128 lines
18 KiB
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128 lines
18 KiB
XML
<document id="6EB3BBBA384029DF161168F35403BED1" ID-DOI="10.14446/FI.2016.78" ID-ISSN="2533-4069" ID-Zenodo-Dep="4773056" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621431330703" checkinUser="felipe" docAuthor="Gardner, James D., Redman, Cory M. & Cifelli, Richard L." docDate="2016" docId="947587F2667BFFC5FC0E1133F9DEF92A" docLanguage="en" docName="FossilImprint.72.1-2.78-107.pdf" docOrigin="Fossil Imprint 72 (1 - 2)" docSource="http://fi.nm.cz/clanek/the-hopping-dead-late-cretaceous-frogs-from-the-middle-late-campanian-judithian-of-western-north-america/" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Anura Gardner & Redman & Cifelli 2016" docType="treatment" docVersion="3" lastPageNumber="93" masterDocId="684CFF8A6675FFCAFFC91418F909FF88" masterDocTitle="The Hopping Dead: Late Cretaceous Frogs From The Middle - Late Campanian (Judithian) Of Western North America" masterLastPageNumber="107" masterPageNumber="78" pageNumber="92" updateTime="1698921899666" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="F868C3A2E5BEF48162F6B7DF4D66E0BA">The Hopping Dead: Late Cretaceous Frogs From The Middle - Late Campanian (Judithian) Of Western North America</mods:title>
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<mods:namePart id="FA023BCB60E9CE128BD4CF77C26E8E45">Gardner, James D.</mods:namePart>
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<mods:roleTerm id="937774A6B6475268A5CC0CFF9D545CB8">Author</mods:roleTerm>
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<mods:namePart id="4CCB0DF2A238810CF42D5A49CB5520B3">Redman, Cory M.</mods:namePart>
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<mods:name id="D0739062BE5C1847DE5F17EDBF0C0566" type="personal">
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<mods:roleTerm id="EDE50FD20BA2C591575FF7CC548789DE">Author</mods:roleTerm>
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<mods:namePart id="F31B3BB281E7CF24AF40A8282B0AD29B">Cifelli, Richard L.</mods:namePart>
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<mods:typeOfResource id="2CC0BAD17AB12B9AB1F94C29044AD827">text</mods:typeOfResource>
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<mods:title id="947BD48E179FD1E6A139F1092438FED2">Fossil Imprint</mods:title>
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<mods:date id="0A4E0071F7DEB4995696281E8156DC3D">2016</mods:date>
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<mods:number id="F63344B88F10DC46B65D8DEABD7D491A">2016-08-15</mods:number>
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<mods:number id="DF803E6A61BE91F036CAE5C42183F53F">72</mods:number>
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<mods:number id="80CF2CCD518D8D31F7334AAFAAA907D2">1 - 2</mods:number>
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<mods:classification id="DEBA49AA67E76E2522B5971B516FDBAB">journal article</mods:classification>
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<mods:identifier id="425B83BD91D0ABA82B0A2BF98D9EEF7E" type="DOI">10.14446/FI.2016.78</mods:identifier>
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<mods:identifier id="5F8ABAE1DDC1A0B72F572C52AC3AFA58" type="ISSN">2533-4069</mods:identifier>
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<subSubSection id="54C6656F667BFFC4FC0E1133FC16FACA" box="[967,1311,1323,1346]" pageId="14" pageNumber="92" type="nomenclature">
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<paragraph id="1C6336E4667BFFC4FC0E1133FC16FACA" blockId="14.[967,1311,1323,1346]" box="[967,1311,1323,1346]" pageId="14" pageNumber="92">
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<heading id="472B8188667BFFC4FC0E1133FC16FACA" box="[967,1311,1323,1346]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="92" reason="2">
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<emphasis id="2EA8EAF6667BFFC4FC0E1133FC16FACA" bold="true" box="[967,1311,1323,1346]" pageId="14" pageNumber="92">
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<taxonomicName id="DBDC4D67667BFFC4FC0E1133FD18FACA" ID-CoL="PW" authorityName="Gardner & Redman & Cifelli" authorityYear="2016" box="[967,1041,1323,1346]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="92" phylum="Chordata" rank="order">Anura</taxonomicName>
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indeterminate morph 2
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="54C6656F667BFFC5FBFF1143F9DEF92A" lastPageId="15" lastPageNumber="93" pageId="14" pageNumber="92" type="description">
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<paragraph id="1C6336E4667BFFC4FBFF1143FDB8FAF9" blockId="14.[1078,1201,1371,1393]" box="[1078,1201,1371,1393]" pageId="14" pageNumber="92">
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(
