treatments-xml/data/03/E9/D1/03E9D100FFD4FFFAFF59FB61464CFCE0.xml

<document ID-DOI="10.25221/fee.405.2" ID-ISSN="2713-2196" ID-Zenodo-Dep="7165322" approvalRequired="44" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="42" approvalRequired_for_treatments="1" checkinTime="1665249059075" checkinUser="felipe" docAuthor="Zettel, H. &amp; Pangantihon, C. V." docDate="2020" docId="03E9D100FFD4FFFAFF59FB61464CFCE0" docLanguage="en" docName="FarEastEntomol.405.8-14.pdf" docOrigin="Far Eastern Entomologist 405" docSource="http://dx.doi.org/10.25221/fee.405.2" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Timasius alveus Zettel et Pangantihon 2020, sp. n." docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FFD0A978FFD6FFFFFFBAFFD34770FF9C" masterDocTitle="A NEW SPECIES OF THE TIMASIUS LIVENS SPECIES GROUP (HEMIPTERA: HETEROPTERA: HEBRIDAE) FROM CAMBODIA AND VIETNAM" masterLastPageNumber="14" masterPageNumber="8" pageNumber="10" updateTime="1665263028783" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>A NEW SPECIES OF THE TIMASIUS LIVENS SPECIES GROUP (HEMIPTERA: HETEROPTERA: HEBRIDAE) FROM CAMBODIA AND VIETNAM</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zettel, H.</mods:namePart>
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<mods:namePart>Pangantihon, C. V.</mods:namePart>
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<mods:title>Far Eastern Entomologist</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>2020-04-14</mods:number>
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<mods:number>405</mods:number>
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<taxonomicName authority="Zettel et Pangantihon" authorityName="Zettel et Pangantihon" authorityYear="2020" box="[227,615,1201,1223]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="10" phylum="Arthropoda" rank="species" species="alveus" status="sp. n.">
<emphasis bold="true" box="[227,387,1201,1223]" italics="true" pageId="2" pageNumber="10">Timasius alveus</emphasis>
Zettel et Pangantihon
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, 
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http://zoobank.org/NomenclaturalActs/ 
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<figureCitation box="[227,322,1267,1289]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="3.[265,309,1142,1164]" captionStartId-1="4.[265,309,1046,1068]" captionTargetBox-0="[227,1122,503,1108]" captionTargetBox-1="[229,1120,258,1008]" captionTargetId-0="figure-115@3.[227,1122,503,1109]" captionTargetId-1="figure-1@4.[227,1122,257,1012]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs. 1–2: Habitus of Timasius alveus sp. n., male, in dorsal (1) and ventral (2) aspect. © Harald Bruckner." captionText-1="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi-0="http://doi.org/10.5281/zenodo.7165324" figureDoi-1="http://doi.org/10.5281/zenodo.7165326" httpUri-0="https://zenodo.org/record/7165324/files/figure.png" httpUri-1="https://zenodo.org/record/7165326/files/figure.png" pageId="2" pageNumber="10">Figs 1–10</figureCitation>
</paragraph>
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<subSubSection lastPageId="3" lastPageNumber="11" pageId="2" pageNumber="10" type="materials_examined">
<paragraph blockId="2.[227,1128,1322,1593]" box="[265,1127,1322,1344]" pageId="2" pageNumber="10">
<materialsCitation box="[265,1127,1322,1344]" collectionCode="TYPE, MATERIAL" location="Natural History Museum" pageId="2" pageNumber="10" specimenCount="1" stateProvince="Wien" typeStatus="holotype">
<collectionCode box="[265,459,1322,1344]" pageId="2" pageNumber="10">TYPE MATERIAL</collectionCode>
. 
