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<mods:title id="4F38E39F532F1F8517E7CB027564FB91">Notes on the Dianous bimaculatus complex with description of a new species from China (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="8960A1C038783BE12CD6B977E1463BB3">Tang, Liang</mods:namePart>
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<mods:namePart id="9D097FFD615880A238B2D5F2728448C3">Huang, Meng-Chi</mods:namePart>
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<mods:date id="2D6DCE8149B4325AA048921247628543">2018</mods:date>
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<taxonomicName id="4C44C2E3A518FFC4FF7C401EB25D7A0F" ID-CoL="359W3" authority="Cameron, 1927" authorityName="Cameron" authorityYear="1927" box="[151,606,580,607]" class="Insecta" family="Staphylinidae" genus="Dianous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B9306572A518FFC4FF7C401EB25D7A0F" bold="true" box="[151,606,580,607]" pageId="1" pageNumber="561">
<emphasis id="B9306572A518FFC4FF7C401EB19A7A0E" bold="true" box="[151,409,580,606]" italics="true" pageId="1" pageNumber="561">Dianous bimaculatus</emphasis>
<bibRefCitation id="EFD5C491A518FFC4FE4B401FB25D7A0F" author="Cameron, M." box="[416,606,581,607]" pageId="1" pageNumber="561" pagination="6 - 10" refId="ref4366" refString="Cameron, M. (1927) New species of Indian Dianous. The Entomologist's Monthly Magazine, 62, 6 - 10." type="journal article" year="1927">Cameron, 1927</bibRefCitation>
</emphasis>
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<paragraph id="8BFBB960A518FFC4FF7C403DB12D7AD0" blockId="1.[151,606,580,640]" box="[151,302,615,640]" pageId="1" pageNumber="561">
(
<figureCitation id="137FA5E5A518FFC4FF74403DB0EF7AD0" box="[159,236,615,640]" captionStart="FIGURES 16" captionStartId="2.[151,264,1984,2007]" captionTargetBox="[151,1393,370,1953]" captionTargetId="figure@2.[151,1435,370,1970]" captionTargetPageId="2" captionText="FIGURES 16. Habitus. 1 Dianous bimaculatus 2 D. flavoculatus 3 D. hubeiensis 4 D. hainanensis 5 D. siamensis 6 D. strabo. Scale bars: 1mm." httpUri="https://zenodo.org/record/2612511/files/figure.png" pageId="1" pageNumber="561">Figs. 1</figureCitation>
, 7, 8)
</paragraph>
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<taxonomicName id="4C44C2E3A518FFC4FF7C40EBB22E7A96" ID-CoL="359W3" authority="Cameron, 1927: 6" authorityName="Cameron" authorityPageNumber="6" authorityYear="1927" box="[151,557,688,710]" class="Insecta" family="Staphylinidae" genus="Dianous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B9306572A518FFC4FF7C40EBB16C7A96" box="[151,367,688,710]" italics="true" pageId="1" pageNumber="561">Dianous bimaculatus</emphasis>
<treatmentCitation id="0AE59F71A518FFC4FE9E40EAB22E7A96" author="Cameron, M." box="[373,557,688,710]" page="6" pageId="1" pageNumber="561" year="1927">
<bibRefCitation id="EFD5C491A518FFC4FE9E40EAB2177A96" author="Cameron, M." box="[373,532,688,710]" pageId="1" pageNumber="561" pagination="6 - 10" refId="ref4366" refString="Cameron, M. (1927) New species of Indian Dianous. The Entomologist's Monthly Magazine, 62, 6 - 10." type="journal article" year="1927">Cameron, 1927</bibRefCitation>
: 6
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;
<treatmentCitation id="0AE59F71A518FFC4FDD340EAB2EB7A96" author="Puthz, V." box="[568,744,688,710]" page="427" pageId="1" pageNumber="561" year="2000">
<bibRefCitation id="EFD5C491A518FFC4FDD340EAB2B57A96" author="Puthz, V." box="[568,694,688,710]" pageId="1" pageNumber="561" pagination="419 - 559" refId="ref4490" refString="Puthz, V. (2000) The genus Dianous Leach in China (Coleoptera, Staphylinidae). 261. Contribution to the knowledge of Steninae. Revue Suisse de Zoologie, 107 (3), 419 - 559. https: // doi. org / 10.5962 / bhl. part. 80137" type="journal article" year="2000">Puthz, 2000</bibRefCitation>
: 427
