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<document id="9A1E578F9083CB57AD9602FFD34B0527" ID-DOI="10.5852/ejt.2024.947.2633" ID-ISSN="2118-9773" ID-Zenodo-Dep="13365918" ID-ZooBank="D404A79E-E10C-4B42-817B-8A6A635A32AA" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1724418265566" checkinUser="plazi" docAuthor="Kim, Jeong-Kyu, Shimizu, Akira &amp; Pitts, James P." docDate="2024" docId="03ED87CD8D57FF84FDF9DCE3D2D5F97F" docLanguage="en" docName="EJT.2024.947.248-267.pdf" docOrigin="European Journal of Taxonomy 947" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2633/12117" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lissocnemis Kohl 1907" docType="treatment" docVersion="3" lastPageNumber="253" masterDocId="FFD4FFB58D54FF81FF99DE3BD01DFF9C" masterDocTitle="First record of the genus Lissocnemis Kohl, 1907 (Insecta: Hymenoptera: Pompilidae: Ctenocerinae) from Korea, with a new species and an unrecorded species" masterLastPageNumber="267" masterPageNumber="248" pageNumber="251" updateTime="1724960312330" updateUser="julia" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D8F618AFFEEA0C7F21936604932E2E1D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="78DDD9FE6A1AD4DD3947E43C3161EA4A">
<mods:title id="C43EE889158155A30F8B8326241AA316">First record of the genus Lissocnemis Kohl, 1907 (Insecta: Hymenoptera: Pompilidae: Ctenocerinae) from Korea, with a new species and an unrecorded species</mods:title>
</mods:titleInfo>
<mods:name id="BC1FDC23ACF8AD1DF2CFBB4B4907BCD1" type="personal">
<mods:role id="133FBC2A4A91A323D2BBAED83E59B538">
<mods:roleTerm id="854E0F09DC7AA38E10637B5CD1BB842E">Author</mods:roleTerm>
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<mods:namePart id="24B8337159A16EA55ABB1816DDDAEACA">Kim, Jeong-Kyu</mods:namePart>
<mods:affiliation id="41D73629147B8BAB4DF20EE932731CE1">Department of Bio Environment Health, Dongnam Health University, Suwon-si, Gyeonggi-do, 16328 Korea</mods:affiliation>
<mods:nameIdentifier id="4FB11F823E972BED21436A01DA4AB79F" type="URL">urn:lsid:zoobank.org:author:9202D5DB-0D26-4100-9D3F-C22CD0DA8D07</mods:nameIdentifier>
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<mods:role id="FE0A79E66FFEEAD572710A7845DE29B1">
<mods:roleTerm id="057D3D490665B55B2E9D5230AB09DEF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BACFA4F16331AE2B92217236414534DF">Shimizu, Akira</mods:namePart>
<mods:affiliation id="1B10CE34793D22B9829BDB63EFFCC5CC">Department of Biological Science, Tokyo Metropolitan University, Hachioji, Tokyo, 192 - 0397 Japan &amp; Research Institute of Evolutionary Biology, Inc., Setagaya-ku, Tokyo, 158 - 0098 Japan</mods:affiliation>
<mods:nameIdentifier id="D173CE68BF8678752B5E0772F9D4AED4" type="URL">urn:lsid:zoobank.org:author:F015CF83-2089-4DFF-AF69-535579E5307F</mods:nameIdentifier>
<mods:nameIdentifier id="B68980C3F5FA031CA272CA87E0B1C4EA" type="email">aquilashimizu7@gmail.com</mods:nameIdentifier>
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<mods:name id="5ACA8D6EE5614F9AB6E22405C6E24C0D" type="personal">
<mods:role id="93D4D71C124FAE26BB69FF279A04C35C">
<mods:roleTerm id="D4ED14B7AE48B86BC5A930179B602F06">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4ABBCF88D7F8AF0E76B1E6CC8D5A75D">Pitts, James P.</mods:namePart>
<mods:affiliation id="A2DD91F98ADE6B125CA0FA454097DA8A">Department of Biology, Utah State University, Logan, UT, 84322 - 5305 USA.</mods:affiliation>
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<mods:nameIdentifier id="110A93A963DD4165CE60B65C0F2A9826" type="email">james.pitts@usu.edu</mods:nameIdentifier>
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<mods:typeOfResource id="EEEB157A93E725514294DE9E76ADAF32">text</mods:typeOfResource>
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<mods:title id="73CBD7FC08D5DBF845FADB0A08E5AD64">European Journal of Taxonomy</mods:title>
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<mods:date id="0080BA6934A25BA84628A886FDA781FC">2024</mods:date>
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<mods:number id="587E6826031B77C58BCDCCB0645AEC7F">2024-08-23</mods:number>
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<mods:detail id="AE27A1124588CF413E5DB87A06119BF6" type="volume">
<mods:number id="31DFFA6FA91E598E3ABC5B9F78172002">947</mods:number>
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<mods:start id="0492C927AF6559978E88E792CB713C09">248</mods:start>
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<mods:url id="0967FFE725F2BEFD9FE9013FDD08A2FE">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2633/12117</mods:url>
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<treatment id="03ED87CD8D57FF84FDF9DCE3D2D5F97F" LSID="urn:lsid:plazi:treatment:03ED87CD8D57FF84FDF9DCE3D2D5F97F" httpUri="http://treatment.plazi.org/id/03ED87CD8D57FF84FDF9DCE3D2D5F97F" lastPageId="5" lastPageNumber="253" pageId="3" pageNumber="251">
<subSubSection id="C35E65508D57FF82FDF9DCE3D3C9FD6E" box="[608,980,728,754]" pageId="3" pageNumber="251" type="nomenclature">
<paragraph id="8BFB36DB8D57FF82FDF9DCE3D3C9FD6E" blockId="3.[608,980,728,754]" box="[608,980,728,754]" pageId="3" pageNumber="251">
<heading id="D0B381B78D57FF82FDF9DCE3D3C9FD6E" box="[608,980,728,754]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="251" reason="2">
Genus
<taxonomicName id="4C444D588D57FF82FD2BDCE3D3C9FD6E" ID-CoL="5FFH" ID-ENA="2066996" authority="Kohl, 1907" authorityName="Kohl" authorityYear="1907" box="[690,980,728,754]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D57FF82FD2BDCE3D355FD6E" bold="true" box="[690,840,728,754]" italics="true" pageId="3" pageNumber="251">Lissocnemis</emphasis>
<bibRefCitation id="EFD54B2A8D57FF82FCC9DCE3D3C9FD6E" author="Kohl F. F." box="[848,980,728,754]" pageId="3" pageNumber="251" pagination="169 - 279" refId="ref11340" refString="Kohl F. F. 1907 [1906]. Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 71 (1): 169 - 279." type="book chapter" year="1907">Kohl, 1907</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D57FF82FF24DD21D192FC04" pageId="3" pageNumber="251" type="reference_group">
<paragraph id="8BFB36DB8D57FF82FF24DD21D200FCA8" blockId="3.[189,541,793,820]" box="[189,541,793,820]" pageId="3" pageNumber="251">
<treatmentCitationGroup id="AB5411F58D57FF82FF24DD21D200FCA8" box="[189,541,793,820]" pageId="3" pageNumber="251">