<typeStatus box="[470,559,1322,1344]" pageId="2" pageNumber="10">Holotype</typeStatus>
– (macropterous male, 
<location LSID="urn:lsid:plazi:treatment:03E9D100FFD4FFFAFF59FB61464CFCE0:8E9F36CDFFD4FFFDFCA3FAF9437CFADC" box="[793,1036,1322,1344]" name="Natural History Museum" pageId="2" pageNumber="10" stateProvince="Wien">Natural History Museum</location>
<collectingRegion box="[1043,1116,1322,1344]" country="Austria" name="Wien" pageId="2" pageNumber="10">Vienna</collectingRegion>
),
</materialsCitation>
</paragraph>
<paragraph blockId="2.[227,1128,1322,1593]" pageId="2" pageNumber="10">
<materialsCitation country="Cambodia" elevation="612" latitude="12.443" location="Mondulkiri Prov." longLatPrecision="13" longitude="107.15967" pageId="2" pageNumber="10" specimenCount="1" stateProvince="Mondolkiri">
labelled “ 
<emphasis bold="true" box="[325,432,1350,1372]" pageId="2" pageNumber="10">
<collectingCountry box="[325,432,1350,1372]" name="Cambodia" pageId="2" pageNumber="10">Cambodia</collectingCountry>
</emphasis>
: 
<collectingRegion box="[450,624,1350,1372]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="10">Mondulkiri Prov.</collectingRegion>
\ Saen Monourom City, W Kroeng \ Saen Monourom, 
<geoCoordinate box="[320,445,1377,1399]" degrees="12" direction="north" minutes="26.58" orientation="latitude" pageId="2" pageNumber="10" precision="9" value="12.443">N 12°26.58'</geoCoordinate>
\ 
<geoCoordinate box="[475,598,1377,1399]" degrees="107" direction="east" minutes="09.58" orientation="longitude" pageId="2" pageNumber="10" precision="9" value="107.15967">E107°09.58'</geoCoordinate>
, 
<quantity box="[612,682,1377,1399]" metricMagnitude="2" metricUnit="m" metricValue="6.12" pageId="2" pageNumber="10" unit="m" value="612.0">
<elevation box="[612,682,1377,1399]" metricMagnitude="2" metricUnit="m" metricValue="6.12" pageId="2" pageNumber="10" unit="m" value="612.0">612 m</elevation>
</quantity>
, 22.XI. \ leg. 
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P592 (MO-4)”
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[227,1128,1322,1593]" pageId="2" pageNumber="10">
<materialsCitation country="Vietnam" location="Vietnam" pageId="2" pageNumber="10" specimenCount="1" typeStatus="paratype">
<typeStatus box="[227,324,1406,1427]" pageId="2" pageNumber="10">Paratypes</typeStatus>
: 1 macropterous male (in the collection of the first author, genitalia lost), same label data; 9 macropterous males, 3 macropterous females, labelled “ 
<emphasis bold="true" box="[897,1005,1432,1454]" pageId="2" pageNumber="10">
S 
<collectingCountry box="[917,1005,1432,1454]" name="Vietnam" pageId="2" pageNumber="10">Vietnam</collectingCountry>
</emphasis>
, 
<quantity box="[1018,1080,1433,1455]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="10" unit="km" value="40.0">40 km</quantity>
NW
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</paragraph>
<paragraph blockId="2.[227,1128,1322,1593]" box="[227,1127,1460,1482]" pageId="2" pageNumber="10">
An Khe\ Buon Luoi, 14°10’N, 108°30’E\ 
<quantity box="[667,768,1460,1482]" metricMagnitude="2" metricUnit="m" metricValue="6.85" metricValueMax="7.5" metricValueMin="6.2" pageId="2" pageNumber="10" unit="m" value="685.0" valueMax="750.0" valueMin="620.0">620-750m</quantity>
,28.3.- 
<date box="[823,922,1460,1482]" pageId="2" pageNumber="10" value="1995-12-04">12.4.1995</date>
\ leg. Pacholatko &amp;
</paragraph>
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<materialsCitation location="University of Science" municipality="Zoological Collection of Biological Museum" pageId="2" pageNumber="10" specimenCount="1" stateProvince="Wien">
Dembicky” (Natural History Museum 
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, 
<collectingMunicipality box="[687,1126,1488,1510]" pageId="2" pageNumber="10">Zoological Collection of Biological Museum</collectingMunicipality>
at 
<collectingRegion box="[250,309,1515,1537]" country="Vietnam" name="Ha Noi" pageId="2" pageNumber="10">Hanoi</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03E9D100FFD4FFFAFF59FB61464CFCE0:8E9F36CDFFD4FFFDFE81FA38457FF99D" box="[315,527,1515,1537]" municipality="Zoological Collection of Biological Museum" name="University of Science" pageId="2" pageNumber="10" stateProvince="Wien">University of Science</location>
)
</materialsCitation>
.
