[199,321,1265,1290]" pageId="2">Diagnosis.</emphasis>
A species of the genus 
<taxonomicName id="D39D17A9C009FF9EFDA7FB4D43854F55" box="[583,720,1266,1289]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" phylum="Chordata" rank="genus">
<emphasis id="26E9B038C009FF9EFDA7FB4D43854F55" box="[583,720,1266,1289]" italics="true" pageId="2">Pristimantis</emphasis>
</taxonomicName>
, assigned to the 
<taxonomicName id="D39D17A9C009FF9EFC69FB4D45364F55" box="[905,1123,1266,1289]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" phylum="Chordata" rank="species" species="myersi">
<emphasis id="26E9B038C009FF9EFC69FB4D45364F55" box="[905,1123,1266,1289]" italics="true" pageId="2">Pristimantis myersi</emphasis>
</taxonomicName>
species group (
<emphasis id="26E9B038C009FF9EFAF1FB4D441A4F55" box="[1297,1359,1266,1289]" italics="true" pageId="2">sensu</emphasis>
<bibRefCitation id="700C11DBC009FF9EFAB6FB4E40324F72" author="Lynch" pageId="2" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology, and biogeography. The University of Kansas Natural History Museum, Special Publications, 23, 1 - 236. http: // dx. doi. org / 10.5962 / bhl. title. 16289" type="journal article" year="1997">Lynch &amp; Duellman 1997</bibRefCitation>
, and 
<bibRefCitation id="700C11DBC009FF9EFE45FAAA43D54F72" author="Hedges" box="[421,640,1301,1326]" pageId="2" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New World direct-developing frogs (Anura: Terrarana): Molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">
Hedges 
<emphasis id="26E9B038C009FF9EFDE1FAA943664F71" box="[513,563,1301,1325]" italics="true" pageId="2">et al</emphasis>
. 2008
</bibRefCitation>
), by its small size (SVL &lt;
<quantity id="D365C1CFC009FF9EFC55FAA9455F4F72" box="[949,1034,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="2" unit="mm" value="28.0">28 mm</quantity>
), robust body, relative narrow head and short snout, absence of cranial crests, tympanum differentiated, vocal slits and vomerine teeth present, limbs short, FI shorter than FII, TV slightly longer than TIII and not extending to the proximal edge of the distal subarticular tubercle of TIV, digital discs narrow and rounded. The new species is readily diagnosable by the combination of the following characters: (1) body small (SVL in adult males = 
<quantity id="D365C1CFC009FF9EFBF3FA1945EC4FE1" box="[1043,1209,1446,1470]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" metricValueMax="1.97" metricValueMin="1.49" pageId="2" unit="mm" value="17.3" valueMax="19.7" valueMin="14.9">14.9–19.7 mm</quantity>
; females unknown); (2) dorsal skin shagreen, with a barely visible middorsal raphe, scapular and dorsolateral folds; (3) ventral skin areolate on chest and belly, smooth to slightly areolate on throat; (4) tympanum small (1/2–2/5 of ED), welldefined, slightly higher than long; tympanic membrane differentiated; tympanic annulus prominent anteroventrally, posterodorsally concealed beneath skin; (5) snout acuminate in dorsal view, rounded in profile; (6) 
<emphasis id="26E9B038C009FF9EFAA5F98A41A04C2D" italics="true" pageId="2">canthus rostralis</emphasis>
weakly concave in dorsal view, nearly angular in cross-section; (7) upper eyelid with one large and few low tubercles; (8) choanae large, round, not concealed by palatal shelf of maxillary arch; (9) dentigerous processes of vomers low, triangular, slightly smaller than choanae, arranged in V-shaped, posterior and medial to choanae, each bearing one to four teeth; (10) tongue oval, longer than wide, posterior 1/4 free, posterior edge feebly notched; (11) males with prominent vocal slits; (12) males without nuptial pads on thumbs; (13) FI shorter than FII; (14) preaxial keels only on FII and FIII, very weak and barely visible; (15) two to four large ulnar tubercles longitudinally aligned on the outer ventrolateral surface of forearm; (16) finger disc and pad absent on FI, pads present and disc slightly expanded on FII to FIV; (17) calcar tubercles absent; heel with few, low tubercles; (18) four to five fold-like tarsal tubercles aligned on the outer ventrolateral surface of tarsus; (19) inner tarsal fold short, low and straight, extending on the distal fourth of the tarsus; (20) outer metatarsal tubercle large, round, swollen; inner metatarsal tubercle oval, elongate, about two times the size of the outer; (21) toes without lateral fringes or keels; (21) toes not webbed; (22) all toes with slightly expanded discs and digital pads; (23) TV slightly longer than TIII; (24) throat white reticulate of black, rest of ventral surfaces pale orange finely punctuated of black.
</paragraph>
<paragraph id="14226C2AC008FF9FFF27FF60435F4D58" blockId="3.[151,1437,151,2013]" pageId="3">
<emphasis id="26E9B038C008FF9FFF27FF6043024AA4" bold="true" box="[199,599,223,248]" pageId="3">Comparison with similar species.</emphasis>
<taxonomicName id="D39D17A9C008FF9FFD80FF5F42174AAB" box="[608,834,224,247]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFD80FF5F42174AAB" box="[608,834,224,247]" italics="true" pageId="3">Pristimantis munozi</emphasis>
</taxonomicName>
has a small but well-defined tympanum and slightly expanded discs on FII to IV, these traits differentiate 
<taxonomicName id="D39D17A9C008FF9FFD0EFEBA420E4B40" box="[750,859,261,284]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFD0EFEBA420E4B40" box="[750,859,261,284]" italics="true" pageId="3">P. munozi</emphasis>
</taxonomicName>
from 
<taxonomicName id="D39D17A9C008FF9FFC41FEBA45764B40" box="[929,1059,261,284]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="scopaeus">
<emphasis id="26E9B038C008FF9FFC41FEBA45764B40" box="[929,1059,261,284]" italics="true" pageId="3">P. scopaeus</emphasis>
</taxonomicName>
, which lacks tympanum and disc on FII-IV. In 
<taxonomicName id="D39D17A9C008FF9FFED2FE9640F54B63" box="[306,416,296,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFED2FE9640F54B63" box="[306,416,296,320]" italics="true" pageId="3">P. munozi</emphasis>
</taxonomicName>
the tips of finger and toe discs are rounded and present an inner metatarsal tubercle two times the size of the outer and distally not free, which differs notably from the acuminate tips of the discs and the larger (ca. four times the size of the outer) and distally free inner metatarsal tubercle of 
<taxonomicName id="D39D17A9C008FF9FFB86FECE459D4BD4" box="[1126,1224,368,392]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="hectus">
<emphasis id="26E9B038C008FF9FFB86FECE459D4BD4" box="[1126,1224,368,392]" italics="true" pageId="3">P. hectus</emphasis>
</taxonomicName>
, 
<taxonomicName id="D39D17A9C008FF9FFB36FECE44C04BD4" box="[1238,1429,368,392]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="lucidosignatus">
<emphasis id="26E9B038C008FF9FFB36FECE44C04BD4" box="[1238,1429,368,392]" italics="true" pageId="3">P. lucidosignatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="D39D17A9C008FF9FFF29FE2A407B4BF0" box="[201,302,405,428]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="onorei">
<emphasis id="26E9B038C008FF9FFF29FE2A407B4BF0" box="[201,302,405,428]" italics="true" pageId="3">P. onorei</emphasis>
</taxonomicName>
. Adult males of 
<taxonomicName id="D39D17A9C008FF9FFE14FE2A43374BF0" box="[500,610,405,428]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFE14FE2A43374BF0" box="[500,610,405,428]" italics="true" pageId="3">P. munozi</emphasis>
</taxonomicName>
have prominent vocal slits while there are absent in 
<taxonomicName id="D39D17A9C008FF9FFB28FE2A44194BF0" box="[1224,1356,404,428]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="xeniolum">
<emphasis id="26E9B038C008FF9FFB28FE2A44194BF0" box="[1224,1356,404,428]" italics="true" pageId="3">P. xeniolum</emphasis>
</taxonomicName>
and 
<taxonomicName id="D39D17A9C008FF9FFA67FE2A41BB4B8C" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="floridus">
<emphasis id="26E9B038C008FF9FFA67FE2A41BB4B8C" italics="true" pageId="3">P. floridus</emphasis>
</taxonomicName>
; additionally, the new species differs from 
<taxonomicName id="D39D17A9C008FF9FFD31FE0642064B8C" box="[721,851,440,464]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="xeniolum">
<emphasis id="26E9B038C008FF9FFD31FE0642064B8C" box="[721,851,440,464]" italics="true" pageId="3">P. xeniolum</emphasis>
</taxonomicName>
by having odontophores and vomerine teeth (absent in the latter), and from 
<taxonomicName id="D39D17A9C008FF9FFE4AFE6243754BA8" box="[426,544,476,500]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="floridus">
<emphasis id="26E9B038C008FF9FFE4AFE6243754BA8" box="[426,544,476,500]" italics="true" pageId="3">P. floridus</emphasis>
</taxonomicName>
by having prominent dorsolateral folds, and discs only slightly expanded on fingers and toes (
<emphasis id="26E9B038C008FF9FFE84FDBE40834844" box="[356,470,512,536]" italics="true" pageId="3">P. f l o r i d u s</emphasis>
lacks dorsolateral folds and presents wide discs, about twice the width of the last phalange). Prominent dorsolateral folds, weak preaxial keels on FII and III, and four to five fold-like tarsal tubercles aligned on the outer ventrolateral surface of tarsus distinguishes to 
<taxonomicName id="D39D17A9C008FF9FFC13FDF6450A4803" box="[1011,1119,584,608]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFC13FDF6450A4803" box="[1011,1119,584,608]" italics="true" pageId="3">P. munozi</emphasis>
</taxonomicName>
from 
<taxonomicName id="D39D17A9C008FF9FFB44FDF6445C483C" box="[1188,1289,585,608]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="repens">
<emphasis id="26E9B038C008FF9FFB44FDF6445C483C" box="[1188,1289,585,608]" italics="true" pageId="3">P. repens</emphasis>
</taxonomicName>
, which lacks dorsolateral folds the dorsum, keels on fingers and tubercles on the tarsus. 