<treatmentCitation id="0AE510CA8D57FF82FF24DD21D204FCA8" author="Kohl F. F." box="[189,537,793,820]" page="265" pageId="3" pageNumber="251" year="1907">
<taxonomicName id="4C444D588D57FF82FF24DD21D204FCA8" ID-CoL="5FFH" authority="Kohl, 1907: 265" authorityName="Kohl" authorityPageNumber="265" authorityYear="1907" box="[189,537,793,820]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D57FF82FF24DD21D153FCAF" box="[189,334,794,819]" italics="true" pageId="3" pageNumber="251">Lissocnemis</emphasis>
<bibRefCitation id="EFD54B2A8D57FF82FECCDD22D204FCA8" author="Kohl F. F." box="[341,537,793,820]" pageId="3" pageNumber="251" pagination="169 - 279" refId="ref11340" refString="Kohl F. F. 1907 [1906]. Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 71 (1): 169 - 279." type="book chapter" year="1907">Kohl, 1907: 265</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24DD60D192FC04" blockId="3.[189,1399,859,920]" pageId="3" pageNumber="251">
<treatmentCitationGroup id="AB5411F58D57FF82FF24DD60D192FC04" pageId="3" pageNumber="251">
<taxonomicName id="4C444D588D57FF82FF24DD60D148FCE9" ID-CoL="3WSN" authorityName="Turner" authorityYear="1917" box="[189,341,859,885]" class="Insecta" family="Pompilidae" genus="Cryptosalius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D57FF82FF24DD60D148FCE9" box="[189,341,859,885]" italics="true" pageId="3" pageNumber="251">Cryptosalius</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AE510CA8D57FF82FEE8DD67D22FFCE9" author="Turner R. E." box="[369,562,859,886]" page="76" pageId="3" pageNumber="251" year="1917">
<bibRefCitation id="EFD54B2A8D57FF82FEE8DD67D22FFCE9" author="Turner R. E." box="[369,562,859,886]" pageId="3" pageNumber="251" pagination="53 - 84" refId="ref11661" refString="Turner R. E. 1917. New species of Hymenoptera in the British Museum. Transactions of the Entomological Society of London 1917: 53 - 84. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01402. x" type="journal article" year="1917">Turner 1917: 76</bibRefCitation>
</treatmentCitation>
. [In part:
<taxonomicName id="4C444D588D57FF82FD3ADD60D3F4FCE9" authority="Turner, 1920" authorityName="Turner" authorityYear="1920" box="[675,1001,859,886]" class="Insecta" family="Pompilidae" genus="Cryptosalius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B930EAC98D57FF82FD3ADD60D356FCE9" box="[675,843,859,885]" italics="true" pageId="3" pageNumber="251">C. tonkinensis</emphasis>
<bibRefCitation id="EFD54B2A8D57FF82FCCBDD67D3F4FCE9" author="Turner R. E." box="[850,1001,859,886]" pageId="3" pageNumber="251" pagination="84 - 98" refId="ref11709" refString="Turner R. E. 1920. On Indo-Chinese Hymenoptera collected by R. Vitalis de Salvaza. IV. Annals and Magazine of Natural History (9) 5: 84 - 98. https: // doi. org / 10.1080 / 00222932008632344" type="journal article" year="1920">Turner, 1920</bibRefCitation>
</taxonomicName>
is transferred to
<taxonomicName id="4C444D588D57FF82FB36DD67D55DFCE9" ID-CoL="5FFH" authorityName="Kohl" authorityYear="1907" box="[1199,1344,860,885]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D57FF82FB36DD67D55DFCE9" box="[1199,1344,860,885]" italics="true" pageId="3" pageNumber="251">Lissocnemis</emphasis>
</taxonomicName>
(see the remarks).]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D57FF82FF24DDFBD469FB94" pageId="3" pageNumber="251" type="type_taxon">
<paragraph id="8BFB36DB8D57FF82FF24DDFBD144FC45" blockId="3.[189,345,959,986]" box="[189,345,959,986]" pageId="3" pageNumber="251">
<heading id="D0B381B78D57FF82FF24DDFBD144FC45" bold="true" box="[189,345,959,986]" fontSize="11" level="3" pageId="3" pageNumber="251" reason="3">
<emphasis id="B930EAC98D57FF82FF24DDFBD144FC45" bold="true" box="[189,345,959,986]" pageId="3" pageNumber="251">
<typeStatus id="54FF88798D57FF82FF24DDFBD0E6FC46" box="[189,251,960,986]" pageId="3" pageNumber="251">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24DDD5D469FB94" blockId="3.[189,1140,1006,1032]" box="[189,1140,1006,1032]" pageId="3" pageNumber="251">
<taxonomicName id="4C444D588D57FF82FF24DDD5D286FB94" ID-CoL="6XCRN" authority="Kohl, 1907" authorityName="Kohl" authorityYear="1907" box="[189,667,1006,1032]" class="Insecta" family="Pompilidae" genus="Salius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="species" species="irrasus" subGenus="Lissocnemis">
<emphasis id="B930EAC98D57FF82FF24DDD5D118FB94" box="[189,261,1006,1032]" italics="true" pageId="3" pageNumber="251">Salius</emphasis>
(
<emphasis id="B930EAC98D57FF82FE8EDDD4D1B5FB94" box="[279,424,1007,1032]" italics="true" pageId="3" pageNumber="251">Lissocnemis</emphasis>
)
<emphasis id="B930EAC98D57FF82FE20DDD4D210FB94" box="[441,525,1007,1032]" italics="true" pageId="3" pageNumber="251">irrasus</emphasis>
<bibRefCitation id="EFD54B2A8D57FF82FD8DDDD5D286FB94" author="Kohl F. F." box="[532,667,1006,1032]" pageId="3" pageNumber="251" pagination="169 - 279" refId="ref11340" refString="Kohl F. F. 1907 [1906]. Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 71 (1): 169 - 279." type="book chapter" year="1907">Kohl, 1907</bibRefCitation>
</taxonomicName>
, by original designation and monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D57FF82FF24DA14D13FFB07" pageId="3" pageNumber="251" type="diagnosis">
<paragraph id="8BFB36DB8D57FF82FF24DA14D12AFBD5" blockId="3.[189,311,1071,1097]" box="[189,311,1071,1097]" pageId="3" pageNumber="251">
<heading id="D0B381B78D57FF82FF24DA14D12AFBD5" bold="true" box="[189,311,1071,1097]" fontSize="11" level="3" pageId="3" pageNumber="251" reason="3">
<emphasis id="B930EAC98D57FF82FF24DA14D12AFBD5" bold="true" box="[189,311,1071,1097]" pageId="3" pageNumber="251">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24DA64D13FFB07" blockId="3.[189,1398,1118,1179]" pageId="3" pageNumber="251">
<taxonomicName id="4C444D588D57FF82FF24DA64D153FBE4" authorityName="Kohl" authorityYear="1907" box="[189,334,1119,1144]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D57FF82FF24DA64D153FBE4" box="[189,334,1119,1144]" italics="true" pageId="3" pageNumber="251">Lissocnemis</emphasis>
</taxonomicName>
is distinguished from other genera of
<taxonomicName id="4C444D588D57FF82FC86DA65D3A7FBE4" authorityName="Arnold" authorityYear="1934" box="[799,954,1118,1144]" class="Insecta" family="Pompilidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="subFamily" subFamily="Ctenocerinae">Ctenocerinae</taxonomicName>
by the combination of the following features.