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<figureCitation id="84E72A61667BFFC4FBF41143FDA0FAF9" box="[1085,1193,1371,1393]" captionStart="Text-fig" captionStartId="14.[106,185,633,654]" captionTargetBox="[106,1459,152,589]" captionTargetId="figure-656@14.[124,1457,168,590]" captionTargetPageId="14" captionText="Text-fig. 9. Maxillae ofAnura indeterminate morph 2 from the middle – late Campanian (Judithian) ofAlberta, Canada. Both specimens are from the Irvine locality in the Dinosaur Park Formation. Except where noted otherwise, images are photographs and depict specimens lightly dusted with ammonium chloride to enhance details and texture. Images at same magnification; see scale bar. a, b – incomplete left maxilla, UALVP 40191, in labial (a) and lingual (b) views. c–g – incomplete left maxilla, UALVP 40192, in labial (c, d) and lingual (e, f) views and, with anterior to right, in dorsal (g) view (images in d and f are scanning electron micrographs). Arrow (f) points at groove extending diagonally along lingual surface of pars facialis below margo orbitalis." figureDoi="http://doi.org/10.5281/zenodo.4773087" httpUri="https://zenodo.org/record/4773087/files/figure.png" pageId="14" pageNumber="92">Text-fig. 9</figureCitation>
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)
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</paragraph>
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<paragraph id="1C6336E4667BFFC4FC931191FCBAFA28" blockId="14.[820,1459,1417,1472]" box="[858,1459,1417,1440]" pageId="14" pageNumber="92">M a t e r i a l a n d o c c u r r e n c e s: Two maxillae from</paragraph>
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<paragraph id="1C6336E4667BFFC4FCFD11B1FC8DFA48" blockId="14.[820,1459,1417,1472]" box="[820,1412,1449,1472]" pageId="14" pageNumber="92">
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Dinosaur Park Formation,
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<collectingRegion id="DE18F806667BFFC4FB8C11B1FD9CFA48" box="[1093,1173,1449,1472]" country="Canada" name="Alberta" pageId="14" pageNumber="92">Alberta</collectingRegion>
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,
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<collectingCountry id="64CB7674667BFFC4FB5611B1FDE4FA48" box="[1183,1261,1449,1472]" name="Canada" pageId="14" pageNumber="92">Canada</collectingCountry>
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(Appendix 2).
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</paragraph>
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<paragraph id="1C6336E4667BFFC5FC9311C4F85EFBE7" blockId="14.[820,1460,1500,2099]" lastBlockId="15.[128,768,152,1135]" lastPageId="15" lastPageNumber="93" pageId="14" pageNumber="92">
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<materialsCitation id="ACB43CB9667BFFC5FC9311C4F85AFBE7" collectionCode="UALVP" lastPageId="15" lastPageNumber="93" pageId="14" pageNumber="92" specimenCode="UALVP 40191, UALVP 40192" specimenCount="2">
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D e s c r i p t i o n: The
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<specimenCount id="0ADAFD6D667BFFC4FB9C11C4FDFAFA7B" box="[1109,1267,1500,1523]" pageId="14" pageNumber="92" type="generic">two specimens</specimenCount>
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are both from the Irvine locality and preserve about the posterior two-thirds of a left maxilla.
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<specimenCode id="4C7A9E9F667BFFC4FC171204FD8DF9BB" box="[990,1156,1564,1587]" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40191</specimenCode>
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(
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<figureCitation id="84E72A61667BFFC4FB5F1204FC35F9BB" box="[1174,1340,1564,1587]" captionStart="Text-fig" captionStartId="14.[106,185,633,654]" captionTargetBox="[106,1459,152,589]" captionTargetId="figure-656@14.[124,1457,168,590]" captionTargetPageId="14" captionText="Text-fig. 9. Maxillae ofAnura indeterminate morph 2 from the middle – late Campanian (Judithian) ofAlberta, Canada. Both specimens are from the Irvine locality in the Dinosaur Park Formation. Except where noted otherwise, images are photographs and depict specimens lightly dusted with ammonium chloride to enhance details and texture. Images at same magnification; see scale bar. a, b – incomplete left maxilla, UALVP 40191, in labial (a) and lingual (b) views. c–g – incomplete left maxilla, UALVP 40192, in labial (c, d) and lingual (e, f) views and, with anterior to right, in dorsal (g) view (images in d and f are scanning electron micrographs). Arrow (f) points at groove extending diagonally along lingual surface of pars facialis below margo orbitalis." figureDoi="http://doi.org/10.5281/zenodo.4773087" httpUri="https://zenodo.org/record/4773087/files/figure.png" pageId="14" pageNumber="92">Text-fig. 9a, b</figureCitation>
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) lacks the posteriormost end of the processus posterior and most of the processus pterygoideus, but anteriorly preserves the base of the processus palatinus.