<taxonomicName id="D39D17A9C008FF9FFBE5FDD245BC48D8" box="[1029,1257,621,644]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFBE5FDD245BC48D8" box="[1029,1257,621,644]" italics="true" pageId="3">Pristimantis munozi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="D39D17A9C008FF9FFA67FDD241A348F4" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="ocreatus">
<emphasis id="26E9B038C008FF9FFA67FDD241A348F4" italics="true" pageId="3">P. ocreatus</emphasis>
</taxonomicName>
by having odontophores on vomer, preaxial keels on FII–III, pads on FII–IV, fold-like tubercles on tarsus, and males with prominent vocal slits (
<taxonomicName id="D39D17A9C008FF9FFDB8FD0A43834890" box="[600,726,693,716]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="ocreatus">
<emphasis id="26E9B038C008FF9FFDB8FD0A43834890" box="[600,726,693,716]" italics="true" pageId="3">P. ocreatus</emphasis>
</taxonomicName>
lacking odontophores on vomers, keels and pads on fingers, tubercles on tarsus, and only with short vocal slits). From 
<taxonomicName id="D39D17A9C008FF9FFCB1FD66455748AC" box="[849,1026,728,752]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="pyrrhomerus">
<emphasis id="26E9B038C008FF9FFCB1FD66455748AC" box="[849,1026,728,752]" italics="true" pageId="3">P. pyrrhomerus</emphasis>
</taxonomicName>
, the new species differs in having prominent dorsolateral folds, preaxial keels on FII–III, weak discs and pads on FII–IV, fold-like tubercles on outer ventrolateral edge of tarsus, dentigerous process of vomers low, males with prominent vocal slits and without red color on groin and concealed parts of thighs (
<taxonomicName id="D39D17A9C008FF9FFD7EFCFA421E4900" box="[670,843,836,860]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="pyrrhomerus">
<emphasis id="26E9B038C008FF9FFD7EFCFA421E4900" box="[670,843,836,860]" italics="true" pageId="3">P. pyrrhomerus</emphasis>
</taxonomicName>
lacks dorsolateral folds and lateral keels on fingers, discs and pads absent on FI–II, outer tarsal tubercles small and sub-conical, prominent dentigerous process, and males whit short vocal slits and red colored groins and hidden parts of thighs). 
<taxonomicName id="D39D17A9C008FF9FFBF1FC3245A749F8" box="[1041,1266,909,932]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFBF1FC3245A749F8" box="[1041,1266,909,932]" italics="true" pageId="3">Pristimantis munozi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="D39D17A9C008FF9FFA67FC3241B7499B" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="myersi">
<emphasis id="26E9B038C008FF9FFA67FC3241B7499B" italics="true" pageId="3">P. myersi</emphasis>
</taxonomicName>
by having dentigerous process on vomers, white ventral color finely spotted of black, with translucent gray groins, and by lacking papilla at tip of snout, lateral keels on toes, and by have a large tubercle on the upper eyelid (
<taxonomicName id="D39D17A9C008FF9FFF40FC4640504E53" box="[160,261,1016,1040]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="myersi">
<emphasis id="26E9B038C008FF9FFF40FC4640504E53" box="[160,261,1016,1040]" italics="true" pageId="3">P. myersi</emphasis>
</taxonomicName>
presents a fleshy papilla at tip of snout, lateral keels on toes, upper eyelid with numerous small tubercles, black ventral color spotted with white, groins with red spots, and lacks dentigerous process of vomers). 
<taxonomicName id="D39D17A9C008FF9FFF77FBFF40224E0B" box="[151,375,1088,1111]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFF77FBFF40224E0B" box="[151,375,1088,1111]" italics="true" pageId="3">Pristimantis munozi</emphasis>
</taxonomicName>
distinguished from 
<emphasis id="26E9B038C008FF9FFDB7FBFE43E54E04" box="[599,688,1088,1112]" italics="true" pageId="3">P. f e s t a e</emphasis>
by having the dorsal skin shagreen with middorsal raphe, scapular and dorsolateral folds, small tympanum, snout acuminate in dorsal view, 
<emphasis id="26E9B038C008FF9FFC39FBDB45654E20" box="[985,1072,1124,1148]" italics="true" pageId="3">canthus</emphasis>
nearly angular in cross-section, enlarged tubercle on the upper eyelid, pads on FII–IV, white venter finely spotted with black, and lacking of lateral keels on toes (
<emphasis id="26E9B038C008FF9FFED6FB1240DA4E98" box="[310,399,1196,1220]" italics="true" pageId="3">P. f e s t a e</emphasis>
have dorsal skin smooth without middorsal raphe nor dorsolateral folds, present a prominent tympanum, snout rounded in dorsal view, 
<emphasis id="26E9B038C008FF9FFD8AFB6F43944EB4" box="[618,705,1232,1256]" italics="true" pageId="3">canthus</emphasis>
rounded, upper eyelid without enlarged tubercles, fingers lacking pads, toes whit lateral keels, and black venter spotted with white and pale cream). 
<taxonomicName id="D39D17A9C008FF9FFBB0FB4A441F4F50" box="[1104,1354,1268,1292]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="gladiator">
<emphasis id="26E9B038C008FF9FFBB0FB4A441F4F50" box="[1104,1354,1268,1292]" italics="true" pageId="3">Pristimantis gladiator</emphasis>
</taxonomicName>
differs from the new species by lacking middorsal raphe and dorsolateral folds, and having only small tubercles on the upper eyelid and a prominent tympanum; additionally, 
<taxonomicName id="D39D17A9C008FF9FFCE4FA8242DD4F08" box="[772,904,1340,1364]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="gladiator">
<emphasis id="26E9B038C008FF9FFCE4FA8242DD4F08" box="[772,904,1340,1364]" italics="true" pageId="3">P. gladiator</emphasis>
</taxonomicName>
shows slightly pointed and asymmetrical discs (discs with tip rounded in 
<taxonomicName id="D39D17A9C008FF9FFE2FFADE436A4F2B" box="[463,575,1376,1400]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C008FF9FFE2FFADE436A4F2B" box="[463,575,1376,1400]" italics="true" pageId="3">P. munozi</emphasis>
</taxonomicName>
). From 
<taxonomicName id="D39D17A9C008FF9FFD42FADE42754F24" box="[674,800,1376,1400]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="bicantus">
<emphasis id="26E9B038C008FF9FFD42FADE42754F24" box="[674,800,1376,1400]" italics="true" pageId="3">P. bicantus</emphasis>
</taxonomicName>
the new species is distinguished by having prominent dorsolateral folds, an enlarged tubercle on the upper eyelid, snout acuminate in dorsal view, prolateral keels on FII– III, ulnar tubercles and outer tarsal tubercles present (
<taxonomicName id="D39D17A9C008FF9FFD1BFA1642224F9C" box="[763,887,1448,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="bicantus">
<emphasis id="26E9B038C008FF9FFD1BFA1642224F9C" box="[763,887,1448,1472]" italics="true" pageId="3">P. bicantus</emphasis>
</taxonomicName>
usually lacks dorsolateral folds, lateral keels on fingers, ulnar and outer tarsal tubercles, and has small tubercles on the upper eyelid, and rounded snout in dorsal view). At the 
<typeStatus id="CB26D288C008FF9FFED4FA4F40314C54" box="[308,356,1520,1544]" pageId="3">type</typeStatus>
locality, 
<emphasis id="26E9B038C008FF9FFE2EFA4E436D4C5B" box="[462,568,1520,1544]" italics="true" pageId="3">P. m u n o z i</emphasis>
is sympatric with 
<taxonomicName id="D39D17A9C008FF9FFCEFFA4E42314C54" box="[783,868,1520,1544]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="leoni">
<emphasis id="26E9B038C008FF9FFCEFFA4E42314C54" box="[783,868,1520,1544]" italics="true" pageId="3">P. leoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D39D17A9C008FF9FFC7DFA4E45424C54" box="[925,1047,1520,1544]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="sirnigeli">
<emphasis id="26E9B038C008FF9FFC7DFA4E45424C54" box="[925,1047,1520,1544]" italics="true" pageId="3">P. sirnigeli</emphasis>
</taxonomicName>
; however, it is distinguished from the former by having one large and rounded tubercle on the upper eyelid, preaxial keels on FII–III, toes without lateral keels, tips of the discs on toes rounded, and fold-like outer tarsal tubercles (
<taxonomicName id="D39D17A9C008FF9FFBD3F98645D24C0C" box="[1075,1159,1592,1616]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="leoni">
<emphasis id="26E9B038C008FF9FFBD3F98645D24C0C" box="[1075,1159,1592,1616]" italics="true" pageId="3">P. leoni</emphasis>
</taxonomicName>
with numerous enlarged tubercles on the upper eyelid, lateral keels absent on the fingers but present on the toes, tips of the discs on toes slightly pointed, and outer tarsal tubercles small and conical). Finally, 
<emphasis id="26E9B038C008FF9FFC55F93E454A4CCB" box="[949,1055,1664,1688]" italics="true" pageId="3">P. m u n o z i</emphasis>
is distinguished from 
<taxonomicName id="D39D17A9C008FF9FFAC2F93E44CE4CC4" box="[1314,1435,1664,1688]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="sirnigeli">
<emphasis id="26E9B038C008FF9FFAC2F93E44CE4CC4" box="[1314,1435,1664,1688]" italics="true" pageId="3">P. sirnigeli</emphasis>
</taxonomicName>
by having a smaller body size in adult males (SVL = 
<quantity id="D365C1CFC008FF9FFD0DF91B42C14CE0" box="[749,916,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" metricValueMax="1.7899999999999998" metricValueMin="1.49" pageId="3" unit="mm" value="16.4" valueMax="17.9" valueMin="14.9">14.9–17.9 mm</quantity>
), prominent dorsolateral folds, relatively short fingers, and by lacking lateral keels on toes (
<taxonomicName id="D39D17A9C008FF9FFD71F976425F4CBC" box="[657,778,1736,1760]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" phylum="Chordata" rank="species" species="sirnigeli">
<emphasis id="26E9B038C008FF9FFD71F976425F4CBC" box="[657,778,1736,1760]" italics="true" pageId="3">P. sirnigeli</emphasis>
</taxonomicName>
with lateral keels on toes, relatively long and thin fingers, and male SVL = 