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D57FF84FF73DAF8D276FC12" lastPageId="5" lastPageNumber="253" pageId="3" pageNumber="251" type="description">
<paragraph id="8BFB36DB8D57FF82FF73DAF8D15BFB41" blockId="3.[188,1400,1219,1597]" box="[234,326,1219,1245]" pageId="3" pageNumber="251">
<emphasis id="B930EAC98D57FF82FF73DAF8D15BFB41" bold="true" box="[234,326,1219,1245]" pageId="3" pageNumber="251">Female</emphasis>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24DADDD1A8F9A1" blockId="3.[188,1400,1219,1597]" pageId="3" pageNumber="251">
Supra-antennal area of frons produced anteriorly (
<figureCitation id="137F2A5E8D57FF82FCBDDADDD39FFA9C" box="[804,898,1254,1280]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3F</figureCitation>
); pronotum elongate but not longer than mesoscutum at midline (
<figureCitation id="137F2A5E8D57FF82FE7ADB32D25AFABF" box="[483,583,1289,1315]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Figs 3G</figureCitation>
,
<figureCitation id="137F2A5E8D57FF82FDCFDB32D266FABF" box="[598,635,1289,1315]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">4A</figureCitation>
) with collar deeply depressed below level of dorsum (
<figureCitation id="137F2A5E8D57FF82FA95DB32D575FABF" box="[1292,1384,1289,1315]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3F</figureCitation>
); propodeum strongly and transversely rugulose at least posteriorly (
<figureCitation id="137F2A5E8D57FF82FC73DB17D452FADB" box="[1002,1103,1324,1351]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Figs 3H</figureCitation>
,
<figureCitation id="137F2A5E8D57FF82FBC6DB16D499FADB" box="[1119,1156,1325,1351]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">4C</figureCitation>
), its declivity rather flattened; hind tibia with minute sparse spines, lacking scale-like integumental projections (teeth) dorsally; apical spines of hind tibia short, of similar length, not splayed out; lateral margin of metasoma, in dorsal view, not very convex (
<figureCitation id="137F2A5E8D57FF82FDCBDBADD2B1FA2C" box="[594,684,1430,1456]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Figs 3I</figureCitation>
,
<figureCitation id="137F2A5E8D57FF82FD24DBADD2F8FA2C" box="[701,741,1430,1456]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">4A</figureCitation>
); T1 petiolate basally (segment narrower immediately behind articulation than its width at articulation itself;
<figureCitation id="137F2A5E8D57FF82FCC5DB82D3A0FA4F" box="[860,957,1465,1491]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3G</figureCitation>
); S2 with distinct transverse groove (
<figureCitation id="137F2A5E8D57FF82FF5FDBE7D103FA6B" box="[198,286,1500,1527]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3J</figureCitation>
); T6 with long, dense, curled pubescence (
<figureCitation id="137F2A5E8D57FF82FCABDBE7D38EFA6B" box="[818,915,1500,1527]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3K</figureCitation>
) and short, sparse, fine setae; all tarsal claws bifid, inner ray thicker than outer ray (
<figureCitation id="137F2A5E8D57FF82FD50DBC4D338F986" box="[713,805,1535,1562]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="3" pageNumber="251">Fig. 3E</figureCitation>
); and FW discal cell1 usually with oval, clear spot subbasally (
<figureCitation id="137F2A5E8D57FF82FED0D818D1B5F9A1" box="[329,424,1571,1597]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF73D85FD137F9E2" blockId="3.[188,1399,1636,1839]" box="[234,298,1636,1662]" pageId="3" pageNumber="251">
<emphasis id="B930EAC98D57FF82FF73D85FD137F9E2" bold="true" box="[234,298,1636,1662]" pageId="3" pageNumber="251">Male</emphasis>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24D8BCD2A9F8B2" blockId="3.[188,1399,1636,1839]" pageId="3" pageNumber="251">
Head in frontal view, rhomboid (
<figureCitation id="137F2A5E8D57FF82FDC2D8BCD2A2F93E" box="[603,703,1671,1698]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="3" pageNumber="251">Figs 1C</figureCitation>
,
<figureCitation id="137F2A5E8D57FF82FD4AD8B3D2EAF93E" box="[723,759,1672,1698]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">4E</figureCitation>
); upper frons broader than lower frons; scape with longitudinal sharp carina ventromedially (
<figureCitation id="137F2A5E8D57FF82FD3FD890D31DF959" box="[678,768,1707,1733]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">Fig. 4E</figureCitation>
); flagellomeres beneath usually arcuately or angularly produced subbasally (
<figureCitation id="137F2A5E8D57FF82FE25D8F5D207F974" box="[444,538,1742,1768]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="3" pageNumber="251">Fig. 4H</figureCitation>
); propodeum densely punctate and minutely reticulate rugulose (
<figureCitation id="137F2A5E8D57FF82FA92D8F5D57AF974" box="[1291,1383,1742,1768]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="3" pageNumber="251">Fig. 1H</figureCitation>
); fore and mid tarsal claws bifid as in female; hind tarsal claws edentate, acutely bent subapically (
<figureCitation id="137F2A5E8D57FF82FA96D8CAD57AF897" box="[1295,1383,1777,1803]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="3" pageNumber="251">Fig. 1E</figureCitation>
), pair of claws not divergent but subparallel.
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24D96ED170F8F3" blockId="3.[189,365,1877,1903]" box="[189,365,1877,1903]" pageId="3" pageNumber="251">
<heading id="D0B381B78D57FF82FF24D96ED170F8F3" bold="true" box="[189,365,1877,1903]" fontSize="11" level="3" pageId="3" pageNumber="251" reason="3">
<emphasis id="B930EAC98D57FF82FF24D96ED170F8F3" bold="true" box="[189,365,1877,1903]" pageId="3" pageNumber="251">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DB8D57FF82FF24D9BED3D8F85E" blockId="3.[189,1398,1924,1986]" pageId="3" pageNumber="251">
We redescribe the genus based on the above four species as a complement to
<bibRefCitation id="EFD54B2A8D57FF82FBDBD9BFD4FBF802" author="Kohl F. F." box="[1090,1254,1924,1950]" pageId="3" pageNumber="251" pagination="169 - 279" refId="ref11340" refString="Kohl F. F. 1907 [1906]. Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 71 (1): 169 - 279." type="book chapter" year="1907">Kohl’s (1907)</bibRefCitation>
description, which was based only on the female of the
<typeStatus id="54FF88798D57FF82FD21D993D2F6F85E" box="[696,747,1960,1986]" pageId="3" pageNumber="251">type</typeStatus>
species,
<taxonomicName id="4C444D588D57FF82FCCFD993D3A0F85D" authorityName="Kohl" authorityYear="1907" box="[854,957,1960,1985]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="251" phylum="Arthropoda" rank="species" species="irrasa">
<emphasis id="B930EAC98D57FF82FCCFD993D3A0F85D" box="[854,957,1960,1985]" italics="true" pageId="3" pageNumber="251">L. irrasa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF73DF32D15BFEBF" blockId="4.[189,978,265,326]" box="[234,326,265,291]" pageId="4" pageNumber="252">
<emphasis id="B930EAC98D50FF85FF73DF32D15BFEBF" bold="true" box="[234,326,265,291]" pageId="4" pageNumber="252">Female</emphasis>
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DF17D3CFFEDA" blockId="4.[189,978,265,326]" box="[189,978,300,326]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DF17D173FED8" baselines="319,319" box="[189,366,300,326]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="4" pageNumber="252">MEASUREMENTS</smallCapsWord>
. Medium-sized wasps,
<quantity id="4CBC9B3E8D50FF85FD1DDF17D307FEDA" box="[644,794,300,326]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="4" pageNumber="252" unit="mm" value="16.0" valueMax="20.0" valueMin="12.0">12 to 20 mm</quantity>
in body length.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DF55D2C1FD58" blockId="4.[188,1401,365,708]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DF55D0E1FE1A" baselines="385,385" box="[189,252,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="4" pageNumber="252">HEAD</smallCapsWord>
. Broader than high. Vertex moderately convex above level of eye tops (
<figureCitation id="137F2A5E8D50FF85FBC3DF56D4A4FE1B" box="[1114,1209,365,391]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3D</figureCitation>
); juncture of its anterior and posterior faces broadly rounded (
<figureCitation id="137F2A5E8D50FF85FD46DFABD327FE36" box="[735,826,400,426]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3F</figureCitation>
). Ocelli forming right- to obtuse-angled triangle (
<figureCitation id="137F2A5E8D50FF85FF5EDF88D135FE52" box="[199,296,435,462]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3C</figureCitation>
). Supra-antennal area of frons, in dorsal view, triangular, trapezoidal, or arcuate. Clypeus short and transverse, as broad as or broader than LID (
<figureCitation id="137F2A5E8D50FF85FCA6DFECD3BCFE6D" box="[831,929,471,497]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Figs 3D</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FC37DFECD3C9FE6D" box="[942,980,471,497]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4B</figureCitation>
), slightly convex medially. Labrum slightly narrower than lower margin of clypeus (
