treatments-xml/data/03/CD/FC/03CDFC30D007FFF5971EFCEFFAB5FA54.xml

<document id="F08F2F7034901192FD4B3EBA44B53212" ID-CLB-Dataset="289477" ID-DOI="10.15553/c2023v781a1" ID-GBIF-Dataset="b65462fc-2200-4f6c-9dd7-0ca2a646fff4" ID-ISSN="2235-3658" ID-Zenodo-Dep="8110231" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1688393673680" checkinUser="felipe" docAuthor="Eggenberg, Stefan, Champoud, Luca, Leibundgut, Mary, Parisod, Christian, Wyss, Louisa &amp; Kozlowski, Gregor" docDate="2023" docId="03CDFC30D007FFF5971EFCEFFAB5FA54" docLanguage="en" docName="Candollea.78.1.1-9.pdf" docOrigin="Candollea 78 (1)" docSource="http://dx.doi.org/10.15553/c2023v781a1" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Calamagrostis lonana Eggenb. &amp; Leibundg." docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFF48448D005FFF29426FFCEFFF1FFF0" masterDocTitle="Calamagrostis lonana (Poaceae): a new grass species from the Pennine Alps (Switzerland)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1709127318818" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B86D50B36A24673E31EDC8D5F1795379">Calamagrostis lonana (Poaceae): a new grass species from the Pennine Alps (Switzerland)</mods:title>
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<mods:namePart id="BF5890E255D032DFE37FA21A9DBAFB7C">Eggenberg, Stefan</mods:namePart>
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<mods:namePart id="800027D70D4C59DCCDC9B80E2ECA9D65">Champoud, Luca</mods:namePart>
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<mods:date id="4F23C2465F82F1F585D51EB3D6180BCD">2023</mods:date>
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<subSubSection id="C37E1EADD007FFF0971EFCEFFA5CFC54" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BDB4D26D007FFF0971EFCEFFA35FCCA" blockId="2.[824,1476,801,826]" box="[824,1476,801,826]" pageId="2" pageNumber="3">
<heading id="D093FA4AD007FFF0971EFCEFFA35FCCA" box="[824,1476,801,826]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="4C6436A5D007FFF0971EFCEFFB00FCCA" authority="Eggenb. &amp; Leibundg." authorityName="Eggenb. &amp; Leibundg." box="[824,1265,801,826]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D007FFF0971EFCEFFBE7FCC9" bold="true" box="[824,1046,801,825]" italics="true" pageId="2" pageNumber="3">Calamagrostis lonana</emphasis>
Eggenb. &amp; Leibundg.
</taxonomicName>
, 
<emphasis id="B9109134D007FFF09120FCECFAA6FCCA" bold="true" box="[1286,1367,802,826]" pageId="2" pageNumber="3">sp. nov.</emphasis>
(
<figureCitation id="135F51A3D007FFF09142FCECFA6BFCCA" box="[1380,1434,802,826]" captionStart="Fig" captionStartId="4.[136,166,1864,1884]" captionTargetBox="[194,1428,324,1792]" captionTargetId="figure-21@4.[170,1440,305,1811]" captionTargetPageId="4" captionText="Fig. 2.– Calamagrostis lonana Eggenb.&amp; Leibundg. A. Habit with stolons within the moss layer; B. Inflorescence (contracted panicle); C. Section of inflorescence with adjacent panicle branches; D. Spikelet peduncle with glumes; E. Glume and palea with callus hairs and awn (dorsally inserted). [Drawings: S. Eggenberg]" figureDoi="http://doi.org/10.5281/zenodo.10722307" httpUri="https://zenodo.org/record/10722307/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
, 
<figureCitation id="135F51A3D007FFF0918CFCECFA49FCCA" box="[1450,1464,802,826]" captionStart="Fig" captionStartId="5.[113,142,1125,1145]" captionTargetBox="[114,1451,276,1101]" captionTargetId="figure-469@5.[639,1451,276,1101]" captionTargetPageId="5" captionText="Fig. 3. – Calamagrostis lonana Eggenb.&amp; Leibundg. A. Habit of the aerial parts with moss cushion of Warnstorfia exannulata (Schimp.) Loeske. B. Upper root space in the moss layer. [Photos: S. Eggenberg]" figureDoi="http://doi.org/10.5281/zenodo.10722311" httpUri="https://zenodo.org/record/10722311/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
).
</heading>
</paragraph>
<paragraph id="8BDB4D26D007FFF09778FC82FA5CFC54" blockId="2.[862,1475,844,932]" pageId="2" pageNumber="3">
<materialsCitation id="3B0C477BD007FFF09778FC82FA59FC54" collectingDate="2019-10-12" collectionCode="BERN!, G, SION" collectorName="Champoud &amp; Eggenberg" country="Switzerland" location="coffune d'Anniviers" pageId="2" pageNumber="3" specimenCode="G00414405" specimenCount="1" stateProvince="Valais" typeStatus="holotype">
<emphasis id="B9109134D007FFF09778FC82FB21FC94" bold="true" box="[862,1232,844,868]" pageId="2" pageNumber="3">
<typeStatus id="54DFF384D007FFF09778FC82FC27FC94" box="[862,982,844,868]" pageId="2" pageNumber="3" type="holotype">Holotypus</typeStatus>
: 
<collectingCountry id="F3730DB6D007FFF097F9FC82FB8FFC94" box="[991,1150,844,868]" name="Switzerland" pageId="2" pageNumber="3">SWITZERLAND</collectingCountry>
. 
<collectingRegion id="49A083C4D007FFF090A1FC82FB3AFC94" box="[1159,1227,844,868]" country="Switzerland" name="Valais" pageId="2" pageNumber="3">Valais</collectingRegion>
:
</emphasis>
<location id="8EBB1BFDD007FFF090F0FC82FA40FC94" LSID="urn:lsid:plazi:treatment:03CDFC30D007FFF5971EFCEFFAB5FA54:8EBB1BFDD007FFF090F0FC82FA40FC94" box="[1238,1457,844,868]" country="Switzerland" name="coffune d'Anniviers" pageId="2" pageNumber="3" stateProvince="Valais">coffune d’Anniviers</location>
, 
<location id="8EBB1BFDD007FFF09778FCA2FBEAFC74" LSID="urn:lsid:plazi:treatment:03CDFC30D007FFF5971EFCEFFAB5FA54:8EBB1BFDD007FFF09778FCA2FBEAFC74" box="[862,1051,876,900]" country="Switzerland" name="Plateau de Lona" pageId="2" pageNumber="3" stateProvince="Valais">Plateau de Lona</location>
, 2588 f, 
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<collectingDate id="EF9E920ED007FFF09084FCA2FAFBFC74" box="[1186,1290,876,900]" pageId="2" pageNumber="3" value="2019-10-12">12.X.2019</collectingDate>
</date>
, 
<emphasis id="B9109134D007FFF0910FFCA2FC03FC54" italics="true" pageId="2" pageNumber="3">
<collectorName id="269128F0D007FFF0910FFCA2FA50FC74" box="[1321,1441,876,900]" pageId="2" pageNumber="3">Champoud</collectorName>
&amp; 
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s.n.
</emphasis>
(
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; iso-: 
<collectionCode id="ED75D5E3D007FFF090B3FC42FB5AFC54" box="[1173,1195,908,932]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="2" pageNumber="3" type="Herbarium">G</collectionCode>
[
<specimenCode id="DBC2E55DD007FFF09091FC42FAC0FC54" box="[1207,1329,908,932]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="2" pageNumber="3" type="Herbarium">G00414405</specimenCode>
]!, 
<collectionCode id="ED75D5E3D007FFF09174FC42FA6DFC54" box="[1362,1436,908,932]" pageId="2" pageNumber="3">SION</collectionCode>
!)
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.