<quantity id="D365C1CFC008FF9FFEB6F95340A84D58" box="[342,509,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" metricValueMax="2.06" metricValueMin="1.86" pageId="3" unit="mm" value="19.6" valueMax="20.6" valueMin="18.6">18.6–20.6 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="5C873FA1C008FF94FF27F8B043DD4CBF" lastPageId="8" pageId="3" type="description">
<paragraph id="14226C2AC008FF98FF27F8B040144B8C" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,464]" lastPageId="4" pageId="3">
<emphasis id="26E9B038C008FF9FFF27F8B043434D74" bold="true" box="[199,534,1807,1832]" pageId="3">
Description of the 
<typeStatus id="CB26D288C008FF9FFE49F8B043474D74" box="[425,530,1807,1832]" pageId="3" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male of 
<quantity id="D365C1CFC008FF9FFD27F8B042784D7B" box="[711,813,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="3" unit="mm" value="17.6">17.6 mm</quantity>
SVL. Body short and robust (
<figureCitation id="8CA670AFC008FF9FFB69F8B045804D74" box="[1161,1237,1807,1832]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1822]" captionTargetBox="[151,1436,517,1778]" captionTargetId="figure@4.[151,1436,517,1778]" captionTargetPageId="4" captionText="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="3">Figs. 1</figureCitation>
a–b, 2a–b). Head slightly narrower than body, slightly longer than wide (
<figureCitation id="8CA670AFC008FF9FFCE3F88B42054D11" box="[771,848,1844,1869]" captionStart="FIGURE 2" captionStartId="5.[151,250,1524,1546]" captionTargetBox="[151,1436,301,1502]" captionTargetId="figure@5.[151,1436,301,1503]" captionTargetPageId="5" captionText="FIGURE 2. Pristimantis munozi sp. nov. Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante." httpUri="https://zenodo.org/record/225617/files/figure.png" pageId="3">Figs. 2</figureCitation>
a–b, 3a–b); HeL 39.5% of SVL; greatest head width between angles of jaws 38% of SVL; cranial crests absent. Snout short, acuminate in dorsal view (
<figureCitation id="8CA670AFC008FF9FFAE9F8E844184D2C" box="[1289,1357,1879,1904]" captionStart="FIGURE 3" captionStartId="6.[151,250,1701,1723]" captionTargetBox="[151,1436,481,1677]" captionTargetId="figure@6.[151,1436,481,1679]" captionTargetPageId="6" captionText="FIGURE 3. Pristimantis munozi sp. nov. (holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a, b, e) &amp; F. J. M. Rojas- Runjaic (c, d)." httpUri="https://zenodo.org/record/225618/files/figure.png" pageId="3">Fig. 3</figureCitation>
a), and rounded in profile (
<figureCitation id="8CA670AFC008FF9FFE90F8C340E74DC8" box="[368,434,1916,1941]" captionStart="FIGURE 3" captionStartId="6.[151,250,1701,1723]" captionTargetBox="[151,1436,481,1677]" captionTargetId="figure@6.[151,1436,481,1679]" captionTargetPageId="6" captionText="FIGURE 3. Pristimantis munozi sp. nov. (holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a, b, e) &amp; F. J. M. Rojas- Runjaic (c, d)." httpUri="https://zenodo.org/record/225618/files/figure.png" pageId="3">Fig. 3</figureCitation>
c); larger than eye (ETS/ED = 1.5); distance eye-nostril slightly shorter than eye diameter (EN/ED = 0.8). 
<emphasis id="26E9B038C008FF9FFEB4F81F434C4DE4" box="[340,537,1952,1976]" italics="true" pageId="3">Canthus rostralis</emphasis>
weakly concave in dorsal view; nearly angular in cross-section; loreal region slightly concave, sloping outward to lip. Not protruding lips. Nares not protuberant, with dorsolateral orientation, barely visible from the front and completely observable in dorsal view. Upper eyelid wide, narrower than interocular region (UEW/IO = 0.8), with one enlarged and rounded tubercle on each eyelid and few additional low tubercles. Tympanum small (TD/ED = 0.4) but well-defined, slightly higher than long; tympanic membrane differentiated; tympanic annulus prominent anteroventrally, posterodorsally concealed beneath skin, barely visible in dorsal view. Supratympanic fold absent, with several large and conical postrictal tubercles below tympanum. Choanae large, round, not concealed by palatal shelf of maxillary arch; dentigerous processes of vomers low, triangular, slightly smaller than choanae, arranged in V-shape, posterior and medial to choanae, each bearing threeto-four teeth. Tongue oval, longer than wide, posterior 1/4 free, posterior edge feebly notched. Vocal slits present and prominent.
</paragraph>
<caption id="40E23CA2C00FFF98FF77F8B7445B4D60" httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="4" targetBox="[151,1436,517,1778]" targetPageId="4">
<paragraph id="14226C2AC00FFF98FF77F8B7445B4D60" blockId="4.[151,1436,1800,1852]" pageId="4">
<emphasis id="26E9B038C00FFF98FF77F8B740414D41" bold="true" box="[151,276,1800,1822]" pageId="4">FIGURE 1.</emphasis>
<taxonomicName id="D39D17A9C00FFF98FEFBF8B640B24D42" box="[283,487,1801,1822]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" phylum="Chordata" rank="species" species="munozi" status="sp. nov.">
<emphasis id="26E9B038C00FFF98FEFBF8B640B24D42" box="[283,487,1801,1822]" italics="true" pageId="4">Pristimantis munozi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DDA0D43C00FFF98FE0FF8B743144D41" box="[495,577,1800,1821]" pageId="4" rank="species">
<emphasis id="26E9B038C00FFF98FE0FF8B743144D41" bold="true" box="[495,577,1800,1821]" pageId="4">sp. nov.</emphasis>
</taxonomicNameLabel>
Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F.J.M. Rojas-Runjaic.
</paragraph>
</caption>
<paragraph id="14226C2AC00FFF99FF27F8DB43574AA4" blockId="4.[151,1436,1892,2025]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" pageId="4">
Skin of dorsum and upper eyelids shagreen; middorsal raphe present, but low and barely distinguishable; scapular folds present, sinusoidal and prominent, ending posteriorly in a large conical tubercle; dorsolateral folds straight and prominent, extending from posterior end of scapular folds to level of groins; no paravertebral folds (
<figureCitation id="8CA670AFC00FFF98FF7FF86F41B04DB4" box="[159,229,2000,2025]" captionStart="FIGURE 2" captionStartId="5.[151,250,1524,1546]" captionTargetBox="[151,1436,301,1502]" captionTargetId="figure@5.[151,1436,301,1503]" captionTargetPageId="5" captionText="FIGURE 2. Pristimantis munozi sp. nov. Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante." httpUri="https://zenodo.org/record/225617/files/figure.png" pageId="4">Fig. 2</figureCitation>
a); cloacal sheath absent, small cloacal tubercles present but not conspicuous; flanks and posterior third of dorsum slightly tuberculate; dorsal surface of limbs finely tuberculate. Throat smooth, chest and belly areolate; ventral surface of limbs smooth to slightly areolate (
<figureCitation id="8CA670AFC00EFF99FCECFF0342054A89" box="[780,848,188,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1822]" captionTargetBox="[151,1436,517,1778]" captionTargetId="figure@4.[151,1436,517,1778]" captionTargetPageId="4" captionText="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="5">Fig. 1</figureCitation>
b). Two-to-three large ulnar tubercles along outer ventrolateral surface of forearm.
</paragraph>
<caption id="40E23CA2C00EFF99FF77FA4B40434C14" httpUri="https://zenodo.org/record/225617/files/figure.png" pageId="5" targetBox="[151,1436,301,1502]" targetPageId="5">
<paragraph id="14226C2AC00EFF99FF77FA4B40434C14" blockId="5.[151,1437,1524,1608]" pageId="5">
<emphasis id="26E9B038C00EFF99FF77FA4B40474C55" bold="true" box="[151,274,1524,1546]" pageId="5">FIGURE 2.</emphasis>
<taxonomicName id="D39D17A9C00EFF99FEF8FA4A40B64C56" box="[280,483,1525,1546]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" phylum="Chordata" rank="species" species="munozi" status="sp. nov.">
<emphasis id="26E9B038C00EFF99FEF8FA4A40B64C56" box="[280,483,1525,1546]" italics="true" pageId="5">Pristimantis munozi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DDA0D43C00EFF99FE09FA4A436C4C56" box="[489,569,1525,1546]" pageId="5" rank="species">
<emphasis id="26E9B038C00EFF99FE09FA4A436C4C56" bold="true" box="[489,569,1525,1546]" pageId="5">sp. nov.</emphasis>
</taxonomicNameLabel>
Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante.
</paragraph>
</caption>
<paragraph id="14226C2AC00EFF99FF27F9D040554DF4" blockId="5.[151,1436,1647,2032]" pageId="5">
Hand length 28.6% of SVL. Relative lengths of adpressed fingers III&gt; IV&gt; II&gt; I; adpressed FI does not reach proximal edge of disc pad of FII; nuptial pads absent; adpresed FIV reaches past distal subarticular tubercle of FIII. Preaxial keels only present on FII and III, very weak and barely visible. Finger disc absent on FI, slightly expanded on FII to IV; digital pad absent on FI, present in all other fingers, rounded, almost longer than wide; circumferential groove of digital pads poorly defined laterally and incomplete proximally; distal edge of discs rounded; disc of FIII about 1.4 times wider than distal end of adjacent phalanx. Subarticular tubercles large, longer than wide, and slightly swollen; palmar tubercle large, slightly swollen, deeply bifid, V-shaped; thenar tubercle large, slightly swollen, elongate; supernumerary tubercles present, relative large (between 1/3–1/2 size of subarticular tubercles) (
<figureCitation id="8CA670AFC00EFF99FF7FF83041B74DF4" box="[159,226,1935,1960]" captionStart="FIGURE 3" captionStartId="6.[151,250,1701,1723]" captionTargetBox="[151,1436,481,1677]" captionTargetId="figure@6.[151,1436,481,1679]" captionTargetPageId="6" captionText="FIGURE 3. Pristimantis munozi sp. nov. (holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a, b, e) &amp; F. J. M. Rojas- Runjaic (c, d)." httpUri="https://zenodo.org/record/225618/files/figure.png" pageId="5">Fig. 3</figureCitation>
d).