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<specimenCode id="4C7A9E9F667BFFC4FB881264FDEBF91B" box="[1089,1250,1660,1683]" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40192</specimenCode>
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(
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<figureCitation id="84E72A61667BFFC4FB3A1264FC99F91B" box="[1267,1424,1660,1683]" captionStart="Text-fig" captionStartId="14.[106,185,633,654]" captionTargetBox="[106,1459,152,589]" captionTargetId="figure-656@14.[124,1457,168,590]" captionTargetPageId="14" captionText="Text-fig. 9. Maxillae ofAnura indeterminate morph 2 from the middle – late Campanian (Judithian) ofAlberta, Canada. Both specimens are from the Irvine locality in the Dinosaur Park Formation. Except where noted otherwise, images are photographs and depict specimens lightly dusted with ammonium chloride to enhance details and texture. Images at same magnification; see scale bar. a, b – incomplete left maxilla, UALVP 40191, in labial (a) and lingual (b) views. c–g – incomplete left maxilla, UALVP 40192, in labial (c, d) and lingual (e, f) views and, with anterior to right, in dorsal (g) view (images in d and f are scanning electron micrographs). Arrow (f) points at groove extending diagonally along lingual surface of pars facialis below margo orbitalis." figureDoi="http://doi.org/10.5281/zenodo.4773087" httpUri="https://zenodo.org/record/4773087/files/figure.png" pageId="14" pageNumber="92">Text-fig. 9c–g</figureCitation>
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) is broken farther behind the processus palatinus, but preserves an intact processus posterior and more of the processus pterygoideus. The
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<specimenCount id="0ADAFD6D667BFFC4FC3E12C4FD9BF97B" box="[1015,1170,1756,1779]" pageId="14" pageNumber="92" type="generic">two specimens</specimenCount>
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are similar in size (
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<specimenCode id="4C7A9E9F667BFFC4FA9512C4FA7EF89B" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40191</specimenCode>
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is 6.0 mm long and
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<specimenCode id="4C7A9E9F667BFFC4FB9112E4FDF1F89B" box="[1112,1272,1788,1811]" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40192</specimenCode>
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is 6.7 mm long); the latter is from a slightly larger individual. They also resemble one another in the following features: bone elongate and low in labial or lingual outline; margo orbitalis shallowly concave and moderately elongate; processus zygomaticomaxillaris low and broadly convex dorsally, bearing grooves dorsolingually for contact with squamosal (these features are more prominently developed in the larger
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<specimenCode id="4C7A9E9F667BFFC4FACB13C4FCA1F87B" box="[1282,1448,2012,2035]" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40192</specimenCode>
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); processus posterior only moderately elongate, with its posterodorsal edge moderately (
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<specimenCode id="4C7A9E9F667BFFC4FB541C04FC4FF7BB" box="[1181,1350,2076,2099]" collectionCode="UALVP" pageId="14" pageNumber="92">UALVP 40191</specimenCode>
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) or more steeply (
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<specimenCode id="4C7A9E9F667AFFC5FF141480F88BFF27" box="[221,386,152,175]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40192</specimenCode>
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) declined; and lamina horizontalis moderately deep and lingually wide, with convex lingual surface. Although neither specimen retains any intact teeth, as indicated by their preserved tooth shafts the teeth are small and closely spaced. Judging by the profile along the anteriormost portion of the preserved dorsal edge in
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<specimenCode id="4C7A9E9F667AFFC5FD631520F9CEFEE7" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40191</specimenCode>
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, the processus frontalis would have been taller than the processus zygomatico-maxillaris.