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<subSubSection id="C37E1EADD007FFF59778FC79FAB5FA54" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3" type="description">
<paragraph id="8BDB4D26D007FFF09778FC79FB1FFB5E" blockId="2.[862,1475,951,1199]" pageId="2" pageNumber="3">
<emphasis id="B9109134D007FFF09778FC79FB16FB5F" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C6436A5D007FFF09778FC79FAD4FC3F" authority="Eggenb. &amp; Leibundg." authorityName="Eggenb. &amp; Leibundg." box="[862,1317,951,975]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="lonana">Calamagrostis lonana Eggenb. &amp; Leibundg.</taxonomicName>
differs from all other Central European 
<taxonomicName id="4C6436A5D007FFF09068FC19FAEEFC1F" box="[1102,1311,983,1007]" pageId="2" pageNumber="3">Calamagrostis species</taxonomicName>
by its small size: plants are smaller than 
<quantity id="4C9CE0C3D007FFF09072FC39FB63FBFF" box="[1108,1170,1015,1039]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="cm" value="20.0">20 cm</quantity>
whereas other 
<taxonomicName id="4C6436A5D007FFF09108FC39FC6CFBDF" pageId="2" pageNumber="3">Calamagrostis species</taxonomicName>
reach (30 –) 
<quantity id="4C9CE0C3D007FFF0903AFBD9FB7CFBDF" box="[1052,1165,1046,1071]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="3" unit="cm" value="100.0" valueMax="150.0" valueMin="50.0">50 –150 cm</quantity>
high. Like 
<taxonomicName id="4C6436A5D007FFF09127FBD9FA90FBDF" box="[1281,1377,1047,1071]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="stricta">C. stricta</taxonomicName>
, its callus hairs are distinctly shorter than the lemma and the awn is not protruding from the spikelet but differs by its short panicle with very short branches, short and often glabrous ligule, almost glabrous glumes, and very short anthers
</emphasis>
.
</paragraph>
<paragraph id="8BDB4D26D007FFF19778FB19FD79FE7F" blockId="2.[824,1482,1239,1935]" lastBlockId="3.[113,764,278,399]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Perennial, stoloniferous, light to grey-green grass with loose but deep-reaching root systef, with extravaginal shoot initiation. 
<emphasis id="B9109134D007FFF09781FAD9FC19FADF" box="[935,1000,1303,1327]" italics="true" pageId="2" pageNumber="3">Culms</emphasis>
solitary, glabrous, (10–)13–18(–22) cf high, erect, hairy towards the inflorescence, with 1 – 3 glabrous nodes; sheaths 1 – 5 cf long, glabrous. 
<emphasis id="B9109134D007FFF090D0FA99FAB2FA9F" box="[1270,1347,1367,1391]" italics="true" pageId="2" pageNumber="3">Ligules</emphasis>
0.5 – 1 ff long, glabrous abaxially or pubescent towards the tip. 
<emphasis id="B9109134D007FFF091B5FAB9FC84FA5F" italics="true" pageId="2" pageNumber="3">Leaf blades</emphasis>
80(–120) × 2–3(–3.5) ff, scabrous adaxially, glabrous abaxially, fostly enrolled. 
<emphasis id="B9109134D007FFF09056FA79FAF1FA3F" box="[1136,1280,1463,1487]" italics="true" pageId="2" pageNumber="3">Inflorescences</emphasis>
a dense panicle, (2.4–)3.1 –3.7(–4) cf long, gradually spreading, soon contracting after anthesis; panicle branches 1– 3(–4) frof fain axis, 1 –6 ff long, short ciliate; spikelet stalks 0.5 – 2 ff long. 
<emphasis id="B9109134D007FFF09752F9F9FC3AF9BF" box="[884,971,1591,1615]" italics="true" pageId="2" pageNumber="3">Spikelets</emphasis>
2.5–3.2(–3.6) ff long, lanceolate, 1-flowered, light to grey-green, fostly with a purplish tinge towards the tip. 
<emphasis id="B9109134D007FFF09742F9B9FC43F97F" box="[868,946,1655,1679]" italics="true" pageId="2" pageNumber="3">Glumes</emphasis>
narrowly lanceolate, long acufinate, the lower 2.6–3.1 ff long, 1-veined, glabrous, the upper 2.7–3.4 ff long, 3-veined, the profinent fedian vein often ending in an erect tip, fifbriate dorsally, otherwise glabrous; callus short, rounded, covered with hairs 1–1.4 ff long; axillary process tiny, 0.3 – 0.5 ff long, hairy, sofetifes absent. 
<emphasis id="B9109134D007FFF0971EF8F6FC7CF8BF" box="[824,909,1848,1871]" italics="true" pageId="2" pageNumber="3">Lemmas</emphasis>
2.1–2.4(–2.7) ff long, alfost as long as the glufes, 5-veined, pubescent, ragged distally and ciliate like the veins; awn 1.5–1.9 ff long, straight, erect, ciliate, inserted c. 0.6 ff above the leffa base. 
<taxonomicName id="4C6436A5D006FFF1955AFED9FE4EFEDF" authorityName="Alekseev" authorityYear="1994" box="[380,447,279,303]" class="Insecta" family="Buprestidae" genus="Paleas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9109134D006FFF1955AFED9FE4EFEDF" box="[380,447,279,303]" italics="true" pageId="3" pageNumber="4">Paleas</emphasis>
</taxonomicName>
1.3 – 1.6 ff long, delicateskinned, glabrous. 
<emphasis id="B9109134D006FFF19563FEF9FE66FEBF" box="[325,407,311,335]" italics="true" pageId="3" pageNumber="4">Anthers</emphasis>
0.9 – 1.4 ff long, light yellow. 
<emphasis id="B9109134D006FFF19457FE99FF30FE9E" box="[113,193,343,366]" italics="true" pageId="3" pageNumber="4">Ovaries</emphasis>
0.2 ff long, refaining iffature, narrow, with two terfinal styles and feathery stigfas c. 1 ff long.
</paragraph>
<paragraph id="8BDB4D26D006FFF194B1FE79FECAFDFF" blockId="3.[113,764,439,527]" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF194B1FE79FEF2FE3F" box="[151,259,439,463]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
. – The epithet 
<taxonomicName id="4C6436A5D006FFF19583FE79FE1AFE3F" box="[421,491,439,463]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF19583FE79FE1AFE3F" box="[421,491,439,463]" italics="true" pageId="3" pageNumber="4">lonana</emphasis>
</taxonomicName>
refers to the nafe of the Lona plateau in the Pennine Alps (Valais), where the species was found in 2018.
</paragraph>
<paragraph id="8BDB4D26D006FFF194B1FDF9FD1EFC5F" blockId="3.[113,764,567,943]" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF194B1FDF9FEE9FDBF" box="[151,280,567,591]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
. – Based on fultiple visits of the Lona area, 
<taxonomicName id="4C6436A5D006FFF19457FD99FEB9FD9F" box="[113,328,599,623]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF19457FD99FEB9FD9F" box="[113,328,599,623]" italics="true" pageId="3" pageNumber="4">Calamagrostis lonana</emphasis>
</taxonomicName>
seefs to be restricted to the alluvial plain of the Torrent de Lona and has not been found elsewhere to date. The plain lies at an altitude of 2580 f and extends along the Torrent de Lona, which flows in a west-easterly direction. It is about 720 f long and 200 f wide at its widest point. The distribution area of 
<taxonomicName id="4C6436A5D006FFF1956BFD39FE44FCFF" box="[333,437,759,783]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF1956BFD39FE44FCFF" box="[333,437,759,783]" italics="true" pageId="3" pageNumber="4">C. lonana</emphasis>
</taxonomicName>
is located within this alluvial plain with a faxifuf extension of 600 × 
<specimenCount id="9D6286AFD006FFF19671FCD9FD6EFCDF" box="[599,671,791,815]" pageId="3" pageNumber="4" type="generic">150 f.</specimenCount>
Within this area, several subpopulations of 
<taxonomicName id="4C6436A5D006FFF195CCFCF9FDBFFCBF" box="[490,590,823,847]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF195CCFCF9FDBFFCBF" box="[490,590,823,847]" italics="true" pageId="3" pageNumber="4">C. lonana</emphasis>
</taxonomicName>
could be found. They are very unevenly distributed and dissifilar in density. It cannot be excluded that further localities will be reported frof other high altitude alluvial plains in the Pennine Alps.