</paragraph>
<paragraph id="14226C2AC00EFF9AFF27F80C42954BF0" blockId="5.[151,1436,1647,2032]" lastBlockId="6.[151,1436,151,429]" lastPageId="6" pageId="5">
<collectingCountry id="6C8A2CBAC00EFF99FF27F80C41AA4D90" box="[199,255,1971,1996]" name="India" pageId="5">Hind</collectingCountry>
limbs relatively long (thigh length 50.5% SVL; shank length 56.3% SVL; foot length 50.5% SVL); heels overlap when shanks are held perpendicular to sagittal plane; relative length of adpressed toes IV&gt; V&gt; III&gt; II&gt; I; several small, low, and rounded tubercles on heel, calcar tubercle absent; four to five fold-like tarsal tubercles aligned on the outer ventrolateral surface of tarsus; inner tarsal fold short, low and straight, extending on the distal fourth of the tarsus. Tip of TV barely extends past distal edge of mid subarticular tubercle of TIV; tip of TIII reaches distal half of mid subarticular tubercle of TIV. Disc of TIV wider than disc of FIII (F3D/T4D = 0.8); all toes with slightly expanded discs and digital pads; circumferential grooves of discs absent. Toes without lateral fringes or keels; webbing absent. Outer metatarsal tubercle large, round, swollen; inner metatarsal tubercle oval, about two times the size of outer metatarsal tubercle. Supernumerary plantar tubercles scarce, small, low, and round. Subarticular tubercles large, round and slightly swollen (
<figureCitation id="8CA670AFC00DFF9AFC81FE2B42F64BF0" box="[865,931,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1701,1723]" captionTargetBox="[151,1436,481,1677]" captionTargetId="figure@6.[151,1436,481,1679]" captionTargetPageId="6" captionText="FIGURE 3. Pristimantis munozi sp. nov. (holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a, b, e) &amp; F. J. M. Rojas- Runjaic (c, d)." httpUri="https://zenodo.org/record/225618/files/figure.png" pageId="6">Fig. 3</figureCitation>
e).
</paragraph>
<caption id="40E23CA2C00DFF9AFF77F91A40774CA4" httpUri="https://zenodo.org/record/225618/files/figure.png" pageId="6" targetBox="[151,1436,481,1677]" targetPageId="6">
<paragraph id="14226C2AC00DFF9AFF77F91A40774CA4" blockId="6.[151,1435,1700,1784]" pageId="6">
<emphasis id="26E9B038C00DFF9AFF77F91A40414CE6" bold="true" box="[151,276,1701,1723]" pageId="6">FIGURE 3.</emphasis>
<taxonomicName id="D39D17A9C00DFF9AFEFCF91940BD4CE7" box="[284,488,1702,1723]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" phylum="Chordata" rank="species" species="munozi" status="sp. nov.">
<emphasis id="26E9B038C00DFF9AFEFCF91940BD4CE7" box="[284,488,1702,1723]" italics="true" pageId="6">Pristimantis munozi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DDA0D43C00DFF9AFE10F91A43144CE6" box="[496,577,1701,1722]" pageId="6" rank="species">
<emphasis id="26E9B038C00DFF9AFE10F91A43144CE6" bold="true" box="[496,577,1701,1722]" pageId="6">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, MZUTI 1782, male). Dorsal (a), ventral (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 1 mm. Photos: L. Bustamante (a,b,e) &amp; F.J.M. Rojas- Runjaic (c,d).
</paragraph>
</caption>
<paragraph id="14226C2AC00DFF9BFF27F89B4282483C" blockId="6.[151,1436,1828,2033]" lastBlockId="7.[151,1436,151,2013]" lastPageId="7" pageId="6">
<emphasis id="26E9B038C00DFF9AFF27F89B437A4D61" bold="true" box="[199,559,1828,1853]" pageId="6">
Color of the 
<typeStatus id="CB26D288C00DFF9AFE91F89B408D4D61" box="[369,472,1828,1853]" pageId="6" type="holotype">holotype</typeStatus>
in life
</emphasis>
(based on field notes and color digital photographs; 
<figureCitation id="8CA670AFC00DFF9AFB58F89B44524D61" box="[1208,1287,1828,1853]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1822]" captionTargetBox="[151,1436,517,1778]" captionTargetId="figure@4.[151,1436,517,1778]" captionTargetPageId="4" captionText="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="6">Figs. 1</figureCitation>
a–b). Dorsal background color grayish brown, with small and irregular black spots scattered on the dorsal surface of head, upper eyelids, and dorsum; several conspicuous black spots flanking externally the scapular fold. All tubercles and folds on dorsum of head and body with reddish brown coloration. 
<emphasis id="26E9B038C00DFF9AFC9FF82E451D4DF5" box="[895,1096,1937,1961]" italics="true" pageId="6">Canthus rostralis</emphasis>
with a black canthal stripe, extending from tip of snout to anterior corner of eye. Lips with vertical black and brown bars, except for cream bar just below eye. A conspicuous dark supratympanic stripe extends from upper and posterior parts of the tympanum and fails to reach the shoulder; the inferior border of this stripe is also delineated of creamish white. Flank grayish brown, slightly lighter than dorsum, with scattered small black spots and with one or two diffuse diagonal cream stripes. Groin background translucent gray, with several small immaculate white dots. Upper arm light brown; forearm grayish brown with two-to-three irregular blackish brown cross-bands in the distal half, more intensely pigmented the proximal one. Hand and fingers light brown with three-to-four ill-defined pale brown cross-bands; tips of finger discs reddish brown. 
<collectingCountry id="6C8A2CBAC00CFF9BFDC4FEF343094B39" box="[548,604,332,357]" name="India" pageId="7">Hind</collectingCountry>
limbs background grayish brown, becoming paler distally (on feet), with three to four blackish brown cross-bands on thighs, shanks, tarsi and feet; blackish cross-bands on thighs laterally bordered with light lines; background color of the inner surface of thighs translucent gray, densely spotted of black and with scattered small white dots; transversal rows of tubercles on shanks pigmented of reddish brown. Toes pale orange; toe discs dorsally grayish, bordered by reddish brown coloration. Throat white, finely reticulated with black; chest, belly, undersurface forelimbs of thighs and shanks pale orange, finely punctuated of black; tarsi ventrally grayish brown; palms and soles pale orange; subarticular tubercles of feet bright orange. Iris golden with fine black venation and a broad horizontal copper band; pupil ring golden.
</paragraph>
<paragraph id="14226C2AC00CFF9BFF27FDD343764901" blockId="7.[151,1436,151,2013]" pageId="7">
<emphasis id="26E9B038C00CFF9BFF27FDD3432C48D9" bold="true" box="[199,633,620,645]" pageId="7">
Color of the 
<typeStatus id="CB26D288C00CFF9BFEBCFDD3409648D9" box="[348,451,620,645]" pageId="7" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
(after sixteen months, 
<date id="60234AEAC00CFF9BFC99FDD242BB48D8" box="[889,1006,620,644]" pageId="7" value="2013-06">June 2013</date>
; 
<figureCitation id="8CA670AFC00CFF9BFC19FDD3451048D9" box="[1017,1093,620,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1524,1546]" captionTargetBox="[151,1436,301,1502]" captionTargetId="figure@5.[151,1436,301,1503]" captionTargetPageId="5" captionText="FIGURE 2. Pristimantis munozi sp. nov. Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante." httpUri="https://zenodo.org/record/225617/files/figure.png" pageId="7">Figs. 2</figureCitation>
a–b, 3a–e). Dorsal background almost uniformly gray; reddish brown coloration of dorsal tubercles and folds is replaced by gray and light gray; dorsal and lateral black spots turns more dark and conspicuous. Flanks turn grayish, lighter towards lower portion; diagonal light bands of flanks disappear. Groins and ventral surfaces turn white, with black reticulation. Palms and soles white, with diffuse pink tone and several small black blotches. Fingers and toes with dorsal dark gray, transversal bars, and ventrally colored of white, densely spotted of black. Iris grayish, finely punctuated with black; horizontal copper band turns black.
</paragraph>
<paragraph id="14226C2AC00CFF9BFF27FCD8440349F8" blockId="7.[151,1436,151,2013]" pageId="7">
<emphasis id="26E9B038C00CFF9BFF27FCD8437B49DC" bold="true" box="[199,558,871,896]" pageId="7">
Measurements of the 
<typeStatus id="CB26D288C00CFF9BFE27FCD8437B49DC" box="[455,558,871,896]" pageId="7" type="holotype">holotype</typeStatus>
</emphasis>
(in mm). SVL: 17.6; ThL: 8.9; SL: 9.9; FL: 8.9; HaL: 5.0; HeL: 6.9; HW: 6.7; 
<collectingCountry id="6C8A2CBAC00CFF9BFF77FC32419249F8" box="[151,199,909,932]" name="India" pageId="7">InD</collectingCountry>
: 1.9; IOD: 2.1; UEW: 1.7; EN: 1.7; ED: 2.2; ETS: 3.1; TD: 0.9; F3D: 0.6; T4D: 0.8; F1L: 2.6; F2L: 3.3.
</paragraph>
<paragraph id="14226C2AC00CFF9BFF27FC1044C94E69" blockId="7.[151,1436,151,2013]" pageId="7">
<emphasis id="26E9B038C00CFF9BFF27FC10406A4994" bold="true" box="[199,319,943,968]" pageId="7">Variation.</emphasis>
Females of 
<taxonomicName id="D39D17A9C00CFF9BFE3AFC0F4394499B" box="[474,705,944,967]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C00CFF9BFE3AFC0F4394499B" box="[474,705,944,967]" italics="true" pageId="7">Pristimantis munozi</emphasis>
</taxonomicName>
are unknown; then, sexual dimorphism is not evaluated. We mention, however, that females of all other species of 
<taxonomicName id="D39D17A9C00CFF9BFD19FC6A428149B0" box="[761,980,981,1004]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" phylum="Chordata" rank="species" species="myersi">
<emphasis id="26E9B038C00CFF9BFD19FC6A428149B0" box="[761,980,981,1004]" italics="true" pageId="7">Pristimantis myersi</emphasis>
</taxonomicName>
group are larger than males, and several of them have red blotches on the groins (gender-specific coloration). Thus, we suppose that adult females of 
<taxonomicName id="D39D17A9C00CFF9BFA67FC4641BD4E68" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C00CFF9BFA67FC4641BD4E68" italics="true" pageId="7">P. munozi</emphasis>
</taxonomicName>
may be are larger than males and may have red flash coloration on the groins, as other species of the group.