<figureCitation id="137F2A5E8D50FF85FD76DFC1D356FD88" box="[751,843,506,532]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3D</figureCitation>
), moderately to fairly exposed beneath clypeus. Antennal socket separated from frontoclypeal suture by much less than half of its own diameter. Scape almost straight, or slightly curved outward with lateral face slightly concave. Malar space very short (
<figureCitation id="137F2A5E8D50FF85FF51DC58D13DFDE2" box="[200,288,611,638]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3F</figureCitation>
). Mandible short and stout with strong tooth subapically on inner margin. Maxillary palpomeres 4–6 not much longer than palpomere 3. Gena, in dorsal view, rather strongly receding posteriorly but not very thin (
<figureCitation id="137F2A5E8D50FF85FEACDC91D18CFD58" box="[309,401,682,708]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3C</figureCitation>
). Occipital suture complete.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DCD7D39DFB83" blockId="4.[188,1400,747,1055]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DCD7D127FC98" baselines="767,767" box="[189,314,748,774]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesosoma" pageId="4" pageNumber="252">MESOSOMA</smallCapsWord>
. Pronotum with streptaulus complete; declivity nearly vertical and flattened (
<figureCitation id="137F2A5E8D50FF85FB7FDCD0D55CFC99" box="[1254,1345,747,773]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3F</figureCitation>
), its juncture with dorsum narrowly rounded; dorsum truncate anteriorly (
<figureCitation id="137F2A5E8D50FF85FC6FDD35D44BFCB5" box="[1014,1110,782,809]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3G</figureCitation>
), rather flattened above, its lateral margin, in dorsal view, barely convex posteriorly, gradually narrowing anteriorly; juncture of dorsum and lateral face narrowly raised or carinate; lateral face vertical and flattened. Mesoscutum narrowly raised posterolaterally; parapsidal sulci divergent anteriorly. Scutellum slightly raised above level of mesoscutum (
<figureCitation id="137F2A5E8D50FF85FE27DDA0D20BFC29" box="[446,534,923,949]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3F</figureCitation>
). Metapostnotum shorter than half of metanotum at midline (
<figureCitation id="137F2A5E8D50FF85FB40DDA0D524FC29" box="[1241,1337,923,949]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Figs 3H</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FADCDDA0D576FC29" box="[1349,1387,923,949]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4C</figureCitation>
). Propodeum with dorsum transversely convex, transversely rugulose or reticulate-rugulae (
<figureCitation id="137F2A5E8D50FF85FB4FDD85D52BFC45" box="[1238,1334,958,985]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Figs 3H</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FADBDD84D57AFC45" box="[1346,1383,959,985]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4C</figureCitation>
), spiracles distanced from anterior margin of propodeum by more than its own length; declivity rather flattened (
<figureCitation id="137F2A5E8D50FF85FEADDA3ED196FB83" box="[308,395,1029,1055]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3F</figureCitation>
) but not distinctly delimited from dorsum.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DA7CD18DFB56" blockId="4.[189,1399,1094,1227]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DA7CD114FBC3" baselines="1114,1114" box="[189,265,1095,1121]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wings" pageId="4" pageNumber="252">WINGS</smallCapsWord>
. FW with three SMCs (
<figureCitation id="137F2A5E8D50FF85FD82DA7DD266FBFD" box="[539,635,1094,1121]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Figs 3A</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FD1FDA7CD2B3FBFD" box="[646,686,1095,1121]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4A</figureCitation>
). Pterostigma long, its posterior base much longer than cross-vein 2
<emphasis id="B930EAC98D50FF85FE9CDA51D12CFB1F" box="[261,305,1130,1155]" italics="true" pageId="4" pageNumber="252">r-rs</emphasis>
. Marginal cell lanceolate, acute at apex. Second abscissa of vein M (lower part of basal vein) strongly curved. Crossvein 2
<emphasis id="B930EAC98D50FF85FD88DAB5D25AFB3B" box="[529,583,1166,1191]" italics="true" pageId="4" pageNumber="252">rs-m</emphasis>
straight. Cross-vein
<emphasis id="B930EAC98D50FF85FCA6DAB5D369FB3B" box="[831,884,1166,1191]" italics="true" pageId="4" pageNumber="252">cu-a</emphasis>
originating distally to fork of vein
<emphasis id="B930EAC98D50FF85FA8FDAB6D56AFB3B" box="[1302,1399,1165,1191]" italics="true" pageId="4" pageNumber="252">M+CuA</emphasis>
by its own length.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DAC9D4C6FA27" blockId="4.[188,1398,1265,1468]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DAC9D0EBFA96" baselines="1285,1285" box="[189,246,1266,1292]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="4" pageNumber="252">LEGS</smallCapsWord>
. Fore tarsomere 1 longer than tarsomeres 2–4 combined. Fore femur not swollen, almost as thick as mid femur. Fore tibia lacking stout, decurved spines apicomesially. Both hind coxae separate each other, unlike mid coxae (
<figureCitation id="137F2A5E8D50FF85FE0BDB03D1EDFACE" box="[402,496,1336,1362]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">Fig. 4D</figureCitation>
). Basal ring on mid and hind femora distinct. Mid and hind tibiae with several short spines laterally and dorsally. Orbiculae small (
<figureCitation id="137F2A5E8D50FF85FC8DDB60D370FAE9" box="[788,877,1371,1397]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3E</figureCitation>
), its width ca 0.5× as wide as tarsomere 5; its pecten consisting of fine, straight, divergent setulae, these being much longer than orbiculae themselves. Hind tarsomere 5 with one or a few irregularly arranged spines or without spines beneath.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24DBD8D47BF9BC" blockId="4.[189,1398,1507,1569]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24DBD8D124FA67" baselines="1526,1526" box="[189,313,1507,1533]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metasoma" pageId="4" pageNumber="252">METASOMA</smallCapsWord>
. Rather slender, its width about 1.2 × as wide as mesosoma between tegulae, in dorsal view, not very convex medially. S6 moderately compressed laterally or gently arched.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF73D873D137F9FE" blockId="4.[189,1116,1608,1669]" box="[234,298,1608,1634]" pageId="4" pageNumber="252">
<emphasis id="B930EAC98D50FF85FF73D873D137F9FE" bold="true" box="[234,298,1608,1634]" pageId="4" pageNumber="252">Male</emphasis>
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24D850D441F919" blockId="4.[189,1116,1608,1669]" box="[189,1116,1643,1669]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24D850D173F91F" baselines="1662,1662" box="[189,366,1643,1669]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="4" pageNumber="252">MEASUREMENTS</smallCapsWord>
. Smaller and slenderer than female,
<quantity id="4CBC9B3E8D50FF85FC84D850D3BEF919" box="[797,931,1643,1669]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="4" pageNumber="252" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8 to 13 mm</quantity>
in body length.
</paragraph>
<paragraph id="8BFB36DB8D50FF85FF24D896D4A5F821" blockId="4.[189,1399,1708,1981]" pageId="4" pageNumber="252">
<smallCapsWord id="8D1DA0078D50FF85FF24D896D0E1F959" baselines="1728,1728" box="[189,252,1709,1735]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="4" pageNumber="252">HEAD</smallCapsWord>
. Upper frons, vertex, and gena with numerous long erect setae (
<figureCitation id="137F2A5E8D50FF85FB89D897D485F95A" box="[1040,1176,1708,1734]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Figs 1C–D</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FB3ED897D4D1F95A" box="[1191,1228,1708,1734]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4E</figureCitation>
). Vertex more strongly convex above level of eye tops than in female (
<figureCitation id="137F2A5E8D50FF85FCFED8F4D3DAF975" box="[871,967,1743,1769]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Figs 1C</figureCitation>
vs 3D, 4E vs 4B), chevron-shaped. Inner orbits more or less divergent above. Ocelli forming right- or obtuse-angled triangle (
<figureCitation id="137F2A5E8D50FF85FB7FD8C9D575F891" box="[1254,1384,1778,1805]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Fig. 1D, G</figureCitation>
), anterior ocellus greater than posterior ocelli (
<figureCitation id="137F2A5E8D50FF85FD49D92DD32CF8AC" box="[720,817,1814,1840]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Figs 1D</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FCA6D92DD379F8AC" box="[831,868,1814,1840]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4E</figureCitation>
). Supra-antennal ledge as in female. Clypeus almost truncate apically (
<figureCitation id="137F2A5E8D50FF85FE7ED902D25AF8CF" box="[487,583,1849,1875]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Figs 1C</figureCitation>