</paragraph>
<paragraph id="8BDB4D26D006FFF194B1FC19FB99FD5F" blockId="3.[113,764,983,1231]" lastBlockId="3.[801,1452,279,1231]" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF194B1FC19FF15FC1F" box="[151,228,983,1007]" italics="true" pageId="3" pageNumber="4">Ecology</emphasis>
. – The large alluvial plain of Lona shows a fosaic of dimmerent habitat 
<typeStatus id="54DFF384D006FFF19561FC39FE8EFBFF" box="[327,383,1015,1039]" pageId="3" pageNumber="4">types</typeStatus>
which is typical in dynafic alpine alluvial plains (
<figureCitation id="135F51A3D006FFF1953DFBD9FE91FBDF" box="[283,352,1047,1071]" captionStart="Fig" captionStartId="7.[113,142,1192,1212]" captionTargetBox="[113,1451,276,1168]" captionTargetId="figure-389@7.[113,1451,276,1168]" captionTargetPageId="7" captionText="Fig. 4. – Alpine alluvial plain of Lona, with fens, alluvial and spring vegetation patches on both sides of the meandering Torrent de Lona." figureDoi="http://doi.org/10.5281/zenodo.8110239" httpUri="https://zenodo.org/record/8110239/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). In addition to vegetation-free sand and gravel banks, initial alluvial vegetation (
<emphasis id="B9109134D006FFF19664FBF9FF23FB9F" italics="true" pageId="3" pageNumber="4">Caricion bicolorisatrofuscae</emphasis>
), calcareous springs (
<emphasis id="B9109134D006FFF19596FB99FDC4FB9E" box="[432,565,1111,1134]" italics="true" pageId="3" pageNumber="4">Cratoneurion</emphasis>
) and transitions to fens and transitional bogs (
<emphasis id="B9109134D006FFF195AEFBB9FD0DFB7F" box="[392,764,1143,1167]" italics="true" pageId="3" pageNumber="4">Caricion fuscae, Caricion davallianae,</emphasis>
and 
<emphasis id="B9109134D006FFF194B8FB59FE96FB5F" box="[158,359,1175,1199]" italics="true" pageId="3" pageNumber="4">Caricion lasiocarpae</emphasis>
) can be observed. The vegetation classification of the occurrences of 
<taxonomicName id="4C6436A5D006FFF1959FFB79FD62FB3F" box="[441,659,1207,1231]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF1959FFB79FD62FB3F" box="[441,659,1207,1231]" italics="true" pageId="3" pageNumber="4">Calamagrostis lonana</emphasis>
</taxonomicName>
was done by the six releves shown in 
<tableCitation id="C6E6789DD006FFF1901DFED9FB76FEDF" box="[1083,1159,279,303]" captionStart="Table 1" captionStartId="3.[114,165,1277,1297]" captionTargetBox="[121,1345,1374,1940]" captionTargetId="graphics-615@3.[113,1452,1406,1908]" captionText="Table 1. – Six vegetation releves showing the co-occurrence of Calamagrostis lonana Eggenb.&amp; Leibundg. with other vascular plants. Species names according to JUILLERAT et al. (2017). [Abbreviations: imm. = immature; r = very low abundance; + = low abundance, cover &lt;5 %; 1 = high abundance, cover &lt;5 %; 2 = cover 5–25 %; 3 = cover 25–50%]" pageId="3" pageNumber="4">Table 1</tableCitation>
. The plot size was 10 square fetres for all releves. The abundance was estifated using the Braun-Blanquet cover scale. There is usually an extraordinarily high cover of the foss layer at the sites where 
<taxonomicName id="4C6436A5D006FFF1910DFEB9FA63FE7F" box="[1323,1426,375,399]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF1910DFEB9FA63FE7F" box="[1323,1426,375,399]" italics="true" pageId="3" pageNumber="4">C. lonana</emphasis>
</taxonomicName>
is growing, a feature that is quite striking and constant. The following foss species were identified: 
<taxonomicName id="4C6436A5D006FFF1908AFE76FBB0FE1F" authority="(Hedw.) G. Gaertn. et al." authorityName="G. Gaertn." baseAuthorityName="Hedw." class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="pseudotriquetrum">
<emphasis id="B9109134D006FFF1908AFE76FA5AFE3F" box="[1196,1451,439,463]" italics="true" pageId="3" pageNumber="4">Bryum pseudotriquetrum</emphasis>
(Hedw.) G. Gaertn. et al.
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF19069FE19FC9DFDFF" authority="(Schifp.) Kindb." authorityName="Kindb." baseAuthorityName="Schifp." class="Bryopsida" family="Amblystegiaceae" genus="Calliergon" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="giganteum">
<emphasis id="B9109134D006FFF19069FE19FAC5FE1F" box="[1103,1332,471,495]" italics="true" pageId="3" pageNumber="4">Calliergon giganteum</emphasis>
(Schifp.) Kindb.
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF19750FE39FAD8FDFF" authority="(Mitt.) Kindb. ex G. Roth" authorityName="G. Roth" baseAuthorityName="Mitt." box="[886,1321,503,527]" class="Bryopsida" family="Amblystegiaceae" genus="Calliergon" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="richardsonii">
<emphasis id="B9109134D006FFF19750FE39FBFCFDFF" box="[886,1037,503,527]" italics="true" pageId="3" pageNumber="4">C. richardsonii</emphasis>
(Mitt.) Kindb. ex G. Roth
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF19112FE39FB12FDDE" authority="(Hedw.) Lange &amp; C. E. O. Jensen" authorityName="Lange &amp; C. E. O. Jensen" baseAuthorityName="Hedw." class="Bryopsida" family="Amblystegiaceae" genus="Campylium" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="stellatum">
<emphasis id="B9109134D006FFF19112FE39FC8CFDDF" italics="true" pageId="3" pageNumber="4">Campylium stellatum</emphasis>
(Hedw.) Lange &amp; C.E.O. Jensen
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF190C9FDD9FC29FDBF" authority="(Brid.) Hedenas" authorityName="Hedenas" baseAuthorityName="Brid." class="Bryopsida" family="Amblystegiaceae" genus="Palustriella" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="falcata">
<emphasis id="B9109134D006FFF190C9FDD9FA5DFDDF" box="[1263,1452,535,559]" italics="true" pageId="3" pageNumber="4">Palustriella falcata</emphasis>
(Brid.) Hedenäs
</taxonomicName>
[dofinant], 
<taxonomicName id="4C6436A5D006FFF19056FDF9FC2EFD9F" authority="(Bruch &amp; Schifp.) Schifp." authorityName="Schifp." baseAuthorityName="Bruch &amp; Schifp." class="Bryopsida" family="Bartramiaceae" genus="Philonotis" kingdom="Plantae" order="Bryales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="calcarea">
<emphasis id="B9109134D006FFF19056FDF9FAC7FDBF" box="[1136,1334,567,591]" italics="true" pageId="3" pageNumber="4">Philonotis calcarea</emphasis>
(Bruch &amp; Schifp.) Schifp.
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF197CFFD99FAC1FD9F" authority="Hedw." authorityName="Hedw." box="[1001,1328,599,623]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="juniperinum">
<emphasis id="B9109134D006FFF197CFFD99FB15FD9F" box="[1001,1252,599,623]" italics="true" pageId="3" pageNumber="4">Polytrichum juniperinum</emphasis>
Hedw.
</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF1911CFD99FBA0FD7F" authority="(Schifp.) Hedenas" authorityName="Hedenas" authorityYear="1989" baseAuthorityName="Schifp." class="Bryopsida" family="Amblystegiaceae" genus="Scorpidium" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="cossonii">
<emphasis id="B9109134D006FFF1911CFD99FC83FD7F" italics="true" pageId="3" pageNumber="4">Scorpidium cossonii</emphasis>
(Schifp.) Hedenäs
</taxonomicName>
, and 
<emphasis id="B9109134D006FFF19084FDB9FAD6FD7F" box="[1186,1319,631,655]" italics="true" pageId="3" pageNumber="4">Warnstorf ia</emphasis>
<taxonomicName id="4C6436A5D006FFF19116FDB9FC2EFD5F" authority="(Schifp.) Loeske" authorityName="Loeske" authorityYear="1907" baseAuthorityName="Schifp." class="Bryopsida" family="Amblystegiaceae" genus="Warnstorfia" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="exannulata">
<emphasis id="B9109134D006FFF19116FDB9FA5BFD7F" box="[1328,1450,631,655]" italics="true" pageId="3" pageNumber="4">exannulata</emphasis>
(Schifp.) Loeske
</taxonomicName>
[dofinant].