</paragraph>
<paragraph id="14226C2AC00CFF9BFF27FB8043CC4FC0" blockId="7.[151,1436,151,2013]" pageId="7">
Adult males range from 14.9–19.7 (16.7 ±1.6; 
<emphasis id="26E9B038C00CFF9BFD36FBFE43B14E04" box="[726,740,1089,1112]" italics="true" pageId="7">n</emphasis>
= 8). Variation of morphometric characters is shown in 
<tableCitation id="591F5991C00CFF9BFABEFB8041F34E20" captionStart="TABLE 1" captionStartId="8.[151,239,476,499]" captionTargetBox="[161,1115,558,1336]" captionTargetPageId="8" captionText="TABLE 1. Morphometric variation in adult males (n = 8) of Pristimantis munozi. X? ± SD: mean ± standard deviation; Min – Max: minimum – maximum." httpUri="http://table.plazi.org/id/40E23CA2C003FF94FF77FE6340AC484F" pageId="7" tableUuid="40E23CA2C003FF94FF77FE6340AC484F">Table 1</tableCitation>
. Head length ranges from 39.1%–42.3% SVL (40.6% ± 1.1; 
<emphasis id="26E9B038C00CFF9BFC92FBDA42D54E20" box="[882,896,1125,1148]" italics="true" pageId="7">n</emphasis>
= 8), and maximum head width from 35.4%– 42.6% SVL (38.3% ± 2.2; 
<emphasis id="26E9B038C00CFF9BFE21FB36409A4EFC" box="[449,463,1161,1184]" italics="true" pageId="7">n</emphasis>
= 8). Head can be as long as wide, or slightly longer than wide (HeL/HW: 1.0 –1.1[1.1 ± 0.1; 
<emphasis id="26E9B038C00CFF9BFF04FB1241A74E98" box="[228,242,1197,1220]" italics="true" pageId="7">n</emphasis>
= 8]). The EN/ED ratio is somewhat variable but every eye-naris distance is smaller than eye diameter (0.7–0.9 [0.8 ± 0.1; 
<emphasis id="26E9B038C00CFF9BFE9DFB6E40DE4EB4" box="[381,395,1233,1256]" italics="true" pageId="7">n</emphasis>
= 8]). The width of the upper eyelid is narrower than the interorbital distance, and its ratio ranges from 0.7–0.8 (0.78 ± 0.1; 
<emphasis id="26E9B038C00CFF9BFDFEFB4A43794F50" box="[542,556,1269,1292]" italics="true" pageId="7">n</emphasis>
= 8). The snout length also is slightly variable but always longer than eye diameter (1.2–1.6 [1.4 ± 0.1; 
<emphasis id="26E9B038C00CFF9BFE16FAA643514F6C" box="[502,516,1305,1328]" italics="true" pageId="7">n</emphasis>
= 8]). Hand length ranges from 26.4–29.7% SVL (28.2% ± 1.2; 
<emphasis id="26E9B038C00CFF9BFAF1FAA6444A4F6C" box="[1297,1311,1305,1328]" italics="true" pageId="7">n</emphasis>
= 8). The tympanum is small and its ratio respect to eye has little variation (TD/ED: 0.4–0.5; 
<emphasis id="26E9B038C00CFF9BFBBFFA8245384F08" box="[1119,1133,1341,1364]" italics="true" pageId="7">n</emphasis>
= 8). Thigh length ranges from 49.3%–55.0% SVL (51.9% ± 2.2; 
<emphasis id="26E9B038C00CFF9BFDB9FADE43324F24" box="[601,615,1377,1400]" italics="true" pageId="7">n</emphasis>
= 8); shank length from 53.6–62.1% SVL (58.0% ± 2.8; 
<emphasis id="26E9B038C00CFF9BFB0CFADE45AF4F24" box="[1260,1274,1377,1400]" italics="true" pageId="7">n</emphasis>
= 8), and foot length from 48.5%–54.4% (52.1 ± 2.0; 
<emphasis id="26E9B038C00CFF9BFDB0FA3A430B4FC0" box="[592,606,1413,1436]" italics="true" pageId="7">n</emphasis>
= 8).
</paragraph>
<paragraph id="14226C2AC00CFF9BFF27FA16425D4D74" blockId="7.[151,1436,151,2013]" pageId="7">
In almost all specimens dorsal skin texture is shagreen with scattered tubercles of variable size, except MZUTI 1781, which have dorsal skin nearly smooth, non-tubercular, and with scattered granules on flanks and posterior third of dorsum. Flanks and posterior third of dorsum densely tubercular in MZUTI 1777. Upper eyelid typically with one tubercle; MZUTI 1777 has two tubercles on each upper eyelid. Middorsal raphe absent in MZUTI 1778, but prominent in MZUTI1779. Scapular folds extending anteriorly and reaching tubercles of upper eyelids in MZUTI 1783, nearly straight and continuous with dorsolateral folds in MZUTI 
<date id="60234AEAC00CFF9BFBFCF9E345934C28" box="[1052,1222,1628,1653]" pageId="7" value="1779" valueMax="1781">1779 and 1781</date>
. Dorsolateral folds extending to sacral region (MZUTI 
<date id="60234AEAC00CFF9BFDD2F93F43E14CC4" box="[562,692,1664,1688]" pageId="7" value="1777" valueMax="1778">1777–1778</date>
, 1784), to groins (MZUTI 
<date id="60234AEAC00CFF9BFC0AF93F453F4CC4" box="[1002,1130,1664,1688]" pageId="7" value="1780" valueMax="1781">1780–1781</date>
) or inclusively converging above the vent level (MZUTI 1779, 1783). Ulnar tubercles on forearms range from two-to-four; tarsal tubercles in number of four or five. Throat typically smooth, but slightly areolate in some specimens (MZUTI 1777, 1780, 1784). Number of vomerine teeth somewhat variable, from one or two (MZUTI 
<date id="60234AEAC00CFF9BFBADF95345994D58" box="[1101,1228,1772,1796]" pageId="7" value="1777" valueMax="1778">1777–1778</date>
) to four (MZUTI 1783), but typically three-to-four (MZUTI 
<date id="60234AEAC00CFF9BFD99F8AF43A94D74" box="[633,764,1808,1832]" pageId="7" value="1779" valueMax="1782">1779–1782</date>
).
</paragraph>
<paragraph id="14226C2AC00CFF94FF27F88A41964BF1" blockId="7.[151,1436,151,2013]" lastBlockId="8.[151,1436,151,429]" lastPageId="8" pageId="7">
In almost all specimens examined, the color pattern is similar to that of the 
<typeStatus id="CB26D288C00CFF9BFBA9F88B45FE4D11" box="[1097,1195,1844,1869]" pageId="7" type="holotype">holotype</typeStatus>
(
<figureCitation id="8CA670AFC00CFF9BFB5EF88B445E4D11" box="[1214,1291,1844,1869]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1822]" captionTargetBox="[151,1436,517,1778]" captionTargetId="figure@4.[151,1436,517,1778]" captionTargetPageId="4" captionText="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="7">Figs. 1</figureCitation>
a–b), except specimens MZUTI 
<date id="60234AEAC00CFF9BFE96F8E743744D2C" box="[374,545,1879,1904]" pageId="7" value="1779" valueMax="1781">1779 and 1781</date>
(
<figureCitation id="8CA670AFC00CFF9BFDD3F8E8432B4D2C" box="[563,638,1879,1904]" captionStart="FIGURE 1" captionStartId="4.[151,250,1800,1822]" captionTargetBox="[151,1436,517,1778]" captionTargetId="figure@4.[151,1436,517,1778]" captionTargetPageId="4" captionText="FIGURE 1. Pristimantis munozi sp. nov. Dorsolateral (a) and ventral (b) views of the holotype (MZUTI 1782, male) in life. Dorsolateral (c) and ventral (d) views of the paratype (MZUTI 1781, male) in life. Photos: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225616/files/figure.png" pageId="7">Figs. 1</figureCitation>
c–d, 2c–d), which show striking differences. These specimens have a dorsal background pale ocher (pale gray in preservative), extended from the tip of snout to the vent, and medially to the scapular and dorsolateral folds; this area is bordered by a sinuous black stripe that runs from the tip of snout, through nares, 
<emphasis id="26E9B038C00CFF9BFEA5F87B40C94D80" box="[325,412,1988,2012]" italics="true" pageId="7">canthus</emphasis>
, external border of upper eyelid, external flanks of the scapular an dorsolateral folds to upper vent; the middorsal raphe has a light cream coloration and is subtly bordered with pale brown; some small black spots flanking the middorsal raphe in the interorbital and scapular areas. Ventrally, the background color is white, densely spotted with black; a white cross is formed by two thin lines that intersect at chest level, the longitudinal line extends from the chin to the vent, and the transversal one through the chest, undersurfaces of upper arms, and forearms to end at the wrists. Other thin white line extends from the vent through the inner ventrolateral surfaces of thighs, ventral surfaces of shanks and tarsi, to end at the soles; at the level of the articulation between the shank and tarsi, a second line arises from the former and extends to the center of heel (
<figureCitation id="8CA670AFC003FF94FA8EFED041F24BF1" captionStart="FIGURE 2" captionStartId="5.[151,250,1524,1546]" captionTargetBox="[151,1436,301,1502]" captionTargetId="figure@5.[151,1436,301,1503]" captionTargetPageId="5" captionText="FIGURE 2. Pristimantis munozi sp. nov. Dorsal (a) and ventral (b) views of the holotype (MZUTI 1782, male) in preservative. Dorsal (c) and ventral (d) views of the paratype (MZUTI 1781, male) in preservative. Scale bars represent 10 mm. Photos: L. Bustamante." httpUri="https://zenodo.org/record/225617/files/figure.png" pageId="8">Fig. 2</figureCitation>
d).