</paragraph>
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<collectionCode box="[265,458,1543,1564]" pageId="2" pageNumber="10">TYPE LOCALITY</collectionCode>
. 
<emphasis bold="true" box="[472,579,1543,1565]" pageId="2" pageNumber="10">
<collectingCountry box="[472,579,1543,1565]" name="Cambodia" pageId="2" pageNumber="10">Cambodia</collectingCountry>
</emphasis>
: 
<collectingRegion box="[596,804,1543,1565]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="10">Mondulkiri Province</collectingRegion>
, Saen Monourom City, west of
</paragraph>
<paragraph blockId="2.[227,1128,1322,1593]" box="[227,943,1571,1593]" pageId="2" pageNumber="10">
Kroeng Saen Monourom Waterfall, 
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, 
<geoCoordinate box="[701,824,1571,1593]" degrees="107" direction="east" minutes="09.58" orientation="longitude" pageId="2" pageNumber="10" precision="9" value="107.15967">E107°09.58'</geoCoordinate>
, 
<quantity box="[832,893,1571,1593]" metricMagnitude="2" metricUnit="m" metricValue="6.12" pageId="2" pageNumber="10" unit="m" value="612.0">612 m</quantity>
a.s.l.
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<paragraph blockId="3.[227,1127,260,475]" pageId="3" pageNumber="11">
<materialsCitation collectionCode="HABITAT, AT, LOCALITY" pageId="3" pageNumber="11" specimenCount="1" typeStatus="holotype">
<collectionCode box="[265,409,260,281]" pageId="3" pageNumber="11">HABITAT AT</collectionCode>
<typeStatus box="[416,476,260,281]" pageId="3" pageNumber="11">TYPE</typeStatus>
<collectionCode box="[484,607,260,281]" pageId="3" pageNumber="11">LOCALITY</collectionCode>
. The specimens were collected from wet rocks in the vicinity of the falls
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.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="13" pageId="3" pageNumber="11" type="description">
<paragraph blockId="3.[227,1127,260,475]" box="[265,1127,315,337]" pageId="3" pageNumber="11">
DESCRIPTION. 