</paragraph>
<paragraph id="8BDB4D26D006FFF19761FD79FB88FBFF" blockId="3.[801,1452,279,1231]" pageId="3" pageNumber="4">
Due to the stratification of the vegetation (dofinance of fosses) and the species cofposition of vascular plants and foss species, the assignfent to the habitat 
<typeStatus id="54DFF384D006FFF190D8FD39FADCFCFF" box="[1278,1325,759,783]" pageId="3" pageNumber="4">type</typeStatus>
“calcareous spring forfation” (
<emphasis id="B9109134D006FFF197C1FCD9FB9CFCDE" box="[999,1133,791,814]" italics="true" pageId="3" pageNumber="4">Cratoneurion</emphasis>
) is fost likely. Typical species are 
<emphasis id="B9109134D006FFF1976DFCF6FC80FC9F" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C6436A5D006FFF1976DFCF6FBABFCBF" box="[843,1114,823,847]" class="Bryopsida" family="Bryaceae" genus="Bryum" kingdom="Plantae" order="Bryales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="pseudotriquetrum">Bryum pseudotriquetrum</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF19040FCF9FAC5FCBF" box="[1126,1332,823,847]" class="Bryopsida" family="Amblystegiaceae" genus="Palustriella" kingdom="Plantae" order="Hypnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="falcata">Palustriella falcata</taxonomicName>
, 
<taxonomicName id="4C6436A5D006FFF19119FCF9FC80FC9F" class="Bryopsida" family="Bartramiaceae" genus="Philonotis" kingdom="Plantae" order="Bryales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="calcarea">Philonotis calcarea</taxonomicName>
</emphasis>
and the vascular plants 
<taxonomicName id="4C6436A5D006FFF1905FFC99FACDFC9F" box="[1145,1340,855,879]" class="Liliopsida" family="Poaceae" genus="Agrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="stolonifera">
<emphasis id="B9109134D006FFF1905FFC99FACDFC9F" box="[1145,1340,855,879]" italics="true" pageId="3" pageNumber="4">Agrostis stolonifera</emphasis>
</taxonomicName>
L., 
<taxonomicName id="4C6436A5D006FFF1914FFC99FC3BFC7F" authority="Gren." authorityName="Gren." class="Magnoliopsida" family="Brassicaceae" genus="Arabis" kingdom="Plantae" order="Brassicales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="subcoriacea">
<emphasis id="B9109134D006FFF1914FFC99FC61FC7F" italics="true" pageId="3" pageNumber="4">Arabis subcoriacea</emphasis>
Gren.
</taxonomicName>
, and 
<taxonomicName id="4C6436A5D006FFF19036FCB9FB3AFC7F" box="[1040,1227,887,911]" class="Magnoliopsida" family="Saxifragaceae" genus="Saxifraga" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="aizoides">
<emphasis id="B9109134D006FFF19036FCB9FB3AFC7F" box="[1040,1227,887,911]" italics="true" pageId="3" pageNumber="4">Saxifraga aizoides</emphasis>
</taxonomicName>
L. which were often found in the vicinity of the vegetation surveys. The habitat of 
<taxonomicName id="4C6436A5D006FFF19707FC79FC0DFC3F" box="[801,1020,951,975]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF19707FC79FC0DFC3F" box="[801,1020,951,975]" italics="true" pageId="3" pageNumber="4">Calamagrostis lonana</emphasis>
</taxonomicName>
can thus be relatively well characterised. It is alfost exclusively found in sites with low vascular plant cover and dofinance of fosses.
</paragraph>
<paragraph id="8BDB4D26D006FFF19761FBD9FA5AFB3F" blockId="3.[801,1452,279,1231]" pageId="3" pageNumber="4">
The bedrock across the Lona plateau shows a diversity of different sedifent 
<typeStatus id="54DFF384D006FFF1903CFBF9FBA5FBBF" box="[1050,1108,1079,1103]" pageId="3" pageNumber="4">types</typeStatus>
with both lifestone-rich and lifestone-poor sedifents. In the alluvial plain we feasured an average soil pH value of 6.5. The value was confirfed by laboratory feasurefents of soil safples. The high pH values of the organic soils of the alluvial plain can be explained by
</paragraph>
<paragraph id="8BDB4D26D006FFF19454FB33FAC0FAE1" blockId="3.[113,1329,1276,1348]" box="[114,1329,1276,1297]" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF19454FB33FF37FAE1" bold="true" box="[114,198,1277,1297]" pageId="3" pageNumber="4">Table 1. –</emphasis>
Six vegetation releves showing the co-occurrence of 
<taxonomicName id="4C6436A5D006FFF19695FB33FBC9FAE1" authority="Eggenb. &amp; Leibundg." authorityName="Eggenb. &amp; Leibundg." box="[691,1080,1276,1297]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D006FFF19695FB33FC8BFAE0" box="[691,890,1276,1297]" italics="true" pageId="3" pageNumber="4">Calamagrostis lonana</emphasis>
Eggenb.&amp; Leibundg.
</taxonomicName>
with other vascular plants.
</paragraph>
<paragraph id="8BDB4D26D006FFF19457FAD9FBA8FADB" blockId="3.[113,1329,1276,1348]" box="[113,1113,1303,1323]" pageId="3" pageNumber="4">Species names according to JUILLERAT et al. (2017). [Abbreviations: imm. = immature; r = very low abundance;</paragraph>
<paragraph id="8BDB4D26D006FFF19457FAFEFBE1FAB4" blockId="3.[113,1329,1276,1348]" box="[113,1040,1328,1348]" pageId="3" pageNumber="4">+ = low abundance, cover &lt;5 %; 1 = high abundance, cover &lt;5 %; 2 = cover 5–25 %; 3 = cover 25–50%]</paragraph>
<paragraph id="8BDB4D26D006FFF1945FFA90FCC8F864" pageId="3" pageNumber="4">
<table id="F964BF86D006000D945FFA90FAB0F864" box="[121,1345,1374,1940]" gridcols="7" gridrows="13" pageId="3" pageNumber="4">
<tr id="35544F64D006000D945FFA90FAB0FA82" box="[121,1345,1374,1394]" gridrow="0" pageId="3" pageNumber="4">
<th id="76852618D006000D945FFA90FE84FA82" box="[121,373,1374,1394]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Plot ID</th>
<th id="76852618D006000D95CAFA90FDEBFA82" box="[492,538,1374,1394]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">1</th>
<th id="76852618D006000D96ABFA90FD4BFA82" box="[653,698,1374,1394]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">2</th>
<th id="76852618D006000D9717FA90FCAFFA82" box="[817,862,1374,1394]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">3</th>
<th id="76852618D006000D97F4FA90FBF1FA82" box="[978,1024,1374,1394]" gridcol="4" gridrow="0" pageId="3" pageNumber="4">4</th>
<th id="76852618D006000D9055FA90FB58FA82" box="[1139,1193,1374,1394]" gridcol="5" gridrow="0" pageId="3" pageNumber="4">5</th>
<th id="76852618D006000D9135FA90FAB0FA82" box="[1299,1345,1374,1394]" gridcol="6" gridrow="0" pageId="3" pageNumber="4">6</th>
</tr>
<tr id="35544F64D006000D945FFA45FAB0FA6F" box="[121,1345,1419,1439]" gridrow="1" pageId="3" pageNumber="4">
<th id="76852618D006000D945FFA45FE84FA6F" box="[121,373,1419,1439]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Total cover</th>
<td id="76852618D006000D95CAFA45FDEBFA6F" box="[492,538,1419,1439]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">98 %</td>
<td id="76852618D006000D96ABFA45FD4BFA6F" box="[653,698,1419,1439]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">95%</td>
<td id="76852618D006000D9717FA45FCAFFA6F" box="[817,862,1419,1439]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">95%</td>
<td id="76852618D006000D97F4FA45FBF1FA6F" box="[978,1024,1419,1439]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">90%</td>
<td id="76852618D006000D9055FA45FB58FA6F" box="[1139,1193,1419,1439]" gridcol="5" gridrow="1" pageId="3" pageNumber="4">100%</td>
<td id="76852618D006000D9135FA45FAB0FA6F" box="[1299,1345,1419,1439]" gridcol="6" gridrow="1" pageId="3" pageNumber="4">95 %</td>
</tr>
<tr id="35544F64D006000D945FFA76FAB0FA3C" box="[121,1345,1464,1484]" gridrow="2" pageId="3" pageNumber="4">
<th id="76852618D006000D945FFA76FE84FA3C" box="[121,373,1464,1484]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Cover of herb layer</th>
<td id="76852618D006000D95CAFA76FDEBFA3C" box="[492,538,1464,1484]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">10 %</td>
<td id="76852618D006000D96ABFA76FD4BFA3C" box="[653,698,1464,1484]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">15 %</td>
<td id="76852618D006000D9717FA76FCAFFA3C" box="[817,862,1464,1484]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">20%</td>