</paragraph>
<caption id="40E23CA2C003FF94FF77FE6340AC484F" ID-Table-UUID="40E23CA2C003FF94FF77FE6340AC484F" httpUri="http://table.plazi.org/id/40E23CA2C003FF94FF77FE6340AC484F" pageId="8" targetBox="[161,1115,558,1336]" targetIsTable="true" targetPageId="8">
<paragraph id="14226C2AC003FF94FF77FE6340AC484F" blockId="8.[151,1436,476,531]" pageId="8">
<emphasis id="26E9B038C003FF94FF77FE63405F4BAF" bold="true" box="[151,266,476,499]" pageId="8">TABLE 1.</emphasis>
Morphometric variation in adult males (
<emphasis id="26E9B038C003FF94FD5FFE6343B34BAF" box="[703,742,476,499]" italics="true" pageId="8">n =</emphasis>
8) of 
<taxonomicName id="D39D17A9C003FF94FCC8FE6342AB4BAF" box="[808,1022,476,499]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C003FF94FCC8FE6342AB4BAF" box="[808,1022,476,499]" italics="true" pageId="8">Pristimantis munozi</emphasis>
</taxonomicName>
. X? ± SD: mean ± standard deviation; Min–Max: minimum–maximum.
</paragraph>
</caption>
<paragraph id="14226C2AC003FF94FF41FD9145174F64" pageId="8">
<table id="669D9E8AC0030063FF41FD91450E4F64" box="[161,1115,558,1336]" gridcols="3" gridrows="17" pageId="8">
<tr id="AAAD6E68C0030063FF41FD91450E48C4" box="[161,1115,558,664]" gridrow="0" pageId="8">
<th id="E97C0714C0030063FF41FD91406648C4" box="[161,307,558,664]" gridcol="0" gridrow="0" pageId="8">Morphometric characters SVL</th>
<th id="E97C0714C0030063FDDCFD91427C48C4" box="[572,809,558,664]" gridcol="1" gridrow="0" pageId="8">
Measurements (in mm) X? 
<emphasis id="26E9B038C003FF94FD80FDE643C34833" bold="true" box="[608,662,600,623]" pageId="8">± SD</emphasis>
16.7 ± 1.6
</th>
<th id="E97C0714C0030063FC16FD91450E48C4" box="[1014,1115,558,664]" gridcol="2" gridrow="0" pageId="8">Min–Max 14.9–19.7</th>
</tr>
<tr id="AAAD6E68C0030063FF41FD13450E489E" box="[161,1115,684,706]" gridrow="1" pageId="8">
<th id="E97C0714C0030063FF41FD134066489E" box="[161,307,684,706]" gridcol="0" gridrow="1" pageId="8">ThL</th>
<td id="E97C0714C0030063FDDCFD13427C489E" box="[572,809,684,706]" gridcol="1" gridrow="1" pageId="8">8.6 ± 0.5</td>
<td id="E97C0714C0030063FC16FD13450E489E" box="[1014,1115,684,706]" gridcol="2" gridrow="1" pageId="8">7.9–9.7</td>
</tr>
<tr id="AAAD6E68C0030063FF41FD69450E48B0" box="[161,1115,726,748]" gridrow="2" pageId="8">
<th id="E97C0714C0030063FF41FD69406648B0" box="[161,307,726,748]" gridcol="0" gridrow="2" pageId="8">SL</th>
<td id="E97C0714C0030063FDDCFD69427C48B0" box="[572,809,726,748]" gridcol="1" gridrow="2" pageId="8">9.7 ± 0.5</td>
<td id="E97C0714C0030063FC16FD69450E48B0" box="[1014,1115,726,748]" gridcol="2" gridrow="2" pageId="8">9.1–10.5</td>
</tr>
<tr id="AAAD6E68C0030063FF41FCBF450E494A" box="[161,1115,768,790]" gridrow="3" pageId="8">
<th id="E97C0714C0030063FF41FCBF4066494A" box="[161,307,768,790]" gridcol="0" gridrow="3" pageId="8">FL</th>
<td id="E97C0714C0030063FDDCFCBF427C494A" box="[572,809,768,790]" gridcol="1" gridrow="3" pageId="8">8.7 ± 0.6</td>
<td id="E97C0714C0030063FC16FCBF450E494A" box="[1014,1115,768,790]" gridcol="2" gridrow="3" pageId="8">7.7–9.5</td>
</tr>
<tr id="AAAD6E68C0030063FF41FC95450E491C" box="[161,1115,810,832]" gridrow="4" pageId="8">
<th id="E97C0714C0030063FF41FC954066491C" box="[161,307,810,832]" gridcol="0" gridrow="4" pageId="8">HaL</th>
<td id="E97C0714C0030063FDDCFC95427C491C" box="[572,809,810,832]" gridcol="1" gridrow="4" pageId="8">4.7 ± 0.4</td>
<td id="E97C0714C0030063FC16FC95450E491C" box="[1014,1115,810,832]" gridcol="2" gridrow="4" pageId="8">4.0–5.2</td>
</tr>
<tr id="AAAD6E68C0030063FF41FCEB450E4936" box="[161,1115,852,874]" gridrow="5" pageId="8">
<th id="E97C0714C0030063FF41FCEB40664936" box="[161,307,852,874]" gridcol="0" gridrow="5" pageId="8">HeL</th>
<td id="E97C0714C0030063FDDCFCEB427C4936" box="[572,809,852,874]" gridcol="1" gridrow="5" pageId="8">6.8 ± 0.5</td>
<td id="E97C0714C0030063FC16FCEB450E4936" box="[1014,1115,852,874]" gridcol="2" gridrow="5" pageId="8">6.0–7.7</td>
</tr>
<tr id="AAAD6E68C0030063FF41FCC1450E49C8" box="[161,1115,894,916]" gridrow="6" pageId="8">
<th id="E97C0714C0030063FF41FCC1406649C8" box="[161,307,894,916]" gridcol="0" gridrow="6" pageId="8">HW</th>
<td id="E97C0714C0030063FDDCFCC1427C49C8" box="[572,809,894,916]" gridcol="1" gridrow="6" pageId="8">6.4 ± 0.4</td>
<td id="E97C0714C0030063FC16FCC1450E49C8" box="[1014,1115,894,916]" gridcol="2" gridrow="6" pageId="8">5.9–7.0</td>
</tr>
<tr id="AAAD6E68C0030063FF41FC17450E49E2" box="[161,1115,936,958]" gridrow="7" pageId="8">
<th id="E97C0714C0030063FF41FC17406649E2" box="[161,307,936,958]" gridcol="0" gridrow="7" pageId="8">InD</th>
<td id="E97C0714C0030063FDDCFC17427C49E2" box="[572,809,936,958]" gridcol="1" gridrow="7" pageId="8">1.8 ± 0.1</td>
<td id="E97C0714C0030063FC16FC17450E49E2" box="[1014,1115,936,958]" gridcol="2" gridrow="7" pageId="8">1.7–1.9</td>
</tr>
<tr id="AAAD6E68C0030063FF41FC6D450E49B4" box="[161,1115,978,1000]" gridrow="8" pageId="8">
<th id="E97C0714C0030063FF41FC6D406649B4" box="[161,307,978,1000]" gridcol="0" gridrow="8" pageId="8">UEW</th>
<td id="E97C0714C0030063FDDCFC6D427C49B4" box="[572,809,978,1000]" gridcol="1" gridrow="8" pageId="8">1.6 ± 0.1</td>
<td id="E97C0714C0030063FC16FC6D450E49B4" box="[1014,1115,978,1000]" gridcol="2" gridrow="8" pageId="8">1.5–1.8</td>
</tr>
<tr id="AAAD6E68C0030063FF41FC43450E4E4E" box="[161,1115,1020,1042]" gridrow="9" pageId="8">
<th id="E97C0714C0030063FF41FC4340664E4E" box="[161,307,1020,1042]" gridcol="0" gridrow="9" pageId="8">EN</th>
<td id="E97C0714C0030063FDDCFC43427C4E4E" box="[572,809,1020,1042]" gridcol="1" gridrow="9" pageId="8">1.7 ± 0.2</td>
<td id="E97C0714C0030063FC16FC43450E4E4E" box="[1014,1115,1020,1042]" gridcol="2" gridrow="9" pageId="8">1.5–2.0</td>
</tr>
<tr id="AAAD6E68C0030063FF41FB99450E4E60" box="[161,1115,1062,1084]" gridrow="10" pageId="8">
<th id="E97C0714C0030063FF41FB9940664E60" box="[161,307,1062,1084]" gridcol="0" gridrow="10" pageId="8">ED</th>
<td id="E97C0714C0030063FDDCFB99427C4E60" box="[572,809,1062,1084]" gridcol="1" gridrow="10" pageId="8">2.1 ± 0.3</td>
<td id="E97C0714C0030063FC16FB99450E4E60" box="[1014,1115,1062,1084]" gridcol="2" gridrow="10" pageId="8">1.7–2.8</td>
</tr>
<tr id="AAAD6E68C0030063FF41FBEF450E4E3A" box="[161,1115,1104,1126]" gridrow="11" pageId="8">
<th id="E97C0714C0030063FF41FBEF40664E3A" box="[161,307,1104,1126]" gridcol="0" gridrow="11" pageId="8">ETS</th>
<td id="E97C0714C0030063FDDCFBEF427C4E3A" box="[572,809,1104,1126]" gridcol="1" gridrow="11" pageId="8">3.0 ± 0.2</td>
<td id="E97C0714C0030063FC16FBEF450E4E3A" box="[1014,1115,1104,1126]" gridcol="2" gridrow="11" pageId="8">2.7–3.4</td>
</tr>
<tr id="AAAD6E68C0030063FF41FBC5450E4ECC" box="[161,1115,1146,1168]" gridrow="12" pageId="8">
<th id="E97C0714C0030063FF41FBC540664ECC" box="[161,307,1146,1168]" gridcol="0" gridrow="12" pageId="8">TD</th>
<td id="E97C0714C0030063FDDCFBC5427C4ECC" box="[572,809,1146,1168]" gridcol="1" gridrow="12" pageId="8">1.0 ± 0.2</td>
<td id="E97C0714C0030063FC16FBC5450E4ECC" box="[1014,1115,1146,1168]" gridcol="2" gridrow="12" pageId="8">0.8–1.3</td>