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<specimenCode id="4C7A9E9F667AFFC5FDFC1560FBD7FE07" box="[565,734,376,399]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40192</specimenCode>
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is informative for showing that the processus posterior tapers to a blunt point, that near its posterior end that same process lingually bears a small, shallow facet for contact with the jugal, and that the processus pterygoideus was a welldeveloped, lingually projecting flange with a shallowly concave dorsal surface. The specimens exhibit three differences. First, the posterior end of the tooth row lies approximately in line with (
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<specimenCode id="4C7A9E9F667AFFC5FE001660FB7DFD07" box="[457,628,632,655]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40191</specimenCode>
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) or slightly behind (
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<specimenCode id="4C7A9E9F667AFFC5FF2E1680F894FD27" box="[231,413,664,687]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40192</specimenCode>
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) the level of the processus pterygoideus. Second, a deep groove descends from the margo orbitalis anteroventrally along the lingual face of the pars facialis in
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<specimenCode id="4C7A9E9F667AFFC5FEE816E0F8B6FC87" box="[289,447,760,783]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40192</specimenCode>
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(
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<figureCitation id="84E72A61667AFFC5FE0516E0FB43FC87" box="[460,586,760,783]" captionStart="Text-fig" captionStartId="14.[106,185,633,654]" captionTargetBox="[106,1459,152,589]" captionTargetId="figure-656@14.[124,1457,168,590]" captionTargetPageId="14" captionText="Text-fig. 9. Maxillae ofAnura indeterminate morph 2 from the middle – late Campanian (Judithian) ofAlberta, Canada. Both specimens are from the Irvine locality in the Dinosaur Park Formation. Except where noted otherwise, images are photographs and depict specimens lightly dusted with ammonium chloride to enhance details and texture. Images at same magnification; see scale bar. a, b – incomplete left maxilla, UALVP 40191, in labial (a) and lingual (b) views. c–g – incomplete left maxilla, UALVP 40192, in labial (c, d) and lingual (e, f) views and, with anterior to right, in dorsal (g) view (images in d and f are scanning electron micrographs). Arrow (f) points at groove extending diagonally along lingual surface of pars facialis below margo orbitalis." figureDoi="http://doi.org/10.5281/zenodo.4773087" httpUri="https://zenodo.org/record/4773087/files/figure.png" pageId="15" pageNumber="93">Text-fig. 9f</figureCitation>
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: arrow), whereas no such groove is present in
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<specimenCode id="4C7A9E9F667AFFC5FE051700FB7DFCA7" box="[460,628,792,815]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40191</specimenCode>
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. Finally, the labial surface of the pars facialis in
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<specimenCode id="4C7A9E9F667AFFC5FDFE1720FBD6FCC7" box="[567,735,824,847]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40191</specimenCode>
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is ornamented with low ridges that are arranged in a loosely reticulate pattern and enclose shallow pits, whereas in
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<specimenCode id="4C7A9E9F667AFFC5FF481780F839FC27" box="[129,304,920,943]" collectionCode="UALVP" pageId="15" pageNumber="93">UALVP 40192</specimenCode>
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the labial surface is less obviously ornamented. Instead, the labial surface of the latter specimen has a roughened texture and, when viewed under low angle lighting, extremely low ridges enclosing shallow pits can be seen along the dorsal half of its labial surface. This less pronounced labial ornament does not seem to be an artefact of wear or abrasion
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="1C6336E4667AFFC5FF6F1093F9DEF92A" blockId="15.[129,768,1163,1698]" pageId="15" pageNumber="93">R e m a r k s: The two small, low, and elongate maxillae from Irvine are similar to one another in many respects, but exhibit some differences (i.e., relative positions of posterior end of tooth row; variable presence of groove lingually below margo orbitalis; expression of labial ornament). The presence of more prominently expressed labial ornament in the smaller specimen is at odds with the general trend of cranial ornament (where present) becoming more pronounced with growth in anuran species. Determining whether these maxillae belong to separate species or simply are variants within a single species will require additional specimens. Even with this uncertainty, UALVP 40191 and 40192 are sufficiently distinct that they cannot be assigned to any of the species or other morphs recognized here. Nor are they closely comparable with other maxillary specimens known to us from Upper Cretaceous deposits elsewhere in the Western Interior.</paragraph>
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</subSubSection>
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</treatment>
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</document> |