<td id="76852618D006000D97F4FA76FBF1FA3C" box="[978,1024,1464,1484]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">15 %</td>
<td id="76852618D006000D9055FA76FB58FA3C" box="[1139,1193,1464,1484]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">30%</td>
<td id="76852618D006000D9135FA76FAB0FA3C" box="[1299,1345,1464,1484]" gridcol="6" gridrow="2" pageId="3" pageNumber="4">15 %</td>
</tr>
<tr id="35544F64D006000D945FFA29FAB0FA0B" box="[121,1345,1511,1531]" gridrow="3" pageId="3" pageNumber="4">
<th id="76852618D006000D945FFA29FE84FA0B" box="[121,373,1511,1531]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Cover of moss layer</th>
<td id="76852618D006000D95CAFA29FDEBFA0B" box="[492,538,1511,1531]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">95%</td>
<td id="76852618D006000D96ABFA29FD4BFA0B" box="[653,698,1511,1531]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">95%</td>
<td id="76852618D006000D9717FA29FCAFFA0B" box="[817,862,1511,1531]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">92 %</td>
<td id="76852618D006000D97F4FA29FBF1FA0B" box="[978,1024,1511,1531]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">85%</td>
<td id="76852618D006000D9055FA29FB58FA0B" box="[1139,1193,1511,1531]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">95%</td>
<td id="76852618D006000D9135FA29FAB0FA0B" box="[1299,1345,1511,1531]" gridcol="6" gridrow="3" pageId="3" pageNumber="4">95 %</td>
</tr>
<tr id="35544F64D006000D945FF9DBFAB0F9DA" box="[121,1345,1557,1578]" gridrow="4" pageId="3" pageNumber="4">
<th id="76852618D006000D945FF9DBFE84F9DA" box="[121,373,1557,1578]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF9DBFEBCF9DA" box="[121,333,1557,1578]" italics="true" pageId="3" pageNumber="4">Calamagrostis lonana</emphasis>
</th>
<td id="76852618D006000D95CAF9DBFDEBF9DA" box="[492,538,1557,1578]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">+</td>
<td id="76852618D006000D96ABF9DBFD4BF9DA" box="[653,698,1557,1578]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">1</td>
<td id="76852618D006000D9717F9DBFCAFF9DA" box="[817,862,1557,1578]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">2</td>
<td id="76852618D006000D97F4F9DBFBF1F9DA" box="[978,1024,1557,1578]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">2</td>
<td id="76852618D006000D9055F9DBFB58F9DA" box="[1139,1193,1557,1578]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">+</td>
<td id="76852618D006000D9135F9DBFAB0F9DA" box="[1299,1345,1557,1578]" gridcol="6" gridrow="4" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="35544F64D006000D945FF98DFAB0F9A7" box="[121,1345,1603,1623]" gridrow="5" pageId="3" pageNumber="4">
<th id="76852618D006000D945FF98DFE84F9A7" box="[121,373,1603,1623]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF98DFE84F9A7" box="[121,373,1603,1623]" italics="true" pageId="3" pageNumber="4">Eriophorum angustifolium</emphasis>
</th>
<td id="76852618D006000D95CAF98DFDEBF9A7" box="[492,538,1603,1623]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">1</td>
<td id="76852618D006000D96ABF98DFD4BF9A7" box="[653,698,1603,1623]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">+</td>
<td id="76852618D006000D9717F98DFCAFF9A7" box="[817,862,1603,1623]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">+</td>
<td id="76852618D006000D97F4F98DFBF1F9A7" box="[978,1024,1603,1623]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">+</td>
<td id="76852618D006000D9055F98DFB58F9A7" box="[1139,1193,1603,1623]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">r</td>
<td id="76852618D006000D9135F98DFAB0F9A7" box="[1299,1345,1603,1623]" gridcol="6" gridrow="5" pageId="3" pageNumber="4">1</td>
</tr>
<tr id="35544F64D006000D945FF9BEFAB0F974" box="[121,1345,1648,1668]" gridrow="6" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="76852618D006000D945FF9BEFE84F974" box="[121,373,1648,1668]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Eriophorum scheuchzeri</th>
<td id="76852618D006000D9717F9BEFCAFF974" box="[817,862,1648,1668]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">2</td>
<td id="76852618D006000D97F4F9BEFBF1F974" box="[978,1024,1648,1668]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">1</td>
</tr>
<tr id="35544F64D006000D945FF953FAB0F942" box="[121,1345,1693,1714]" gridrow="7" pageId="3" pageNumber="4" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76852618D006000D945FF953FE84F942" box="[121,373,1693,1714]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF950FE90F941" box="[121,353,1693,1714]" italics="true" pageId="3" pageNumber="4">Eleocharis quinqueflora</emphasis>
</th>
<td id="76852618D006000D95CAF953FDEBF942" box="[492,538,1693,1714]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">1</td>
<td id="76852618D006000D96ABF953FD4BF942" box="[653,698,1693,1714]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">2</td>
</tr>
<tr id="35544F64D006000D945FF905FAB0F92F" box="[121,1345,1739,1759]" gridrow="8" pageId="3" pageNumber="4" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76852618D006000D945FF905FE84F92F" box="[121,373,1739,1759]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF905FEF6F92F" box="[121,263,1739,1759]" italics="true" pageId="3" pageNumber="4">Carex rostrata</emphasis>
</th>
<td id="76852618D006000D95CAF905FDEBF92F" box="[492,538,1739,1759]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">1</td>
<td id="76852618D006000D9055F905FB58F92F" box="[1139,1193,1739,1759]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">3</td>
</tr>
<tr id="35544F64D006000D945FF936FAB0F8FC" box="[121,1345,1784,1804]" gridrow="9" pageId="3" pageNumber="4" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76852618D006000D945FF936FE84F8FC" box="[121,373,1784,1804]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF936FEA5F8FC" box="[121,340,1784,1804]" italics="true" pageId="3" pageNumber="4">Equisetum variegatum</emphasis>
</th>
<td id="76852618D006000D95CAF936FDEBF8FC" box="[492,538,1784,1804]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">+</td>
</tr>
<tr id="35544F64D006000D945FF8E8FAB0F8CA" box="[121,1345,1830,1850]" gridrow="10" pageId="3" pageNumber="4" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76852618D006000D945FF8E8FE84F8CA" box="[121,373,1830,1850]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF8E8FEE8F8CA" box="[121,281,1830,1850]" italics="true" pageId="3" pageNumber="4">Juncus triglumis</emphasis>
</th>
<td id="76852618D006000D95CAF8E8FDEBF8CA" box="[492,538,1830,1850]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">1</td>
<td id="76852618D006000D9135F8E8FAB0F8CA" box="[1299,1345,1830,1850]" gridcol="6" gridrow="10" pageId="3" pageNumber="4">+</td>
</tr>
<tr id="35544F64D006000D945FF89DFAB0F897" box="[121,1345,1875,1895]" gridrow="11" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76852618D006000D945FF89DFE84F897" box="[121,373,1875,1895]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">
<emphasis id="B9109134D006FFF1945FF89DFEB6F897" box="[121,327,1875,1895]" italics="true" pageId="3" pageNumber="4">Polygonum viviparum</emphasis>
</th>
<td id="76852618D006000D9717F89DFCAFF897" box="[817,862,1875,1895]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">r</td>
</tr>
<tr id="35544F64D006000D945FF84EFAB0F864" box="[121,1345,1920,1940]" gridrow="12" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76852618D006000D945FF84EFE84F864" box="[121,373,1920,1940]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">
<taxonomicName id="4C6436A5D006FFF1945FF84EFF0BF864" box="[121,250,1920,1940]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="hastata">
<emphasis id="B9109134D006FFF1945FF84EFF0BF864" box="[121,250,1920,1940]" italics="true" pageId="3" pageNumber="4">Salix hastata</emphasis>
</taxonomicName>
(imm.)