,
<figureCitation id="137F2A5E8D50FF85FDCCD902D267F8CF" box="[597,634,1849,1875]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="4" pageNumber="252">4E</figureCitation>
). Antennal socket more separated from frontoclypeal suture than in female. Scape longer than Fl1 (
<figureCitation id="137F2A5E8D50FF85FDD3D967D2B8F8EA" box="[586,677,1884,1910]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Fig. 1B</figureCitation>
), straight or slightly curved outward, produced apicomesially (
<figureCitation id="137F2A5E8D50FF85FF5FD944D13CF805" box="[198,289,1919,1945]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Fig. 1D</figureCitation>
), densely setose lateroventrally. Mandible short with strong tooth subapically on inner margin. Gena more gently receding posteriorly and thicker than in female (
<figureCitation id="137F2A5E8D50FF85FC52D999D43BF821" box="[971,1062,1954,1981]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="4" pageNumber="252">Fig. 1D</figureCitation>
vs
<figureCitation id="137F2A5E8D50FF85FBD6D999D4B7F821" box="[1103,1194,1954,1981]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[246,1339,271,1816]" captionTargetId="figure-17@9.[246,1342,265,1820]" captionTargetPageId="9" captionText="Fig. 3.Lissocnemis koreana Kim &amp; Shimizu sp. nov., paratype, ♀ (DHU), from Mt Gayasan, Daegok-ri, Haemi-myeon, Seosan-si, CN.A. Whole body, lateral view. B. Antenna.C. Head, dorsal view.D. Head, frontal view. E. Hind pretarsus, posterior view. F. Head and mesosoma, lateral view. G. Mesosoma, dorsal view. H. Posterior part of mesosoma, dorsal view. I. Metasoma, dorsal view. J. Metasoma, ventral view. K. Apical part of metasoma, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365924" httpUri="https://zenodo.org/record/13365924/files/figure.png" pageId="4" pageNumber="252">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DF32D553FE2C" blockId="5.[189,1400,265,432]" pageId="5" pageNumber="253">
<smallCapsWord id="8D1DA0078D51FF84FF24DF32D127FEBD" baselines="284,284" box="[189,314,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesosoma" pageId="5" pageNumber="253">MESOSOMA</smallCapsWord>
. Pronotum with declivity vertical, flattened or concave (
<figureCitation id="137F2A5E8D51FF84FC68DF32D44BFEBF" box="[1009,1110,265,291]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Figs 1A</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FBFEDF32D493FEBF" box="[1127,1166,265,291]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">4H</figureCitation>
), polished; dorsum declivous, gradually narrowing anteriorly (
<figureCitation id="137F2A5E8D51FF84FD4BDF17D326FEDA" box="[722,827,300,326]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Figs 1G</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FCD6DF17D376FEDA" box="[847,875,300,326]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">4I</figureCitation>
), with long, dense, erect pubescence and setae. Disc of scutellum triangular (
<figureCitation id="137F2A5E8D51FF84FDFCDF74D2D5FEF5" box="[613,712,335,361]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Figs 1G</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FD4CDF74D2ECFEF5" box="[725,753,335,361]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">4I</figureCitation>
), more strongly raised than in female. Metapostnotum longer than in female (
<figureCitation id="137F2A5E8D51FF84FE57DF49D22DFE11" box="[462,560,370,397]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Figs 1H</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FDA7DF48D247FE11" box="[574,602,371,397]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">4I</figureCitation>
). Propodeum with pubescence and setae, both very long and dense posterolaterally (
<figureCitation id="137F2A5E8D51FF84FE1CDFADD1FFFE2C" box="[389,482,406,432]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Fig. 1H</figureCitation>
); dorsum longer than declivity (
<figureCitation id="137F2A5E8D51FF84FCC2DFADD3A5FE2C" box="[859,952,406,432]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Fig. 1A</figureCitation>
), gradually merging into declivity.
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DFE3D24EFDA4" blockId="5.[189,1399,471,568]" pageId="5" pageNumber="253">
<smallCapsWord id="8D1DA0078D51FF84FF24DFE3D114FE6C" baselines="491,491" box="[189,265,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wings" pageId="5" pageNumber="253">WINGS</smallCapsWord>
. FW discal cell 1 almost lacking clear spot subbasally. Crossvein cu-a originating distally to separation of vein M +Cu by less than its own length (
<figureCitation id="137F2A5E8D51FF84FCD9DFC1D386FD89" box="[832,923,506,533]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Figs 1F</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FC31DFC0D3CDFD89" box="[936,976,507,533]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">4H</figureCitation>
). HW cross-vein cu-a originating at or distally to fork of vein
<emphasis id="B930EAC98D51FF84FE73DC25D251FDA4" box="[490,588,542,568]" italics="true" pageId="5" pageNumber="253">M+CuA</emphasis>
.
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DC5BD35FFD00" blockId="5.[188,1399,607,668]" pageId="5" pageNumber="253">
<smallCapsWord id="8D1DA0078D51FF84FF24DC5BD0EBFDE4" baselines="627,627" box="[189,246,608,634]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="5" pageNumber="253">LEGS</smallCapsWord>
. Pair of hind coxae closely set unlike in female. Hind tarsomeres narrowing apically. Fore and mid orbiculae smaller than in female; hind orbicula minute.
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DCFFD276FC12" blockId="5.[188,1400,708,910]" pageId="5" pageNumber="253">
<smallCapsWord id="8D1DA0078D51FF84FF24DCFFD124FD40" baselines="727,727" box="[189,313,708,734]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metasoma" pageId="5" pageNumber="253">METASOMA</smallCapsWord>
. Slender (
<figureCitation id="137F2A5E8D51FF84FE28DCFFD213FD42" box="[433,526,708,734]" captionStart="Fig" captionStartId="13.[191,234,1892,1918]" captionTargetBox="[224,1374,271,1848]" captionTargetId="figure-91@13.[212,1376,265,1855]" captionTargetPageId="13" captionText="Fig. 4. Lissocnemis brevipennis (Cameron, 1902). A–D. ♀ (UMUT). E–I. ♂ (DHU). A, G. Whole body, dorsal view. B. Head and antennae, frontal view. C. Posterior part of mesosoma, dorsal view. D. Mid and hind coxae, ventral view. E. Head, frontal view. F. T1, lateral view. H. Whole body, lateral view. I. Mesosoma, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365926" httpUri="https://zenodo.org/record/13365926/files/figure.png" pageId="5" pageNumber="253">Fig. 4G</figureCitation>
), its width almost equal to or lesser than mesosoma width between tegulae. T1 gradually narrowing (
<figureCitation id="137F2A5E8D51FF84FE6CDCDCD248FC9D" box="[501,597,743,769]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[238,1346,279,1785]" captionTargetId="figure-11@6.[238,1349,265,1785]" captionTargetPageId="6" captionText="Fig. 1. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–J. Holotype, ♂ (NIBR). K. Paratype, ♂ (DHU) from Gajwa-dong, Jinju-si, GN, Korea. A. Whole body, lateral view. B. Antenna. C. Head, frontal view. D. Head, dorsal view. E. Hind tarsal claw, lateral view. F. Fore and hind wings. G. Head and thorax, dorsal view. H. Posterior part of mesosoma and T1, dorsal view. I. Metasoma, dorsal view. J. T1, lateral view. K. S3–S4, ventral view (red arrows: a pair of linear oblique tubercles on S4). Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365920" httpUri="https://zenodo.org/record/13365920/files/figure.png" pageId="5" pageNumber="253">Fig. 1H</figureCitation>
) or shortly parallel-sided basally. Subgenital plate parallel-sided basally (
<figureCitation id="137F2A5E8D51FF84FE86DD31D161FCB8" box="[287,380,778,804]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[246,1359,1379,1854]" captionTargetId="figure-394@8.[224,1360,1377,1854]" captionTargetPageId="8" captionText="Fig. 2. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–C. Holotype (NIBR). D. Paratype (DHU) from Godang-ri, Unju-myeon, Wanju-gun, JB, Korea, subgenital plate and genitalia. A. Subgenital plate, ventral view. B. Genitalia, dorsal view. C. Paramere and digitus volsellaris, lateral view. D. T7 and genitalia, dorsal view. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365922" httpUri="https://zenodo.org/record/13365922/files/figure.png" pageId="5" pageNumber="253">Fig. 2A</figureCitation>
), gradually narrowing posteriorly, truncate or rounded apically. Genitalia with paramere short (
<figureCitation id="137F2A5E8D51FF84FE91DD16D177FCDB" box="[264,362,813,839]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[246,1359,1379,1854]" captionTargetId="figure-394@8.[224,1360,1377,1854]" captionTargetPageId="8" captionText="Fig. 2. Lissocnemis koreana Kim &amp; Shimizu sp. nov., ♂. A–C. Holotype (NIBR). D. Paratype (DHU) from Godang-ri, Unju-myeon, Wanju-gun, JB, Korea, subgenital plate and genitalia. A. Subgenital plate, ventral view. B. Genitalia, dorsal view. C. Paramere and digitus volsellaris, lateral view. D. T7 and genitalia, dorsal view. Scale bars=1 mm." figureDoi="http://doi.org/10.5281/zenodo.13365922" httpUri="https://zenodo.org/record/13365922/files/figure.png" pageId="5" pageNumber="253">Figs 2B</figureCitation>