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;
<treatmentCitation id="0AE59F71A518FFC4FD1940EAB3AE7A96" author="Puthz, V." box="[754,941,688,710]" httpUri="http://treatment.plazi.org/id/534B87AEFF86091AFF03BCD5FDA39621" page="1741" pageId="1" pageNumber="561" year="2015">
<bibRefCitation id="EFD5C491A518FFC4FD1940EAB36D7A96" author="Puthz, V." box="[754,878,688,710]" pageId="1" pageNumber="561" pagination="1729 - 1746" refId="ref4546" refString="Puthz, V. (2015) Neue und alte Steninen aus Thailand (Coleoptera, Staphylinidae). 342. Beitrag zur Kenntnis der Steninen. Linzer biologische Beitrage, 47 (2), 1729 - 1746." type="journal article" year="2015">Puthz, 2015</bibRefCitation>
: 1741
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;
<treatmentCitation id="0AE59F71A518FFC4FC5340EAB46B7A96" author="Puthz, V." box="[952,1128,688,710]" httpUri="http://treatment.plazi.org/id/03926B6E4534572F24B3FE49FD66FDBE" page="747" pageId="1" pageNumber="561" year="2016">
<bibRefCitation id="EFD5C491A518FFC4FC5340EAB4367A96" author="Puthz, V." box="[952,1077,688,710]" pageId="1" pageNumber="561" pagination="705 - 778" refId="ref4586" refString="Puthz, V. (2016) Ubersicht uber die Arten der Gattung Dianous Leach group II (Coleoptera, Staphylinidae). 347. Beitrag zur Kenntnis der Steninen. Linzer biologische Beitrage, 48 (1), 705 - 778." type="journal article" year="2016">Puthz, 2016</bibRefCitation>
: 747
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.
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</paragraph>
</subSubSection>
<subSubSection id="C35EEAEBA518FFC4FF7C40A8B5047B5B" box="[151,1287,754,779]" pageId="1" pageNumber="561" type="materials_examined">
<paragraph id="8BFBB960A518FFC4FF7C40A8B5047B5B" blockId="1.[151,1436,754,959]" box="[151,1287,754,779]" pageId="1" pageNumber="561">
<emphasis id="B9306572A518FFC4FF7C40A8B1D57B5B" bold="true" box="[151,470,754,779]" pageId="1" pageNumber="561">
Material examined.
<materialsCitation id="3B2CB33DA518FFC4FE6C40A8B1D57B5B" ID-GBIF-Occurrence="2029196009" box="[391,470,754,779]" collectingDate="1993-05-02" collectorName="A. Kleeber" country="Nepal" location="Nepal" pageId="1" pageNumber="561" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F353F9F0A518FFC4FE6C40A8B1D27B5B" box="[391,465,754,779]" name="Nepal" pageId="1" pageNumber="561">Nepal</collectingCountry>
:
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</emphasis>
<specimenCount id="9D4272E9A518FFC4FE3640A9B2017B5B" box="[477,514,755,779]" pageId="1" pageNumber="561" type="male">1♂</specimenCount>
, Umg Shivalaya, Ufer Kimti Khols,
<date id="FFFA9FA0A518FFC4FC7640A9B4077B5B" box="[925,1028,755,779]" pageId="1" pageNumber="561" value="1993-05-02">
<collectingDate id="EFBE6648A518FFC4FC7640A9B4077B5B" box="[925,1028,755,779]" pageId="1" pageNumber="561" value="1993-05-02">2.V.1993</collectingDate>
</date>
,
<collectorName id="26B1DCB6A518FFC4FBFB40A9B4887B5B" box="[1040,1163,754,779]" pageId="1" pageNumber="561">A. Kleeber</collectorName>
leg. (cPut)
</paragraph>
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<subSubSection id="C35EEAEBA518FFC4FF2C414CB2527B03" pageId="1" pageNumber="561" type="description">
<paragraph id="8BFBB960A518FFC4FF2C414CB2527B03" blockId="1.[151,1436,754,959]" pageId="1" pageNumber="561">
<emphasis id="B9306572A518FFC4FF2C414CB17E7B7F" bold="true" box="[199,381,790,815]" pageId="1" pageNumber="561">Measurements.</emphasis>
BL:
<quantity id="4CBC1485A518FFC4FE53414CB2137B7E" box="[440,528,790,815]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="561" unit="mm" value="5.7">5.7 mm</quantity>
, FL:
<quantity id="4CBC1485A518FFC4FDA6414DB2A57B7E" box="[589,678,791,815]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="561" unit="mm" value="2.8">2.8 mm</quantity>
. Head 1.01 times as wide as elytra, pronotum 1.25 times as long as wide, elytra 1.16 times as long as wide.