</tr>
<tr id="AAAD6E68C0030063FF41FB1B450E4EE6" box="[161,1115,1188,1210]" gridrow="13" pageId="8">
<th id="E97C0714C0030063FF41FB1B40664EE6" box="[161,307,1188,1210]" gridcol="0" gridrow="13" pageId="8">F3D</th>
<td id="E97C0714C0030063FDDCFB1B427C4EE6" box="[572,809,1188,1210]" gridcol="1" gridrow="13" pageId="8">0.6 ± 0.0</td>
<td id="E97C0714C0030063FC16FB1B450E4EE6" box="[1014,1115,1188,1210]" gridcol="2" gridrow="13" pageId="8">0.5–0.7</td>
</tr>
<tr id="AAAD6E68C0030063FF41FB71450E4EB8" box="[161,1115,1230,1252]" gridrow="14" pageId="8">
<th id="E97C0714C0030063FF41FB7140664EB8" box="[161,307,1230,1252]" gridcol="0" gridrow="14" pageId="8">T4D</th>
<td id="E97C0714C0030063FDDCFB71427C4EB8" box="[572,809,1230,1252]" gridcol="1" gridrow="14" pageId="8">0.7 ± 0.1</td>
<td id="E97C0714C0030063FC16FB71450E4EB8" box="[1014,1115,1230,1252]" gridcol="2" gridrow="14" pageId="8">0.5–0.8</td>
</tr>
<tr id="AAAD6E68C0030063FF41FB47450E4F52" box="[161,1115,1272,1294]" gridrow="15" pageId="8">
<th id="E97C0714C0030063FF41FB4740664F52" box="[161,307,1272,1294]" gridcol="0" gridrow="15" pageId="8">F1L</th>
<td id="E97C0714C0030063FDDCFB47427C4F52" box="[572,809,1272,1294]" gridcol="1" gridrow="15" pageId="8">2.6 ± 0.2</td>
<td id="E97C0714C0030063FC16FB47450E4F52" box="[1014,1115,1272,1294]" gridcol="2" gridrow="15" pageId="8">2.3–2.9</td>
</tr>
<tr id="AAAD6E68C0030063FF41FA9D450E4F64" box="[161,1115,1314,1336]" gridrow="16" pageId="8">
<th id="E97C0714C0030063FF41FA9D40664F64" box="[161,307,1314,1336]" gridcol="0" gridrow="16" pageId="8">F2L</th>
<td id="E97C0714C0030063FDDCFA9D427C4F64" box="[572,809,1314,1336]" gridcol="1" gridrow="16" pageId="8">3.2 ± 0.2</td>
<td id="E97C0714C0030063FC16FA9D450E4F64" box="[1014,1115,1314,1336]" gridcol="2" gridrow="16" pageId="8">2.9–3.5</td>
</tr>
</table>
</paragraph>
<paragraph id="14226C2AC003FF94FF27FAC243DD4CBF" blockId="8.[151,1436,1405,2015]" pageId="8">
<emphasis id="26E9B038C003FF94FF27FAC243FC4FCA" bold="true" box="[199,681,1405,1430]" pageId="8">
Call (adult male, MZUTI 1777; 
<figureCitation id="8CA670AFC003FF94FDB3FAC243CF4FCA" box="[595,666,1405,1430]" captionStart="FIGURE 4" captionStartId="9.[151,250,1750,1772]" captionTargetBox="[221,1371,197,1719]" captionTargetId="figure@9.[188,1398,193,1729]" captionTargetPageId="9" captionText="FIGURE 4. Vocalizations of Pristimantis munozi, adult male, MZUTI 1777. (a) Waveform of a call series with four calls. (b) Waveform and (c) Spectrogram of a single, pulsed call. (d) Power-spectrum of a call, showing peak of dominant frequency." httpUri="https://zenodo.org/record/225619/files/figure.png" pageId="8">Fig. 4</figureCitation>
).
</emphasis>
The advertisement call of 
<taxonomicName id="D39D17A9C003FF94FC04FAC045924FCA" box="[996,1223,1407,1430]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C003FF94FC04FAC045924FCA" box="[996,1223,1407,1430]" italics="true" pageId="8">Pristimantis munozi</emphasis>
</taxonomicName>
is composed by a series of calls, each with a duration of 1.589– 
<geoCoordinate id="71A90AEDC003FF94FD44FA1D43B14FE7" box="[676,740,1442,1467]" direction="south" orientation="latitude" pageId="8" precision="55" value="-2.492">2.492</geoCoordinate>
s (mean = 2.160 ± 
<geoCoordinate id="71A90AEDC003FF94FC2BFA1D454A4FE6" box="[971,1055,1442,1466]" direction="south" orientation="latitude" pageId="8" precision="55" value="-0.424">0.424 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FBCEFA1C45144FE6" box="[1070,1089,1443,1466]" italics="true" pageId="8">N</emphasis>
= 6) and containing 3–5 calls (mean = 3.833 ± 0.753, 
<emphasis id="26E9B038C003FF94FE50FA7640964FBC" box="[432,451,1481,1504]" italics="true" pageId="8">N</emphasis>
= 6), the first of which is always longer than the remaining calls (length of 1st call = 0.160–0.200, mean = 0.187 ± 
<geoCoordinate id="71A90AEDC003FF94FE09FA52436E4C58" box="[489,571,1517,1541]" direction="south" orientation="latitude" pageId="8" precision="55" value="-0.014">0.014 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FDA9FA5243094C58" box="[585,604,1517,1540]" italics="true" pageId="8">N</emphasis>
= 6; length of subsequent calls = 0.116–0.153, mean = 0.151 ± 
<geoCoordinate id="71A90AEDC003FF94FAC9FA5244294C58" box="[1321,1404,1517,1541]" direction="south" orientation="latitude" pageId="8" precision="55" value="-0.027">0.027 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FA69FA5244C94C58" box="[1417,1436,1517,1540]" italics="true" pageId="8">N</emphasis>
= 16). Series of calls are produced every 1.589– 
<geoCoordinate id="71A90AEDC003FF94FD4DF9AF43AF4C75" box="[685,762,1552,1577]" direction="south" orientation="latitude" pageId="8" precision="55" value="-18.992">18.992</geoCoordinate>
s (mean = 9.870 ± 
<geoCoordinate id="71A90AEDC003FF94FC32F9AF45714C74" box="[978,1060,1552,1577]" direction="south" orientation="latitude" pageId="8" precision="55" value="-6.424">6.424 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FBD0F9AE45164C74" box="[1072,1091,1553,1576]" italics="true" pageId="8">N</emphasis>
= 5). Within a call series, calls are emitted with an interval of 0.501– 
<geoCoordinate id="71A90AEDC003FF94FDA1F98B43D54C11" box="[577,640,1588,1613]" direction="south" orientation="latitude" pageId="8" precision="55" value="-0.783">0.783</geoCoordinate>
s (mean = 0.608 ± 
<geoCoordinate id="71A90AEDC003FF94FCBEF98B42E44C10" box="[862,945,1588,1613]" direction="south" orientation="latitude" pageId="8" precision="55" value="-0.075">0.075 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FC5EF98A42844C10" box="[958,977,1589,1612]" italics="true" pageId="8">N</emphasis>
= 16). Each call is strongly pulsed, with the first call having more pulsed than the subsequent calls (1st call with 16–20 pulses, mean = 18.667 ± 
<geoCoordinate id="71A90AEDC003FF94FAF5F9E444334C2F" box="[1301,1382,1627,1652]" direction="south" orientation="latitude" pageId="8" precision="55" value="-2.066">2.066 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FA93F9E344D34C2F" box="[1395,1414,1628,1651]" italics="true" pageId="8">N</emphasis>
= 6; subsequent calls with16–20 pulses, mean = 13.467 ± 
<geoCoordinate id="71A90AEDC003FF94FCD9F93F42DA4CCB" box="[825,911,1664,1688]" direction="south" orientation="latitude" pageId="8" precision="55" value="-1.119">1.119 s</geoCoordinate>
, 
<emphasis id="26E9B038C003FF94FC41F93F42E14CCB" box="[929,948,1664,1687]" italics="true" pageId="8">N</emphasis>
= 16). The dominant frequency (also de fundamental frequency) is at 
<date id="60234AEAC003FF94FE3EF91843084CE3" box="[478,605,1703,1727]" pageId="8" value="2198" valueMax="2248">2198–2248</date>
Hz (mean = 2230 ± 16.3 Hz, 
<emphasis id="26E9B038C003FF94FC4AF91842E84CE2" box="[938,957,1703,1726]" italics="true" pageId="8">N</emphasis>
= 21). Finally, the 1st harmonic is at 3776– 4357 Hz (mean = 4143.3 ± 225 Hz, 
<emphasis id="26E9B038C003FF94FDCBF974436B4CBE" box="[555,574,1739,1762]" italics="true" pageId="8">N</emphasis>
= 22).