,
<figureCitation id="137F2A5E8D51FF84FEE1DD16D181FCD4" box="[376,412,813,840]" captionStart="Fig" captionStartId="16.[189,232,1963,1989]" captionTargetBox="[200,1396,1263,1921]" captionTargetId="figure-381@16.[189,1398,1262,1924]" captionTargetPageId="16" captionText="Fig. 5. Lissocnemis brevipennis(Cameron, 1902), ♂ (DHU), subgenital plate and genitalia. A. Subgenital plate, ventral view. B. Genitalia, dorsal view. Scale bars =1mm." figureDoi="http://doi.org/10.5281/zenodo.13365928" httpUri="https://zenodo.org/record/13365928/files/figure.png" pageId="5" pageNumber="253">5B</figureCitation>
), wedge-shaped apically, not exceeding beyond apex of digitus volsellaris; digitus volsellaris broadened apically, club-shaped; basal hooklet single; parapenial lobe variable in shape, not exceeding beyond apex of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D51FF84FF24DD8ED332FBBD" pageId="5" pageNumber="253" type="distribution">
<paragraph id="8BFB36DB8D51FF84FF24DD8ED145FC53" blockId="5.[189,344,949,975]" box="[189,344,949,975]" pageId="5" pageNumber="253">
<heading id="D0B381B78D51FF84FF24DD8ED145FC53" bold="true" box="[189,344,949,975]" fontSize="11" level="3" pageId="5" pageNumber="253" reason="3">
<emphasis id="B930EAC98D51FF84FF24DD8ED145FC53" bold="true" box="[189,344,949,975]" pageId="5" pageNumber="253">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DDDFD332FBBD" blockId="5.[189,1399,996,1057]" pageId="5" pageNumber="253">
South Asia (
<collectingCountry id="F353764B8D51FF84FECFDDDFD185FC62" box="[342,408,996,1022]" name="India" pageId="5" pageNumber="253">India</collectingCountry>
), Southeast Asia (
<collectingCountry id="F353764B8D51FF84FDE1DDDFD2E9FC62" box="[632,756,996,1022]" name="Singapore" pageId="5" pageNumber="253">Singapore</collectingCountry>
,
<collectingCountry id="F353764B8D51FF84FC9DDDDFD377FC62" box="[772,874,996,1022]" name="Vietnam" pageId="5" pageNumber="253">Vietnam</collectingCountry>
,
<taxonomicName id="4C444D588D51FF84FCE3DDDFD406FC62" authority=", Borneo" authorityName="Borneo" box="[890,1051,996,1022]" class="Insecta" family="Pompilidae" genus="Java" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">Java, Borneo</taxonomicName>
,
<collectingCountry id="F353764B8D51FF84FBB5DDDFD4A8FC62" box="[1068,1205,996,1022]" name="Philippines" pageId="5" pageNumber="253">Philippines</collectingCountry>
), and
<collectingCountry id="F353764B8D51FF84FA9DDDDFD56AFC62" box="[1284,1399,996,1022]" name="China" pageId="5" pageNumber="253">East Asia</collectingCountry>
(
<collectingCountry id="F353764B8D51FF84FF5CDA3CD13DFBBD" box="[197,288,1031,1057]" name="Taiwan" pageId="5" pageNumber="253">Taiwan</collectingCountry>
, Eastern-central
<collectingCountry id="F353764B8D51FF84FE7DDA3CD230FBBD" box="[484,557,1031,1057]" name="China" pageId="5" pageNumber="253">China</collectingCountry>
,
<collectingCountry id="F353764B8D51FF84FDA0DA3CD2CDFBBD" box="[569,720,1031,1057]" name="South Korea" pageId="5" pageNumber="253">South Korea</collectingCountry>
,
<collectingCountry id="F353764B8D51FF84FD42DA3CD33EFBBD" box="[731,803,1031,1057]" name="Japan" pageId="5" pageNumber="253">Japan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E65508D51FF84FF24DA73D2D5F97F" pageId="5" pageNumber="253" type="discussion">
<paragraph id="8BFB36DB8D51FF84FF24DA73D12DFBFE" blockId="5.[189,304,1096,1122]" box="[189,304,1096,1122]" pageId="5" pageNumber="253">
<heading id="D0B381B78D51FF84FF24DA73D12DFBFE" bold="true" box="[189,304,1096,1122]" fontSize="11" level="3" pageId="5" pageNumber="253" reason="3">
<emphasis id="B930EAC98D51FF84FF24DA73D12DFBFE" bold="true" box="[189,304,1096,1122]" pageId="5" pageNumber="253">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DA4CD1F5FB67" blockId="5.[187,1399,1143,1275]" pageId="5" pageNumber="253">
<collectingCountry id="F353764B8D51FF84FF24DA4CD0FFFB0D" box="[189,226,1143,1169]" name="American Samoa" pageId="5" pageNumber="253">AS</collectingCountry>
examined the
<typeStatus id="54FF88798D51FF84FE1EDA4CD1F3FB0D" box="[391,494,1143,1169]" pageId="5" pageNumber="253" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C444D588D51FF84FD94DA4CD332FB0D" authorityName="Turner" authorityYear="1920" box="[525,815,1143,1169]" class="Insecta" family="Pompilidae" genus="Cryptosalius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B930EAC98D51FF84FD94DA4CD332FB0D" box="[525,815,1143,1169]" italics="true" pageId="5" pageNumber="253">Cryptosalius tonkinensis</emphasis>
</taxonomicName>
(
<specimenCount id="9D42FD528D51FF84FCDADA4CD34AFB0D" box="[835,855,1143,1169]" count="1" pageId="5" pageNumber="253" type="female">♀</specimenCount>
, Chapa,
<collectingCountry id="F353764B8D51FF84FC20DA4CD412FB0D" box="[953,1039,1143,1169]" name="Vietnam" pageId="5" pageNumber="253">Tonkin</collectingCountry>
,
<date id="FFFA101B8D51FF84FB81DA4CD4E4FB0D" box="[1048,1273,1143,1169]" pageId="5" pageNumber="253" value="1916-05" valueMax="1916-06-31" valueMin="1916-05-00">May and June 1916</date>
) deposited in the collection of the NHML and concluded that the species should belong to
<taxonomicName id="4C444D588D51FF84FBD7DAA0D4C2FB28" authorityName="Kohl" authorityYear="1907" box="[1102,1247,1179,1204]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FBD7DAA0D4C2FB28" box="[1102,1247,1179,1204]" italics="true" pageId="5" pageNumber="253">Lissocnemis</emphasis>
</taxonomicName>
. We propose the following nomenclatural change:
<taxonomicName id="4C444D588D51FF84FDE8DA85D422FB4B" authority="(Turner, 1920)" authorityName="Kim &amp; Shimizu &amp; Pitts" authorityYear="2024" baseAuthorityName="Turner" baseAuthorityYear="1920" box="[625,1087,1213,1239]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="tonkinensis" status="comb. nov.">
<emphasis id="B930EAC98D51FF84FDE8DA85D390FB4B" box="[625,909,1213,1239]" italics="true" pageId="5" pageNumber="253">Lissocnemis tonkinensis</emphasis>
(
<bibRefCitation id="EFD54B2A8D51FF84FC04DA86D42AFB4B" author="Turner R. E." box="[925,1079,1213,1239]" pageId="5" pageNumber="253" pagination="84 - 98" refId="ref11709" refString="Turner R. E. 1920. On Indo-Chinese Hymenoptera collected by R. Vitalis de Salvaza. IV. Annals and Magazine of Natural History (9) 5: 84 - 98. https: // doi. org / 10.1080 / 00222932008632344" type="journal article" year="1920">Turner, 1920</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357B28D51FF84FBDFDA86D4DAFB44" box="[1094,1223,1213,1240]" pageId="5" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
=
<taxonomicName id="4C444D588D51FF84FB46DA86D1F9FB67" authority="Turner, 1920" authorityName="Turner" authorityYear="1920" class="Insecta" family="Pompilidae" genus="Cryptosalius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B930EAC98D51FF84FB46DA86D15FFB66" italics="true" pageId="5" pageNumber="253">Cryptosalius tonkinensis</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FED0DADAD1F9FB67" author="Turner R. E." box="[329,484,1249,1275]" pageId="5" pageNumber="253" pagination="84 - 98" refId="ref11709" refString="Turner R. E. 1920. On Indo-Chinese Hymenoptera collected by R. Vitalis de Salvaza. IV. Annals and Magazine of Natural History (9) 5: 84 - 98. https: // doi. org / 10.1080 / 00222932008632344" type="journal article" year="1920">Turner, 1920</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFB36DB8D51FF84FF24DB18D2D5F97F" blockId="5.[187,1399,1314,1763]" pageId="5" pageNumber="253">
<taxonomicName id="4C444D588D51FF84FF24DB18D1CAFAA0" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[189,471,1314,1340]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FF24DB18D127FAA0" box="[189,314,1315,1340]" italics="true" pageId="5" pageNumber="253">Dinagenia</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FEDADB19D1CAFAA0" author="Banks N." box="[323,471,1314,1340]" pageId="5" pageNumber="253" pagination="1 - 117" refId="ref11043" refString="Banks N. 1934. The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences 69: 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
(type species:
<taxonomicName id="4C444D588D51FF84FD13DB18D385FAA0" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[650,920,1314,1340]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FD13DB18D2E1FAA0" box="[650,764,1314,1340]" italics="true" pageId="5" pageNumber="253">D. apollo</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FC9ADB19D385FAA0" author="Banks N." box="[771,920,1314,1340]" pageId="5" pageNumber="253" pagination="1 - 117" refId="ref11043" refString="Banks N. 1934. The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences 69: 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
) includes two species,
<taxonomicName id="4C444D588D51FF84FB34DB18D0E4FAC3" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FB34DB18D4DCFAA0" box="[1197,1217,1315,1340]" italics="true" pageId="5" pageNumber="253">D</emphasis>
.