</paragraph>
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<paragraph id="8BFBB960A518FFC4FF2C4104B3897B27" blockId="1.[151,1436,754,959]" box="[199,906,862,887]" pageId="1" pageNumber="561">
<emphasis id="B9306572A518FFC4FF2C4104B1637B27" bold="true" box="[199,352,862,887]" pageId="1" pageNumber="561">Distribution.</emphasis>
<collectingCountry id="F353F9F0A518FFC4FE8C4104B1A97B27" box="[359,426,862,887]" name="China" pageId="1" pageNumber="561">China</collectingCountry>
(
<collectingRegion id="49807782A518FFC4FE524105B2167B27" box="[441,533,863,887]" country="China" name="Yunnan" pageId="1" pageNumber="561">Yunnan</collectingRegion>
),
<collectingCountry id="F353F9F0A518FFC4FDCC4104B2607B27" box="[551,611,862,887]" name="India" pageId="1" pageNumber="561">India</collectingCountry>
,
<collectingCountry id="F353F9F0A518FFC4FD844104B2B67B27" box="[623,693,862,887]" name="Nepal" pageId="1" pageNumber="561">Nepal</collectingCountry>
,
<collectingCountry id="F353F9F0A518FFC4FD544104B3267B27" box="[703,805,862,887]" name="Thailand" pageId="1" pageNumber="561">Thailand</collectingCountry>
,
<collectingCountry id="F353F9F0A518FFC4FCDA4104B3857B27" box="[817,902,862,887]" name="Bhutan" pageId="1" pageNumber="561">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35EEAEBA518FFC4FF2C41D8B4247BEF" pageId="1" pageNumber="561" type="discussion">
<paragraph id="8BFBB960A518FFC4FF2C41D8B4247BEF" blockId="1.[151,1436,754,959]" pageId="1" pageNumber="561">
<emphasis id="B9306572A518FFC4FF2C41D8B1387BCB" bold="true" box="[199,315,898,923]" pageId="1" pageNumber="561">Remarks.</emphasis>
The species is similar to
<taxonomicName id="4C44C2E3A518FFC4FDB541D9B3087BCB" authorityName="Puthz" authorityYear="1997" box="[606,779,899,923]" class="Insecta" family="Staphylinidae" genus="Dianous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="species" species="flavoculatus">
<emphasis id="B9306572A518FFC4FDB541D9B3087BCB" box="[606,779,899,923]" italics="true" pageId="1" pageNumber="561">D. flavoculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C44C2E3A518FFC4FCAF41D9B3EE7BCB" authorityName="Puthz" authorityYear="1997" box="[836,1005,899,923]" class="Insecta" family="Staphylinidae" genus="Dianous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B9306572A518FFC4FCAF41D9B3EE7BCB" box="[836,1005,899,923]" italics="true" pageId="1" pageNumber="561">D. hainanensis</emphasis>
</taxonomicName>
in aedeagus, but can be distinguished from them by smaller elytral spots and much larger expulsion clasps of aedeagus.
</paragraph>
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</treatment>
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