</th>
<td id="76852618D006000D9717F84EFCAFF864" box="[817,862,1920,1940]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">r</td>
</tr>
</table>
</paragraph>
<caption id="DF1B1DAED001FFF694AEF886FE9DF858" ID-DOI="http://doi.org/10.5281/zenodo.10722307" ID-Zenodo-Dep="10722307" httpUri="https://zenodo.org/record/10722307/files/figure.png" pageId="4" pageNumber="5" startId="4.[136,166,1864,1884]" targetBox="[194,1428,324,1792]" targetPageId="4">
<paragraph id="8BDB4D26D001FFF694AEF886FE9DF858" blockId="4.[136,1353,1862,1960]" pageId="4" pageNumber="5">
<emphasis id="B9109134D001FFF694AEF886FF3CF8AC" bold="true" box="[136,205,1864,1884]" pageId="4" pageNumber="5">Fig. 2.–</emphasis>
<taxonomicName id="4C6436A5D001FFF694F4F886FD55F8AC" authority="Eggenb. &amp; Leibundg. A. Habit" box="[210,676,1862,1884]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D001FFF694F4F886FE68F8AA" box="[210,409,1862,1884]" italics="true" pageId="4" pageNumber="5">Calamagrostis lonana</emphasis>
Eggenb.&amp; Leibundg. 
<emphasis id="B9109134D001FFF6967CF886FD9CF8AC" bold="true" box="[602,621,1864,1884]" pageId="4" pageNumber="5">A.</emphasis>
Habit
</taxonomicName>
with stolons within the moss layer; 
<emphasis id="B9109134D001FFF697CBF886FBF1F8AC" bold="true" box="[1005,1024,1864,1884]" pageId="4" pageNumber="5">B.</emphasis>
Inflorescence (contracted panicle); 
<emphasis id="B9109134D001FFF694AEF8AFFF6AF885" bold="true" box="[136,155,1889,1909]" pageId="4" pageNumber="5">C.</emphasis>
Section of inflorescence with adjacent panicle branches; 
<emphasis id="B9109134D001FFF69694F8AFFD34F885" bold="true" box="[690,709,1889,1909]" pageId="4" pageNumber="5">D.</emphasis>
Spikelet peduncle with glumes; 
<emphasis id="B9109134D001FFF697C8F8AFFBF0F885" bold="true" box="[1006,1025,1889,1909]" pageId="4" pageNumber="5">E.</emphasis>
Glume and palea with callus hairs and awn (dorsally inserted). [Drawings: S. Eggenberg]
</paragraph>
</caption>
<caption id="DF1B1DAED000FFF79457FBABFEB0FB5B" ID-DOI="http://doi.org/10.5281/zenodo.10722311" ID-Zenodo-Dep="10722311" httpUri="https://zenodo.org/record/10722311/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,142,1125,1145]" targetBox="[114,1451,276,1101]" targetPageId="5">
<paragraph id="8BDB4D26D000FFF79457FBABFEB0FB5B" blockId="5.[113,1409,1123,1195]" pageId="5" pageNumber="6">
<emphasis id="B9109134D000FFF79457FBABFF46FB89" bold="true" box="[113,183,1125,1145]" pageId="5" pageNumber="6">Fig. 3. –</emphasis>
<taxonomicName id="4C6436A5D000FFF7949DFBABFD7FFB89" authority="Eggenb. &amp; Leibundg. A. Habit" box="[187,654,1123,1145]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D000FFF7949DFBABFE73FB87" box="[187,386,1123,1145]" italics="true" pageId="5" pageNumber="6">Calamagrostis lonana</emphasis>
Eggenb.&amp; Leibundg. 
<emphasis id="B9109134D000FFF79665FBABFDA6FB89" bold="true" box="[579,599,1125,1145]" pageId="5" pageNumber="6">A.</emphasis>
Habit
</taxonomicName>
of the aerial parts with moss cushion of 
<taxonomicName id="4C6436A5D000FFF79025FBABFF74FB62" authority="(Schimp.) Loeske. B." authorityName="Loeske. B." baseAuthorityName="Schimp." class="Bryopsida" family="Amblystegiaceae" genus="Warnstorfia" kingdom="Plantae" order="Hypnales" pageId="5" pageNumber="6" phylum="Bryophyta" rank="species" species="exannulata">
<emphasis id="B9109134D000FFF79025FBABFB2BFB87" box="[1027,1242,1123,1145]" italics="true" pageId="5" pageNumber="6">Warnstorfia exannulata</emphasis>
(Schimp.) Loeske. 
<emphasis id="B9109134D000FFF79457FBB0FF74FB62" bold="true" box="[113,133,1150,1170]" pageId="5" pageNumber="6">B.</emphasis>
</taxonomicName>
Upper root space in the moss layer. [Photos: S. Eggenberg]
</paragraph>
</caption>
<paragraph id="8BDB4D26D000FFF79457FB02FDE5F974" blockId="5.[113,765,1228,1956]" pageId="5" pageNumber="6">
widespread calcareous sedifents in the area. The exafined soil profiles of 
<taxonomicName id="4C6436A5D000FFF7952FFB22FE11FAF4" box="[265,480,1260,1284]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D000FFF7952FFB22FE11FAF4" box="[265,480,1260,1284]" italics="true" pageId="5" pageNumber="6">Calamagrostis lonana</emphasis>
</taxonomicName>
growth sites always show a sifilar structure: a 10–15 cf thick foss layer (
<figureCitation id="135F51A3D000FFF79669FAC2FD7AFAD4" box="[591,651,1292,1316]" captionStart="Fig" captionStartId="5.[113,142,1125,1145]" captionTargetBox="[114,1451,276,1101]" captionTargetId="figure-469@5.[639,1451,276,1101]" captionTargetPageId="5" captionText="Fig. 3. – Calamagrostis lonana Eggenb.&amp; Leibundg. A. Habit of the aerial parts with moss cushion of Warnstorfia exannulata (Schimp.) Loeske. B. Upper root space in the moss layer. [Photos: S. Eggenberg]" figureDoi="http://doi.org/10.5281/zenodo.10722311" httpUri="https://zenodo.org/record/10722311/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) is directly followed by a layer of grey alluvial loaf fore than 50 cf deep, interfixed with dead fosses probably frof earlier inundations. The pH value refains unchanged over the entire profile depth (pH 6.5). Roots of 
<taxonomicName id="4C6436A5D000FFF79513FA43FE67FA54" box="[309,406,1420,1444]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D000FFF79513FA43FE67FA54" box="[309,406,1420,1444]" italics="true" pageId="5" pageNumber="6">C. lonana</emphasis>
</taxonomicName>
were found both in the foss layer (
<figureCitation id="135F51A3D000FFF7945FFA62FF47FA34" box="[121,182,1452,1476]" captionStart="Fig" captionStartId="5.[113,142,1125,1145]" captionTargetBox="[114,1451,276,1101]" captionTargetId="figure-469@5.[639,1451,276,1101]" captionTargetPageId="5" captionText="Fig. 3. – Calamagrostis lonana Eggenb.&amp; Leibundg. A. Habit of the aerial parts with moss cushion of Warnstorfia exannulata (Schimp.) Loeske. B. Upper root space in the moss layer. [Photos: S. Eggenberg]" figureDoi="http://doi.org/10.5281/zenodo.10722311" httpUri="https://zenodo.org/record/10722311/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) and in the alluvial loaf and can be detected down to a depth of about 40 cf. The root fass of 
<taxonomicName id="4C6436A5D000FFF7962EFA02FD65FA14" box="[520,660,1484,1508]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B9109134D000FFF7962EFA02FD65FA14" box="[520,660,1484,1508]" italics="true" pageId="5" pageNumber="6">Calamagrostis</emphasis>
</taxonomicName>
in a fore closely exafined soil safple was between 
<quantity id="4C9CE0C3D000FFF79662FA22FD72F9F5" box="[580,643,1516,1541]" metricMagnitude="-4" metricUnit="kg" metricValue="1.1" pageId="5" pageNumber="6" unit="g" value="0.11">0.11 g</quantity>
(top layer) and 
<quantity id="4C9CE0C3D000FFF79486F9C2FF15F9D5" box="[160,228,1548,1573]" metricMagnitude="-5" metricUnit="kg" metricValue="2.0" pageId="5" pageNumber="6" unit="g" value="0.02">0.02 g</quantity>
or 
<quantity id="4C9CE0C3D000FFF7952FF9C2FEBCF9D5" box="[265,333,1548,1573]" metricMagnitude="-5" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="6" unit="g" value="0.04">0.04 g</quantity>
(lower layers) per 
<quantity id="4C9CE0C3D000FFF79632F9C2FDBEF9D5" box="[532,591,1548,1573]" metricMagnitude="-1" metricUnit="kg" metricValue="1.0" pageId="5" pageNumber="6" unit="g" value="100.0">100 g</quantity>
of soil faterial. Whether during inspections at the beginning of August, in Septefber or at the beginning of October the soil was always found to be cofpletely water saturated.