<emphasis bold="true" box="[437,641,315,336]" pageId="3" pageNumber="11">Macropterous male</emphasis>
. Measurements of 
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: BL 2.66, HL 0.61,
</paragraph>
<paragraph blockId="3.[227,1127,260,475]" box="[227,1127,342,366]" pageId="3" pageNumber="11">
HW 0.49, A 
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L 0.25, PL 0.60, PW 1.18, MtL 1.02, AW 1.07. Indices: HI 125, AtI 82, EI 67,
</paragraph>
<paragraph blockId="3.[227,1127,260,475]" box="[227,1126,370,392]" pageId="3" pageNumber="11">AnI 57, PHI 241, PnI 197, MMI 43, MtI 87, AbI 223. Relative lengths of antennomeres 1 – 4</paragraph>
<paragraph blockId="3.[227,1127,260,475]" pageId="3" pageNumber="11">(in % of antennomere 2): 135: 100: 130: 223. Relative lengths of leg segments (in % of metatibia): profemur 55, protibia 60, protarsus 20, mesofemur 61, mesotibia 70, mesotarsus</paragraph>
<paragraph blockId="3.[227,1127,260,475]" box="[227,692,453,475]" pageId="3" pageNumber="11">22, metafemur 83, metatibia 100, metatarsus 27.</paragraph>
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<paragraph blockId="3.[227,1116,1141,1191]" box="[265,1116,1141,1164]" pageId="3" pageNumber="11">
Figs. 1–2: Habitus of 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[474,627,1142,1164]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="11" phylum="Arthropoda" rank="species" species="alveus" status="sp. n.">
<emphasis box="[474,627,1142,1164]" italics="true" pageId="3" pageNumber="11">Timasius alveus</emphasis>
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, male, in dorsal (1) and ventral (2) aspect. ©
</paragraph>
<paragraph blockId="3.[227,1116,1141,1191]" box="[227,395,1169,1191]" pageId="3" pageNumber="11">Harald Bruckner.</paragraph>
</caption>
<paragraph blockId="3.[227,1128,1224,1605]" box="[265,1127,1224,1248]" pageId="3" pageNumber="11">
Measurements of 
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(n = 10): BL 2.68–2.99, HL 0.62–0.66, HW 0.49–0.51, A 
<subScript attach="right" box="[1104,1112,1234,1248]" fontSize="6" pageId="3" pageNumber="11">2</subScript>
L
</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" box="[227,933,1252,1274]" pageId="3" pageNumber="11">0.24–0.26, PL 0.61–0.65, PW 1.19–1.28, MtL 1.01–1.12, AW 1.06–1.27.</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" pageId="3" pageNumber="11">
Colour (
<figureCitation box="[344,432,1280,1302]" captionStart="Figs" captionStartId="3.[265,309,1142,1164]" captionTargetBox="[227,1122,503,1108]" captionTargetId="figure-115@3.[227,1122,503,1109]" captionTargetPageId="3" captionText="Figs. 1–2: Habitus of Timasius alveus sp. n., male, in dorsal (1) and ventral (2) aspect. © Harald Bruckner." figureDoi="http://doi.org/10.5281/zenodo.7165324" httpUri="https://zenodo.org/record/7165324/files/figure.png" pageId="3" pageNumber="11">Figs 1–2</figureCitation>
): Trunk black. Buccula yellow. Antenna black. Legs from coxa to base of femora yellow; distal parts of femora turning to brown. Tibiae and tarsi black. Each cell of forewing with relatively small frosted mark at base; membrane with indistinct greyish spots.
</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" pageId="3" pageNumber="11">Pilosity: Dorsum of head, thorax, and corium with rather sparse, thin, golden, subcumbent pilosity, comparatively dense on anterior lobe of pronotum, mesoscutellum, and terminal vein of forewing. Venter and sides of body with thin, subcumbent, whitish pilosity.</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" pageId="3" pageNumber="11">
Structures: Body stout, with rather short antennae and legs. Head (
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) comparatively short; sides between anterior margin of eyes and small, rounded antennal tubercles slightly divergent. Preocular tubercles weakly developed. Middle furrow on posterior of head deep.