<emphasis id="B930EAC98D51FF84FB48DB19D500FAA0" box="[1233,1309,1314,1340]" italics="true" pageId="5" pageNumber="253">apollo</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FABFDB19D0E4FAC3" author="Banks N." pageId="5" pageNumber="253" pagination="1 - 117" refId="ref11043" refString="Banks N. 1934. The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences 69: 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="F353764B8D51FF84FEE3DB7ED1E3FAC3" box="[378,510,1349,1375]" name="Philippines" pageId="5" pageNumber="253">Philippines</collectingCountry>
and
<taxonomicName id="4C444D588D51FF84FDA6DB7DD37CFAC3" authority="Banks, 1938" authorityName="Banks" authorityYear="1938" box="[575,865,1349,1376]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="B930EAC98D51FF84FDA6DB7DD249FAC3" box="[575,596,1350,1375]" italics="true" pageId="5" pageNumber="253">D</emphasis>
.
<emphasis id="B930EAC98D51FF84FDFEDB7DD2A0FAC3" box="[615,701,1350,1375]" italics="true" pageId="5" pageNumber="253">satyrus</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FD50DB7ED37CFAC3" author="Banks N." box="[713,865,1349,1375]" pageId="5" pageNumber="253" refId="ref11079" refString="Banks N. 1938. The Psammocharidae from Singapore. Proceedings of the Entomological Society of Washington 40: 2361 - 249." type="journal volume" year="1938">Banks, 1938</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F353764B8D51FF84FC28DB7ED431FAC3" box="[945,1068,1349,1375]" name="Singapore" pageId="5" pageNumber="253">Singapore</collectingCountry>
.
<bibRefCitation id="EFD54B2A8D51FF84FBA5DB7DD4FDFAC3" author="Haupt H." box="[1084,1248,1349,1376]" pageId="5" pageNumber="253" pagination="33 - 48" refId="ref11265" refString="Haupt H. 1938. Psammocharidae vom unteren Yang-tse. Notes d'Entomologie Chinoise 5: 33 - 48." type="journal article" year="1938">Haupt (1938)</bibRefCitation>
,
<bibRefCitation id="EFD54B2A8D51FF84FB68DB7DD14FFA1E" author="Matsumoto R. &amp; Miyake H. M. &amp; Yamazaki K. &amp; Aono T. &amp; Shimizu A." pageId="5" pageNumber="253" pagination="95 - 98" refId="ref11417" refString="Matsumoto R., Miyake H. M., Yamazaki K., Aono T. &amp; Shimizu A. 2018. Discovery of a new, adventive, spider Wasp, Lissocnemis brevipennis, in Japan (Hymenoptera: Pompilidae: Ctenocerinae). Japanese Journal of Systematic Entomology 24: 95 - 98." type="journal article" year="2018">
Matsumoto
<emphasis id="B930EAC98D51FF84FF24DB52D0E7FA1E" box="[189,250,1384,1410]" italics="true" pageId="5" pageNumber="253">et al.</emphasis>
(2018)
</bibRefCitation>
and Michael C. Day (unpubl.) pointed out the genus was probably a junior synonym of
<taxonomicName id="4C444D588D51FF84FF26DBB7D153FA39" authorityName="Kohl" authorityYear="1907" box="[191,334,1420,1445]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FF26DBB7D153FA39" box="[191,334,1420,1445]" italics="true" pageId="5" pageNumber="253">Lissocnemis</emphasis>
</taxonomicName>
. JPP examined the
<typeStatus id="54FF88798D51FF84FDB7DBB0D2BDFA39" box="[558,672,1419,1445]" pageId="5" pageNumber="253" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="4C444D588D51FF84FD5ADBB7D32DFA39" authorityName="Banks" authorityYear="1934" box="[707,816,1419,1445]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FD5ADBB7D32DFA39" box="[707,816,1419,1445]" italics="true" pageId="5" pageNumber="253">D. apollo</emphasis>
</taxonomicName>
(
<specimenCount id="9D42FD528D51FF84FCDDDBB7D345FA3A" box="[836,856,1420,1446]" count="1" pageId="5" pageNumber="253" type="female">♀</specimenCount>
, “Surigao Mindanao Baker /
<taxonomicName id="4C444D588D51FF84FB33DBB7D0F6FA55" authority="Bks" authorityName="Banks" authorityYear="1934" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FB33DBB7D56BFA39" box="[1194,1398,1419,1445]" italics="true" pageId="5" pageNumber="253">Dinagenia apollo</emphasis>
Bks
</taxonomicName>
type / Type No. 51162
<collectingCountry id="F353764B8D51FF84FE61DB94D234FA55" box="[504,553,1455,1481]" name="United States of America" pageId="5" pageNumber="253">U.S.</collectingCountry>
N.M. / USNMENT 01546711 /
<taxonomicName id="4C444D588D51FF84FC00DB94D4AAFA55" authority="Bks." authorityName="Bks." box="[921,1207,1455,1481]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FC00DB94D467FA55" box="[921,1146,1455,1481]" italics="true" pageId="5" pageNumber="253">Lissocnemis apollo</emphasis>
Bks.