</paragraph>
<paragraph id="8BDB4D26D000FFF794B1F942FC8BFAB5" blockId="5.[113,765,1228,1956]" lastBlockId="5.[801,1452,1228,1349]" pageId="5" pageNumber="6">The clifate conditions are typical for dry central alpine areas with annual precipitation suf of only approx. 1000 ff (METEOSWISS, 2022). Most of the annual precipitation falls in suffer, with a faxifuf in July. The annual fean tefperatures of around 0° C are characteristic of the alpine belt, fonthly averages rise to 6° C in suffer (METEOSWISS, 2022). However, the basin forf of the alluvial plain regularly leads to the forfation of cold air depositions, so that tefperatures in the alluvial plain are probably well below the fean tefperatures of the area. Geoforphological landforfs also reveal periglacial conditions across the whole Lona plateau. Mean tefperatures are low and frost can occur throughout the year.</paragraph>
<paragraph id="8BDB4D26D000FFF79761FAA2FB20FA54" blockId="5.[801,1451,1388,1444]" pageId="5" pageNumber="6">
<emphasis id="B9109134D000FFF79761FAA2FC41FA74" box="[839,944,1388,1412]" italics="true" pageId="5" pageNumber="6">Phenology</emphasis>
. – Flowering late suffer (frof August) and fruiting early autufn (frof Septefber).
</paragraph>
<paragraph id="8BDB4D26D000FFF49761FA02FED6FEBF" blockId="5.[801,1452,1484,1957]" lastBlockId="6.[136,786,279,335]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B9109134D000FFF79761FA02FC69FA14" box="[839,920,1484,1508]" italics="true" pageId="5" pageNumber="6">Threats</emphasis>
. – It can be assufed that the high plateaus of Lona have been used for grazing by sheep and cattle for a long tife (BÄTZING, 2015). The herds can fove freely, there is no fencing. Frof late suffer, when the slopes begin to dry out, the herds often gather in the green and fresh alluvial plains and the alluvial plain of the Torrent de Lona is exposed to increased trafpling in late August and Septefber. Therefore, the question arises to what extent the population of 
<taxonomicName id="4C6436A5D000FFF79707F902FC09F914" box="[801,1016,1740,1764]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D000FFF79707F902FC09F914" box="[801,1016,1740,1764]" italics="true" pageId="5" pageNumber="6">Calamagrostis lonana</emphasis>
</taxonomicName>
fight be seriously dafaged by trafpling and grazing. The trafpling ifpact on the plain is obvious, and tracks refain for a long tife. The trafpling tracks reveal a lower disturbance in fossy and 
<taxonomicName id="4C6436A5D000FFF79059F8E3FB1EF8B4" authorityName="de Jussieu" authorityYear="1789" box="[1151,1263,1837,1860]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">
<emphasis id="B9109134D000FFF79059F8E3FB1EF8B4" box="[1151,1263,1837,1860]" italics="true" pageId="5" pageNumber="6">Cyperaceae</emphasis>
</taxonomicName>
-dofinated parts of the plain, the sites where 
<taxonomicName id="4C6436A5D000FFF79047F883FB3BF894" box="[1121,1226,1868,1892]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D000FFF79047F883FB3BF894" box="[1121,1226,1868,1892]" italics="true" pageId="5" pageNumber="6">C. lonana</emphasis>
</taxonomicName>
thrives fost. These parts seef to be less attractive for the herds. Judging frof the current situation, it does not seef necessary to fence out the alluvial plain. However, long-terf fonitoring of trafpling is recoffended.
</paragraph>
<paragraph id="8BDB4D26D003FFF49488FEB6FE4BFC3F" blockId="6.[136,788,375,975]" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF49488FEB6FF1CFE7F" box="[174,237,376,399]" italics="true" pageId="6" pageNumber="7">Notes.</emphasis>
– With its short, hairy axillary process, short callus hairs, and long leffas, 
<taxonomicName id="4C6436A5D003FFF49582FE59FD63FE5F" box="[420,658,407,431]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF49582FE59FD63FE5F" box="[420,658,407,431]" italics="true" pageId="6" pageNumber="7">Calamagrostis lonana</emphasis>
</taxonomicName>
belongs to subgenus 
<taxonomicName id="4C6436A5D003FFF494D6FE76FEBEFE3F" box="[240,335,440,463]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="9" phylum="Tracheophyta" rank="section" section="Deyeuxia">
<emphasis id="B9109134D003FFF494D6FE76FEBEFE3F" box="[240,335,440,463]" italics="true" pageId="6" pageNumber="7">Deyeuxia</emphasis>
</taxonomicName>
. In central Europe, this subgenus includes the species 
<taxonomicName id="4C6436A5D003FFF49525FE19FE6EFE1F" box="[259,415,471,495]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="arundinacea">
<emphasis id="B9109134D003FFF49525FE19FE6EFE1F" box="[259,415,471,495]" italics="true" pageId="6" pageNumber="7">C. arundinacea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6436A5D003FFF4958DFE19FDF2FE1F" box="[427,515,471,495]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="varia">
<emphasis id="B9109134D003FFF4958DFE19FDF2FE1F" box="[427,515,471,495]" italics="true" pageId="6" pageNumber="7">C. varia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C6436A5D003FFF4961BFE19FD6DFE1F" box="[573,668,471,495]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF4961BFE19FD6DFE1F" box="[573,668,471,495]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
</taxonomicName>
. The short, straight awns show a close relationship to 
<taxonomicName id="4C6436A5D003FFF49661FE39FD55FDFF" box="[583,676,503,527]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF49661FE39FD55FDFF" box="[583,676,503,527]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
</taxonomicName>
(GREENE, 1980; ESCALONA, 1988), but it differs frof this species in having sfaller habit, shorter spikelets with shorter callus hairs, and shorter anthers (
<tableCitation id="C6E6789DD003FFF49582FD99FE07FD9F" box="[420,502,599,623]" captionStart="Table 2" captionStartId="6.[136,187,1009,1029]" captionTargetBox="[144,1410,1105,1941]" captionTargetId="graphics-469@6.[136,1474,1137,1909]" captionText="Table 2. – Comparison of morphological characters between Calamagrostis lonana Eggenb.&amp; Leibundg. (own measurements) and the Central European C. stricta (Timm) Koeler (measurements retrieved from HESS et al., 1967–1972;QUINGER, 1987; CONERT, 1998; MÜLLER &amp; al., 2021; PAROLLY &amp; ROHWER, 2016)." httpUri="http://table.plazi.org/id/DF1B1DAED003FFF494AEFC3FFDD4FBC7" pageId="6" pageNumber="7" tableUuid="DF1B1DAED003FFF494AEFC3FFDD4FBC7">Table 2</tableCitation>
). As in 
<taxonomicName id="4C6436A5D003FFF4966FFD99FD15FD9F" box="[585,740,599,623]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="arundinacea">
<emphasis id="B9109134D003FFF4966FFD99FD15FD9F" box="[585,740,599,623]" italics="true" pageId="6" pageNumber="7">C. arundinacea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6436A5D003FFF494AEFDB9FF2EFD7F" box="[136,223,631,655]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="varia">
<emphasis id="B9109134D003FFF494AEFDB9FF2EFD7F" box="[136,223,631,655]" italics="true" pageId="6" pageNumber="7">C. varia</emphasis>
</taxonomicName>
, the spikelets have an axillary process, which distinguishes it frof other Central European 
<taxonomicName id="4C6436A5D003FFF49609FD59FCE2FD5F" box="[559,787,663,687]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9109134D003FFF49609FD59FD4FFD5F" box="[559,702,663,687]" italics="true" pageId="6" pageNumber="7">Calamagrostis</emphasis>
species.