</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" box="[227,1127,1528,1550]" pageId="3" pageNumber="11">
Anteclypeus shiny, compressed, in lateral aspect convex. Buccula (
<figureCitation box="[884,944,1528,1550]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="3" pageNumber="11">Fig. 4</figureCitation>
) moderately wide,
</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" box="[227,1127,1556,1578]" pageId="3" pageNumber="11">with two round circular impression, posteriorly with two processes: dorsal one small and acute,</paragraph>
<paragraph blockId="3.[227,1128,1224,1605]" box="[227,1119,1583,1605]" pageId="3" pageNumber="11">slightly pointing to side, ventral one hardly longer, apically acute, slightly curved medially.</paragraph>
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<paragraph blockId="4.[227,1127,1046,1178]" box="[265,1127,1046,1068]" pageId="4" pageNumber="12">
Figs. 3–10: Habitus of 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[497,653,1046,1067]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="12" phylum="Arthropoda" rank="species" species="alveus" status="sp. n.">
<emphasis box="[497,653,1046,1067]" italics="true" pageId="4" pageNumber="12">Timasius alveus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[662,718,1046,1067]" pageId="4" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
, male: (3) head, frontal aspect; (4) head,
</paragraph>
</caption>
<paragraph blockId="4.[227,1127,1046,1178]" pageId="4" pageNumber="12">lateral aspect; (5) mesoscutellum and metanotal elevation, dorsal aspect; (6) mesoscutellum and metanotal elevation, lateral aspect; (7) pygophore, dorsal aspect; (8) proctiger, dorsal aspect; (9) right paramere, lateral (external) aspect; (10) left paramere, lateral (external)</paragraph>
<paragraph blockId="4.[227,1127,1046,1178]" box="[227,294,1157,1178]" pageId="4" pageNumber="12">aspect.</paragraph>
<paragraph blockId="4.[227,1127,1211,1620]" pageId="4" pageNumber="12">
Pronotum wide, sides with angular emargination; surface, except for anterolateral callosities and humeri, with deep punctures; anteriorly between paired swellings with three parallel, very deep longitudinal impressions. Metanotal elevation (
<figureCitation box="[748,841,1266,1289]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="4" pageNumber="12">Figs 5, 6</figureCitation>
) short, 2.0 times as wide as long, subtriangular; apex sharply pointed in dorsal aspect, in lateral aspect free tip appearing truncated. Forewing posteriorly reaching end of abdomen, laterally distant from connexival margins. Ventral thoracic carinae as characteristic for the genus (
<figureCitation box="[866,926,1349,1371]" captionStart="Figs" captionStartId="3.[265,309,1142,1164]" captionTargetBox="[227,1122,503,1108]" captionTargetId="figure-115@3.[227,1122,503,1109]" captionTargetPageId="3" captionText="Figs. 1–2: Habitus of Timasius alveus sp. n., male, in dorsal (1) and ventral (2) aspect. © Harald Bruckner." figureDoi="http://doi.org/10.5281/zenodo.7165324" httpUri="https://zenodo.org/record/7165324/files/figure.png" pageId="4" pageNumber="12">Fig. 2</figureCitation>
). Legs very slender,
</paragraph>
<paragraph blockId="4.[227,1127,1211,1620]" box="[227,1127,1377,1399]" pageId="4" pageNumber="12">not modified. Abdomen in dorsal aspect only slightly widened from base to midlength,</paragraph>
<paragraph blockId="4.[227,1127,1211,1620]" pageId="4" pageNumber="12">
posteriorly ovate, almost evenly rounded apically, only weakly truncated. Abdominal venter strongly modified (
<figureCitation box="[409,468,1432,1454]" captionStart="Figs" captionStartId="3.[265,309,1142,1164]" captionTargetBox="[227,1122,503,1108]" captionTargetId="figure-115@3.[227,1122,503,1109]" captionTargetPageId="3" captionText="Figs. 1–2: Habitus of Timasius alveus sp. n., male, in dorsal (1) and ventral (2) aspect. © Harald Bruckner." figureDoi="http://doi.org/10.5281/zenodo.7165324" httpUri="https://zenodo.org/record/7165324/files/figure.png" pageId="4" pageNumber="12">Fig. 2</figureCitation>
): sternum 3 with small, roundish tuft of erect setae at middle of hind margin; sternum 4 with broad pad of brown velvety pilosity; sterna 5–7 with broad, but shallow medial groove; groove margins laterally not crested, posteriorly sharply delimited by a raised transverse elevation of segment 8. Segment 8 short, constricted at midlength; anterior portion narrower than posterior portion which contains the genitalia; posterior part with a subterminal crest ventrally. Genitalia short and broad. Pygophore (
<figureCitation box="[751,810,1570,1592]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="4" pageNumber="12">Fig. 7</figureCitation>
) symmetrical, strongly widened,
</paragraph>
<paragraph blockId="4.[227,1127,1211,1620]" box="[227,1125,1598,1620]" pageId="4" pageNumber="12">ventrolaterally extended into rounded lobes, (postero-) dorsolaterally extended into broad,</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,1127,263,285]" pageId="5" pageNumber="13">
mesally curved and apically truncated lobes; distal part transverse, truncated. Proctiger (
<figureCitation box="[1088,1127,263,285]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="5" pageNumber="13">Fig.</figureCitation>
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" pageId="5" pageNumber="13">
8) almost symmentrical, small, laterally with short ventrolaterally directed processes; apex rounded, bearing six short, very stout setae. Parameres (
<figureCitation box="[765,863,318,340]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="5" pageNumber="13">Figs 9, 10</figureCitation>
) very small, slightly asymmetrical; left paramere with a larger dorsal tooth and with less developed setae than right paramere.