</paragraph>
</subSubSection>
<subSubSection id="5C873FA1C003FF96FF27F95141864B40" lastPageId="10" pageId="8" type="distribution">
<paragraph id="14226C2AC003FF94FF27F95140854D13" blockId="8.[151,1436,1405,2015]" pageId="8">
<emphasis id="26E9B038C003FF94FF27F95140354D5B" bold="true" box="[199,352,1774,1799]" pageId="8">Distribution.</emphasis>
<taxonomicName id="D39D17A9C003FF94FE8DF95043014D5A" box="[365,596,1775,1798]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C003FF94FE8DF95043014D5A" box="[365,596,1775,1798]" italics="true" pageId="8">Pristimantis munozi</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="CB26D288C003FF94FC6BF95042EE4D5B" box="[907,955,1775,1799]" pageId="8">type</typeStatus>
locality, Reserva Ecológica Verdecocha, (
<geoCoordinate id="71A90AEDC003FF94FF7FF8AD404E4D77" box="[159,283,1810,1835]" direction="south" orientation="latitude" pageId="8" precision="1" value="-0.096361116">0°5'46.9&quot;S</geoCoordinate>
, 
<geoCoordinate id="71A90AEDC003FF94FECCF8AD40984D77" box="[300,461,1810,1835]" direction="west" orientation="longitude" pageId="8" precision="1" value="-78.60425">78°36'15.3&quot;W</geoCoordinate>
; elevation 
<quantity id="D365C1CFC003FF94FDB1F8AD43FB4D77" box="[593,686,1810,1835]" metricMagnitude="3" metricUnit="m" metricValue="2.851" pageId="8" unit="m" value="2851.0">2851 m</quantity>
), located at northwestern flank of the Volcán Pichincha, in the Pichincha province (
<figureCitation id="8CA670AFC003FF94FE9DF88940974D13" box="[381,450,1846,1871]" captionStart="FIGURE 5" captionStartId="10.[151,250,1150,1172]" captionTargetBox="[234,1352,344,1118]" captionTargetId="figure@10.[226,1361,337,1129]" captionTargetPageId="10" captionText="FIGURE 5. Distribution of Pristimantis munozi sp. nov. in the Ecuadorian Andes. White dot: Reserva Verdecocha (type locality)." httpUri="https://zenodo.org/record/225620/files/figure.png" pageId="8">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="14226C2AC003FF94FF27F8E545A84D83" blockId="8.[151,1436,1405,2015]" pageId="8">
<emphasis id="26E9B038C003FF94FF27F8E5437F4D2F" bold="true" box="[199,554,1882,1907]" pageId="8">Hábitat and natural history.</emphasis>
The vegetation of the 
<typeStatus id="CB26D288C003FF94FCA6F8E442234D2F" box="[838,886,1883,1907]" pageId="8">type</typeStatus>
locality is described as montane cloud forest (
<bibRefCitation id="700C11DBC003FF94FF7EF8C140D64DCB" author="Valencia" box="[158,387,1918,1943]" pageId="8" refString="Valencia, R., Ceron, C., Palacios, W. &amp; Sierra, R. (1999) Las formaciones naturales de la sierra del Ecuador. In: Sierra, R. (Ed.), Propuesta preliminar de un sistema de clasificacion de vegetacion para el Ecuador continental. Proyecto INEFA / GEF- BIRF y EcoCiencia, Quito, pp. 79 - 108." type="book chapter" year="1999">
Valencia 
<emphasis id="26E9B038C003FF94FEE8F8C0406C4DCA" box="[264,313,1918,1942]" italics="true" pageId="8">et al</emphasis>
. 1999
</bibRefCitation>
). The region has a bi-seasonal climatic regime, and present a bimodal pattern of rainfall, with two rainy periods in February-April and September-November (with peaks of maximum rainfall on March-April and October), and two dry periods in June-August and December-January (
<bibRefCitation id="700C11DBC003FF94FC00F87945A44D83" author="Neill" box="[992,1265,1990,2015]" pageId="8" refString="Neill, D. A. &amp; Jorgensen, P. M. (1999) Climates. In: Jorgensen, P. M. &amp; Leon-Yanez, S. (Eds.), Catalogue of the vascular plants of Ecuador. Monographies in Systematic Botany from the Missouri Botanical Garden, 75, 8 - 13." type="journal article" year="1999">Neill &amp; Jørgensen 1999</bibRefCitation>
).
</paragraph>
<caption id="40E23CA2C002FF95FF77F969442D4D57" httpUri="https://zenodo.org/record/225619/files/figure.png" pageId="9" targetBox="[221,1371,197,1719]" targetPageId="9">
<paragraph id="14226C2AC002FF95FF77F969442D4D57" blockId="9.[151,1436,1750,1803]" pageId="9">
<emphasis id="26E9B038C002FF95FF77F96940464CB0" bold="true" box="[151,275,1750,1772]" pageId="9">FIGURE 4.</emphasis>
Vocalizations of 
<taxonomicName id="D39D17A9C002FF95FE25F96743C44CB1" box="[453,657,1752,1773]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C002FF95FE25F96743C44CB1" box="[453,657,1752,1773]" italics="true" pageId="9">Pristimantis munozi</emphasis>
</taxonomicName>
, adult male, MZUTI 1777. (a) Waveform of a call series with four calls. (b) Waveform and (c) Spectrogram of a single, pulsed call. (d) Power-spectrum of a call, showing peak of dominant frequency.
</paragraph>
</caption>
<paragraph id="14226C2AC002FF96FF27F88C41864B40" blockId="9.[151,1436,1842,2011]" lastBlockId="10.[151,1436,151,284]" lastPageId="10" pageId="9">
<taxonomicName id="D39D17A9C002FF95FF27F88C40FD4D16" box="[199,424,1843,1866]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C002FF95FF27F88C40FD4D16" box="[199,424,1843,1866]" italics="true" pageId="9">Pristimantis munozi</emphasis>
</taxonomicName>
is a nocturnal and secretive frog, apparently associated to bamboo forests (
<taxonomicName id="D39D17A9C002FF95FB14F88C44314D17" box="[1268,1380,1843,1867]" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" order="Poales" pageId="9" phylum="Tracheophyta" rank="genus">
<emphasis id="26E9B038C002FF95FB14F88C44314D17" box="[1268,1380,1843,1867]" italics="true" pageId="9">Chusquea</emphasis>
</taxonomicName>
sp.). Even though several 
<typeStatus id="CB26D288C002FF95FE61F8E840E94D33" box="[385,444,1879,1903]" pageId="9">types</typeStatus>
of habitat were intensively sampled at Verdecocha (primary forest, scrubland, secondary forest, grassland and bamboo patches), the new species only was detected in a small patch of bamboo forest at 
<quantity id="D365C1CFC002FF95FA84F8C541F84DEA" metricMagnitude="3" metricUnit="m" metricValue="2.851" pageId="9" unit="m" value="2851.0">2851 m</quantity>
(
<figureCitation id="8CA670AFC002FF95FF5EF82140514DEB" box="[190,260,1950,1975]" captionStart="FIGURE 6" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[151,1436,195,1909]" captionTargetId="figure@11.[151,1436,195,1910]" captionTargetPageId="11" captionText="FIGURE 6. General view of the bamboo forest at the Reserva Verdecocha, where Pristimantis munozi sp. nov. inhabits. White arrows indicate the sites where two specimens of the type series were found while calling hidden. Photo: F. J. M. Rojas-Runjaic." httpUri="https://zenodo.org/record/225621/files/figure.png" pageId="9">Fig. 6</figureCitation>
). Numerous males were heard calling between 23:50 and 00:55 h, but due to its cryptic coloration and cryptic behavior (males calling from underneath bamboo leaf litter and mosses, hollows and shelters on the slope of a small seasonal creek) only eight specimens were visually detected and collected. Despite the effort no females were found. At Verdecocha, 
<taxonomicName id="D39D17A9C001FF96FE3DFF0243F34A88" box="[477,678,188,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" phylum="Chordata" rank="species" species="leoni">
<emphasis id="26E9B038C001FF96FE3DFF0243F34A88" box="[477,678,188,212]" italics="true" pageId="10">Pristimantis leoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D39D17A9C001FF96FD00FF02420C4A88" box="[736,857,188,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" phylum="Chordata" rank="species" species="sirnigeli">
<emphasis id="26E9B038C001FF96FD00FF02420C4A88" box="[736,857,188,212]" italics="true" pageId="10">P. sirnigeli</emphasis>
</taxonomicName>
are sympatric with 
<emphasis id="26E9B038C001FF96FBDEFF0245FD4A88" box="[1086,1192,189,212]" italics="true" pageId="10">P. m u n o z i</emphasis>
, but none was found syntopically with the new species at the bamboo forest, but in adjacent cloud forest dominated by relatively short trees.
</paragraph>
</subSubSection>
<caption id="40E23CA2C001FF96FF77FBC141A74EEF" httpUri="https://zenodo.org/record/225620/files/figure.png" pageId="10" targetBox="[234,1352,344,1118]" targetPageId="10">
<paragraph id="14226C2AC001FF96FF77FBC141A74EEF" blockId="10.[151,1435,1150,1203]" pageId="10">
<emphasis id="26E9B038C001FF96FF77FBC140434EC8" bold="true" box="[151,278,1150,1172]" pageId="10">FIGURE 5.</emphasis>
Distribution of 
<taxonomicName id="D39D17A9C001FF96FE24FBC043C64EC8" box="[452,659,1151,1172]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" phylum="Chordata" rank="species" species="munozi" status="sp. nov.">
<emphasis id="26E9B038C001FF96FE24FBC043C64EC8" box="[452,659,1151,1172]" italics="true" pageId="10">Pristimantis munozi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DDA0D43C001FF96FD7EFBC043A74EC8" box="[670,754,1151,1172]" pageId="10" rank="species">
<emphasis id="26E9B038C001FF96FD7EFBC043A74EC8" bold="true" box="[670,754,1151,1172]" pageId="10">sp. nov.</emphasis>
</taxonomicNameLabel>
in the Ecuadorian Andes. White dot: Reserva Verdecocha (type locality).
</paragraph>
</caption>
<subSubSection id="5C873FA1C001FF96FF27FB65436E4FB3" pageId="10" type="etymology">
<paragraph id="14226C2AC001FF96FF27FB65436E4FB3" blockId="10.[151,1436,1242,1519]" pageId="10">
<emphasis id="26E9B038C001FF96FF27FB65401B4EAF" bold="true" box="[199,334,1242,1267]" pageId="10">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="D39D17A9C001FF96FDAAFB6443CE4EAE" box="[586,667,1243,1266]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" phylum="Chordata" rank="species" species="munozi">
<emphasis id="26E9B038C001FF96FDAAFB6443CE4EAE" box="[586,667,1243,1266]" italics="true" pageId="10">munozi</emphasis>
</taxonomicName>
is a patronym honoring Jésus Muñoz Fuente, prominent Spanish botanist of the Real Jardín Botánico of 
<collectingCountry id="6C8A2CBAC001FF96FDB5FB4143CD4F4B" box="[597,664,1278,1303]" name="Spain" pageId="10">Spain</collectingCountry>
, founder and Director of the Master’s Program in “Biodiversidad en Áreas Tropicales y su Conservation” (MBATC) of the Universidad Internacional Menéndez Pelayo, 
<collectingCountry id="6C8A2CBAC001FF96FB16FA9D44634F67" box="[1270,1334,1314,1339]" name="Spain" pageId="10">Spain</collectingCountry>
(UIMP), and research affiliate of the Centro para la Investigación de la Biodiversidad y Cambio Climático, BioCamb, of the Universidad Tecnológica Indoamérica, 
<collectingCountry id="6C8A2CBAC001FF96FDB7FAD543E34FDF" box="[599,694,1386,1411]" name="Ecuador" pageId="10">Ecuador</collectingCountry>
. Through this program the Dr. Muñoz Fuente has formed over a hundred of professionals (including the two first authors of this work), contributing significantly to the study and conservation of the Neotropical biodiversity. We make recognition to his admirable work, support, and friendship, naming this new species in his honor.
</paragraph>
</subSubSection>
</treatment>
</document>