</taxonomicName>
<specimenCount id="9D42FD528D51FF84FB25DB94D4CFFA55" box="[1212,1234,1455,1481]" count="1" pageId="5" pageNumber="253" type="female">♀</specimenCount>
det.M.C.Day, 1977”) and
<taxonomicName id="4C444D588D51FF84FEDBDBE8D1A4FA70" authorityName="Banks" authorityYear="1938" box="[322,441,1491,1516]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="B930EAC98D51FF84FEDBDBE8D1A4FA70" box="[322,441,1491,1516]" italics="true" pageId="5" pageNumber="253">D. satyrus</emphasis>
</taxonomicName>
(
<specimenCount id="9D42FD528D51FF84FE55DBE9D1FDFA70" box="[460,480,1490,1516]" count="1" pageId="5" pageNumber="253" type="female">♀</specimenCount>
, “
<collectingCountry id="F353764B8D51FF84FE6EDBE9D26CFA70" box="[503,625,1490,1516]" name="Singapore" pageId="5" pageNumber="253">Singapore</collectingCountry>
Coll. Baker /
<taxonomicName id="4C444D588D51FF84FC96DBE8D405FA70" authority="Bks" authorityName="Banks" authorityYear="1938" box="[783,1048,1490,1516]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="B930EAC98D51FF84FC96DBE8D3F8FA70" box="[783,997,1491,1516]" italics="true" pageId="5" pageNumber="253">Dinagenia satyrus</emphasis>
Bks
</taxonomicName>
type / Type No. 56021 USNM / USNMENT 01559147 / 17915 /
<taxonomicName id="4C444D588D51FF84FDC2DBCDD393F993" authority="Banks" authorityName="Banks" box="[603,910,1525,1551]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FDC2DBCDD323F993" box="[603,830,1525,1551]" italics="true" pageId="5" pageNumber="253">Lissocnemis apollo</emphasis>
Banks
</taxonomicName>
<specimenCount id="9D42FD528D51FF84FC01DBCDD3B3F98C" box="[920,942,1526,1552]" count="1" pageId="5" pageNumber="253" type="female">♀</specimenCount>
det.M.C.Day, 1977”) deposited in the collection of the NMNH and concluded that
<taxonomicName id="4C444D588D51FF84FD5ED822D359F9AE" authorityName="Banks" authorityYear="1934" box="[711,836,1561,1586]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FD5ED822D359F9AE" box="[711,836,1561,1586]" italics="true" pageId="5" pageNumber="253">Dinagenia</emphasis>
</taxonomicName>
should be a junior synonym of
<taxonomicName id="4C444D588D51FF84FB21D822D554F9AE" authorityName="Kohl" authorityYear="1907" box="[1208,1353,1561,1586]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FB21D822D554F9AE" box="[1208,1353,1561,1586]" italics="true" pageId="5" pageNumber="253">Lissocnemis</emphasis>
</taxonomicName>
. As such, these two species become new combinations in
<taxonomicName id="4C444D588D51FF84FCD9D807D3CDF9C9" authorityName="Kohl" authorityYear="1907" box="[832,976,1596,1621]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAC98D51FF84FCD9D807D3CDF9C9" box="[832,976,1596,1621]" italics="true" pageId="5" pageNumber="253">Lissocnemis</emphasis>
</taxonomicName>
:
<taxonomicName id="4C444D588D51FF84FC79D807D56BF9CA" authority="(Banks, 1934)" authorityName="Kim &amp; Shimizu &amp; Pitts" authorityYear="2024" baseAuthorityName="Banks" baseAuthorityYear="1934" box="[992,1398,1595,1622]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo" status="comb. nov.">
<emphasis id="B930EAC98D51FF84FC79D807D4DBF9C9" box="[992,1222,1595,1621]" italics="true" pageId="5" pageNumber="253">Lissocnemis apollo</emphasis>
(
<bibRefCitation id="EFD54B2A8D51FF84FB4ED800D573F9CA" author="Banks N." box="[1239,1390,1595,1622]" pageId="5" pageNumber="253" pagination="1 - 117" refId="ref11043" refString="Banks N. 1934. The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences 69: 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357B28D51FF84FF27D864D15CF9E5" box="[190,321,1631,1657]" pageId="5" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
(=
<taxonomicName id="4C444D588D51FF84FEF6D864D2C2F9E5" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[367,735,1631,1657]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo">
<emphasis id="B930EAC98D51FF84FEF6D864D25FF9E5" box="[367,578,1631,1657]" italics="true" pageId="5" pageNumber="253">Dinagenia apollo</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FDD0D864D2C2F9E5" author="Banks N." box="[585,735,1631,1657]" pageId="5" pageNumber="253" pagination="1 - 117" refId="ref11043" refString="Banks N. 1934. The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences 69: 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
) and
<taxonomicName id="4C444D588D51FF84FCBFD864D4D5F9E5" authority="(Banks, 1938)" authorityName="Kim &amp; Shimizu &amp; Pitts" authorityYear="2024" baseAuthorityName="Banks" baseAuthorityYear="1938" box="[806,1224,1631,1657]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus" status="comb. nov.">
<emphasis id="B930EAC98D51FF84FCBFD864D40AF9E4" box="[806,1047,1631,1656]" italics="true" pageId="5" pageNumber="253">Lissocnemis satyrus</emphasis>
(
<bibRefCitation id="EFD54B2A8D51FF84FBB0D864D4DDF9E5" author="Banks N." box="[1065,1216,1631,1657]" pageId="5" pageNumber="253" refId="ref11079" refString="Banks N. 1938. The Psammocharidae from Singapore. Proceedings of the Entomological Society of Washington 40: 2361 - 249." type="journal volume" year="1938">Banks, 1938</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357B28D51FF84FB4BD864D54EF9E5" box="[1234,1363,1631,1657]" pageId="5" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
(=
<taxonomicName id="4C444D588D51FF84FF26D8B8D25CF900" authority="Banks, 1938" authorityName="Banks" authorityYear="1938" box="[191,577,1666,1692]" class="Insecta" family="Pompilidae" genus="Dinagenia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="B930EAC98D51FF84FF26D8B8D180F900" box="[191,413,1667,1692]" italics="true" pageId="5" pageNumber="253">Dinagenia satyrus</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FE3FD8B9D25CF900" author="Banks N." box="[422,577,1666,1692]" pageId="5" pageNumber="253" refId="ref11079" refString="Banks N. 1938. The Psammocharidae from Singapore. Proceedings of the Entomological Society of Washington 40: 2361 - 249." type="journal volume" year="1938">Banks, 1938</bibRefCitation>
</taxonomicName>
). Furthermore, close examination of
<typeStatus id="54FF88798D51FF84FB9DD8B9D476F900" box="[1028,1131,1666,1692]" pageId="5" pageNumber="253" type="holotype">holotype</typeStatus>
material has revealed that
<taxonomicName id="4C444D588D51FF84FF68D89DD286F923" authority="Haupt, 1941" authorityName="Haupt" authorityYear="1941" box="[241,667,1701,1728]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="nigricoxis">
<emphasis id="B930EAC98D51FF84FF68D89DD21CF923" box="[241,513,1702,1727]" italics="true" pageId="5" pageNumber="253">Lissocnemis nigricoxis</emphasis>
<bibRefCitation id="EFD54B2A8D51FF84FD90D89DD286F923" author="Haupt H." box="[521,667,1701,1728]" pageId="5" pageNumber="253" pagination="50 - 82" refId="ref11284" refString="Haupt H. 1941. Resultate der Oxford Universitat Expedition nach Sarawak (Borneo), 1932. Beitrag zur Kenntnis der Psammochariden-Fauna. Annals and Magazine of Natural History; including Zoology, Botany and Geology (11) 7: 50 - 82. https: // doi. org / 10.1080 / 00222934108527140" type="journal article" year="1941">Haupt, 1941</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C444D588D51FF84FD4CD89DD46EF923" authority="(Banks, 1938)" authorityName="Kim &amp; Shimizu &amp; Pitts" authorityYear="2024" baseAuthorityName="Banks" baseAuthorityYear="1938" box="[725,1139,1701,1727]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="satyrus" status="comb. nov.">
<emphasis id="B930EAC98D51FF84FD4CD89DD3D8F923" box="[725,965,1702,1727]" italics="true" pageId="5" pageNumber="253">Lissocnemis satyrus</emphasis>
(
<bibRefCitation id="EFD54B2A8D51FF84FC4CD89ED476F923" author="Banks N." box="[981,1131,1701,1727]" pageId="5" pageNumber="253" refId="ref11079" refString="Banks N. 1938. The Psammocharidae from Singapore. Proceedings of the Entomological Society of Washington 40: 2361 - 249." type="journal volume" year="1938">Banks, 1938</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357B28D51FF84FBE2D89ED4E0F95C" box="[1147,1277,1701,1728]" pageId="5" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
are junior synonyms of
<taxonomicName id="4C444D588D51FF84FEC2D8F2D25DF97E" authorityName="Kim &amp; Shimizu &amp; Pitts" authorityYear="2024" baseAuthorityName="Banks" baseAuthorityYear="1934" box="[347,576,1736,1762]" class="Insecta" family="Pompilidae" genus="Lissocnemis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="253" phylum="Arthropoda" rank="species" species="apollo" status="comb. nov.">
<emphasis id="B930EAC98D51FF84FEC2D8F2D25DF97E" box="[347,576,1736,1762]" italics="true" pageId="5" pageNumber="253">Lissocnemis apollo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357B28D51FF84FDDED8F3D2D5F97F" box="[583,712,1736,1763]" pageId="5" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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