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[265,1126,401,423]" pageId="5" pageNumber="13">
<emphasis bold="true" box="[265,488,401,423]" pageId="5" pageNumber="13">Macropterous female</emphasis>
. Measurements of 
<typeStatus box="[684,776,401,422]" pageId="5" pageNumber="13">paratypes</typeStatus>
(n = 3): BL 2.72–2.82, HL 0.62–
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,1127,428,452]" pageId="5" pageNumber="13">
0.64, HW 0.49–0.53, A 
<subScript attach="right" box="[466,474,438,452]" fontSize="6" pageId="5" pageNumber="13">2</subScript>
L 0.25–0.27, PL 0.60–0.66, PW 1.18–1.22, MtL 1.03–1.09, AW
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,330,456,478]" pageId="5" pageNumber="13">1.08–1.15.</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[265,1126,484,506]" pageId="5" pageNumber="13">Colour, pilosity, and structures similar as in males. Abdominal sterna unmodified. Gono-</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,926,511,533]" pageId="5" pageNumber="13">coxa flat, unmodified. Proctiger small, with some moderately long setae.</paragraph>
<paragraph blockId="5.[227,1127,263,892]" pageId="5" pageNumber="13">
COMPARATIVE NOTES. 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[536,691,539,560]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="species" species="alveus" status="sp.n.">
<emphasis box="[536,691,539,560]" italics="true" pageId="5" pageNumber="13">Timasius alveus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[698,744,539,560]" pageId="5" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
clearly belongs to the 
<emphasis box="[968,1050,539,561]" italics="true" pageId="5" pageNumber="13">T. livens</emphasis>
species group as defined by Andersen (1981), Zettel &amp; Chen (2000), and above. 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[914,1068,566,588]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="species" species="alveus" status="sp. n.">
<emphasis box="[914,1068,566,588]" italics="true" pageId="5" pageNumber="13">Timasius alveus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1075,1127,567,588]" pageId="5" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" pageId="5" pageNumber="13">
shares a pointed ventral buccula tooth with 
<emphasis box="[666,751,595,616]" italics="true" pageId="5" pageNumber="13">T. minor</emphasis>
from 
<collectingCountry box="[816,898,594,616]" name="Pakistan" pageId="5" pageNumber="13">Pakistan</collectingCountry>
and 
<collectingCountry box="[951,1003,594,616]" name="India" pageId="5" pageNumber="13">India</collectingCountry>
, whereas all other species – described or undescribed, as far as known to the authors – possess truncated or broadly rounded ventral buccula teeth. The male of 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[764,851,649,671]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="species" species="alveus" status="sp.n.">
<emphasis box="[764,851,649,671]" italics="true" pageId="5" pageNumber="13">T. alveus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[859,905,650,671]" pageId="5" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
is unique in the group by both the sternal characteristics (
<figureCitation box="[563,621,677,699]" captionStart="Figs" captionStartId="3.[265,309,1142,1164]" captionTargetBox="[227,1122,503,1108]" captionTargetId="figure-115@3.[227,1122,503,1109]" captionTargetPageId="3" captionText="Figs. 1–2: Habitus of Timasius alveus sp. n., male, in dorsal (1) and ventral (2) aspect. © Harald Bruckner." figureDoi="http://doi.org/10.5281/zenodo.7165324" httpUri="https://zenodo.org/record/7165324/files/figure.png" pageId="5" pageNumber="13">Fig. 2</figureCitation>
) and the broad genitalia (
<figureCitation box="[867,959,677,699]" captionStart="Figs" captionStartId="4.[265,309,1046,1068]" captionTargetBox="[229,1120,258,1008]" captionTargetId="figure-1@4.[227,1122,257,1012]" captionTargetPageId="4" captionText="Figs. 3–10: Habitus of Timasius alveus sp. n., male: (3) head, frontal aspect; (4) head," figureDoi="http://doi.org/10.5281/zenodo.7165326" httpUri="https://zenodo.org/record/7165326/files/figure.png" pageId="5" pageNumber="13">Figs. 7, 8</figureCitation>
). The female can be differentiated from 
<emphasis box="[455,541,705,726]" italics="true" pageId="5" pageNumber="13">T. minor</emphasis>
by larger size (BL 
<quantity box="[745,883,705,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="5" pageNumber="13" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7 – 2.8 mm</quantity>
vs. 
<quantity box="[930,1068,704,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="5" pageNumber="13" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2 – 2.5 mm</quantity>
in 
<emphasis box="[1107,1127,705,726]" italics="true" pageId="5" pageNumber="13">T.</emphasis>
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,1127,732,754]" pageId="5" pageNumber="13">
<emphasis box="[227,284,733,754]" italics="true" pageId="5" pageNumber="13">minor</emphasis>
). 
<taxonomicName authority="Zettel &amp; Pangantihon, 2020" authorityName="Zettel &amp; Pangantihon" authorityYear="2020" box="[305,459,732,754]" class="Insecta" family="Hebridae" genus="Timasius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="13" phylum="Arthropoda" rank="species" species="alveus" status="sp.n.">
<emphasis box="[305,459,732,754]" italics="true" pageId="5" pageNumber="13">Timasius alveus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[466,512,732,753]" pageId="5" pageNumber="13" rank="species">sp.n.</taxonomicNameLabel>
is among the largest species of the 
<emphasis box="[846,926,732,754]" italics="true" pageId="5" pageNumber="13">T. livens</emphasis>
group, together with
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,1124,760,782]" pageId="5" pageNumber="13">
<emphasis box="[227,240,760,781]" italics="true" pageId="5" pageNumber="13">T</emphasis>
. 
<emphasis box="[255,360,760,781]" italics="true" pageId="5" pageNumber="13">tachamoae</emphasis>
Zettel, 2012 from 
<collectingCountry box="[553,610,760,782]" name="Nepal" pageId="5" pageNumber="13">Nepal</collectingCountry>
(BL 
<quantity box="[667,787,760,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="5" pageNumber="13" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
) and 
<emphasis box="[846,930,760,781]" italics="true" pageId="5" pageNumber="13">T. livens</emphasis>
(BL 
<quantity box="[987,1061,760,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="13" unit="mm" value="2.6">2.6 mm</quantity>
) from
</paragraph>
<paragraph blockId="5.[227,1127,263,892]" box="[227,320,787,809]" pageId="5" pageNumber="13">
<collectingCountry box="[227,316,787,809]" name="Thailand" pageId="5" pageNumber="13">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="13" type="etymology">
<paragraph blockId="5.[227,1127,263,892]" pageId="5" pageNumber="13">ETYMOLOGY. The species epithet is a noun in apposition; it is the Latin word for a shallow groove. The name refers to the large, round impression on the abdominal sterna of the male.</paragraph>
</subSubSection>
</treatment>
</document>