</taxonomicName>
However, this axillary process is very short in 
<taxonomicName id="4C6436A5D003FFF49648FD79FD3AFD3F" box="[622,715,695,719]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF49648FD79FD3AFD3F" box="[622,715,695,719]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
</taxonomicName>
(up to 0.5 ff long). 
<taxonomicName id="4C6436A5D003FFF49539FD19FE04FD1F" box="[287,501,727,751]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF49539FD19FE04FD1F" box="[287,501,727,751]" italics="true" pageId="6" pageNumber="7">Calamagrostis lonana</emphasis>
</taxonomicName>
dimmers frof 
<taxonomicName id="4C6436A5D003FFF4965DFD19FCE2FD1F" box="[635,787,727,751]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="arundinacea">
<emphasis id="B9109134D003FFF4965DFD19FCE2FD1F" box="[635,787,727,751]" italics="true" pageId="6" pageNumber="7">C. arundinacea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6436A5D003FFF49493FD39FEFBFCFF" box="[181,266,759,783]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="varia">
<emphasis id="B9109134D003FFF49493FD39FEFBFCFF" box="[181,266,759,783]" italics="true" pageId="6" pageNumber="7">C. varia</emphasis>
</taxonomicName>
by the short, straight, dorsal awn, which is usually shorter than the leffa. In 
<taxonomicName id="4C6436A5D003FFF49590FCD9FDA5FCDF" box="[438,596,791,815]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="arundinacea">
<emphasis id="B9109134D003FFF49590FCD9FDA5FCDF" box="[438,596,791,815]" italics="true" pageId="6" pageNumber="7">C. arundinacea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6436A5D003FFF496ADFCD9FD12FCDF" box="[651,739,791,815]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="varia">
<emphasis id="B9109134D003FFF496ADFCD9FD12FCDF" box="[651,739,791,815]" italics="true" pageId="6" pageNumber="7">C. varia</emphasis>
</taxonomicName>
, the awn is basally inserted, geniculated, and largely exceeding the leffa. 
<taxonomicName id="4C6436A5D003FFF494C6FC99FE37FC9F" box="[224,454,855,879]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF494C6FC99FE37FC9F" box="[224,454,855,879]" italics="true" pageId="6" pageNumber="7">Calamagrostis lonana</emphasis>
</taxonomicName>
dimmers frof all other Central European species by its sfall habit, which it also faintains in cultivation conditions. Ifportant forphological characteristics are cofpared in 
<tableCitation id="C6E6789DD003FFF49542FC79FE47FC3F" box="[356,438,951,975]" captionStart="Table 2" captionStartId="6.[136,187,1009,1029]" captionTargetBox="[144,1410,1105,1941]" captionTargetId="graphics-469@6.[136,1474,1137,1909]" captionText="Table 2. – Comparison of morphological characters between Calamagrostis lonana Eggenb.&amp; Leibundg. (own measurements) and the Central European C. stricta (Timm) Koeler (measurements retrieved from HESS et al., 1967–1972;QUINGER, 1987; CONERT, 1998; MÜLLER &amp; al., 2021; PAROLLY &amp; ROHWER, 2016)." httpUri="http://table.plazi.org/id/DF1B1DAED003FFF494AEFC3FFDD4FBC7" pageId="6" pageNumber="7" tableUuid="DF1B1DAED003FFF494AEFC3FFDD4FBC7">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BDB4D26D003FFF49778FED9FBDDFD5F" blockId="6.[824,1475,279,975]" pageId="6" pageNumber="7">
The Arctic forfs of 
<taxonomicName id="4C6436A5D003FFF4906CFED9FADAFEDF" box="[1098,1323,279,303]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF4906CFED9FADAFEDF" box="[1098,1323,279,303]" italics="true" pageId="6" pageNumber="7">Calamagrostis stricta</emphasis>
</taxonomicName>
show greater forphological plasticity than the Central European taxa, especially 
<taxonomicName id="4C6436A5D003FFF49789FE99FB06FE9F" box="[943,1271,343,367]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="subSpecies" species="stricta" subSpecies="groenlandica">
<emphasis id="B9109134D003FFF49789FE99FBE4FE9F" box="[943,1045,343,367]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
subsp. 
<emphasis id="B9109134D003FFF4904BFE99FB06FE9F" box="[1133,1271,343,367]" italics="true" pageId="6" pageNumber="7">groenlandica</emphasis>
</taxonomicName>
that shows sifilarities with the new species. According to literature (see Material and fethods), 
<taxonomicName id="4C6436A5D003FFF49069FE59FA63FE5F" box="[1103,1426,407,431]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="subSpecies" species="stricta" subSpecies="groenlandica">
<emphasis id="B9109134D003FFF49069FE59FB45FE5F" box="[1103,1204,407,431]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
subsp. 
<emphasis id="B9109134D003FFF4912CFE59FA63FE5F" box="[1290,1426,407,431]" italics="true" pageId="6" pageNumber="7">groenlandica</emphasis>
</taxonomicName>
is a variable species concerning the habit and the panicle length, displaying panicles as short as (3–)4 cf long. However, the panicles often refain contracted (spreading at anthesis in 
<taxonomicName id="4C6436A5D003FFF4971EFDD9FC6DFDDF" box="[824,924,535,559]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF4971EFDD9FC6DFDDF" box="[824,924,535,559]" italics="true" pageId="6" pageNumber="7">C. lonana</emphasis>
</taxonomicName>
), the callus hairs are usually fore than ½ of leffa length (less than ½ of leffa length in 
<taxonomicName id="4C6436A5D003FFF490C5FDF9FAB9FDBF" box="[1251,1352,567,591]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF490C5FDF9FAB9FDBF" box="[1251,1352,567,591]" italics="true" pageId="6" pageNumber="7">C. lonana</emphasis>
</taxonomicName>
), the ligule is (0.5–)1.5–3(–6) ff long (0.5–1 ff long in 
<taxonomicName id="4C6436A5D003FFF49103FD99FA76FD9F" box="[1317,1415,599,623]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF49103FD99FA76FD9F" box="[1317,1415,599,623]" italics="true" pageId="6" pageNumber="7">C. lonana</emphasis>
</taxonomicName>
), and the glufes are fore or less scabrous (glabrous or fifbriate on the keels in 
<taxonomicName id="4C6436A5D003FFF4979EFD59FBEDFD5F" box="[952,1052,663,687]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF4979EFD59FBEDFD5F" box="[952,1052,663,687]" italics="true" pageId="6" pageNumber="7">C. lonana</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BDB4D26D003FFF49778FD79FA33FC3F" blockId="6.[824,1475,279,975]" pageId="6" pageNumber="7">
Based on its forphology and cytology, the taxon found in the Lona alluvial plain belongs to the 
<taxonomicName id="4C6436A5D003FFF490CBFD19FA33FD1F" box="[1261,1474,727,751]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF490CBFD19FA33FD1F" box="[1261,1474,727,751]" italics="true" pageId="6" pageNumber="7">Calamagrostis stricta</emphasis>
</taxonomicName>
aggregate (syn. 
<taxonomicName id="4C6436A5D003FFF497FDFD39FBB9FCFF" box="[987,1096,759,783]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="neglecta">
<emphasis id="B9109134D003FFF497FDFD39FBB9FCFF" box="[987,1096,759,783]" italics="true" pageId="6" pageNumber="7">C. neglecta</emphasis>
</taxonomicName>
). The aggregate is part of the boreal and arctic flora with relict occurrences in Central Europe. Morphologically, the aggregate is very variable and therefore the taxonofy of the whole cofplex is controversially discussed. 
<taxonomicName id="4C6436A5D003FFF497A0FCB9FBACFC7F" box="[902,1117,887,911]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF497A0FCB9FBACFC7F" box="[902,1117,887,911]" italics="true" pageId="6" pageNumber="7">Calamagrostis lonana</emphasis>
</taxonomicName>
shows the safe tetraploid level as all other non-apofictic taxa (2 
<emphasis id="B9109134D003FFF4905EFC56FB77FC5F" box="[1144,1158,920,943]" italics="true" pageId="6" pageNumber="7">n</emphasis>
= 28, x = 7; ELVEN et al., 2023) and dimmers frof higher-ploidy taxa collected in and around the
</paragraph>
<caption id="DF1B1DAED003FFF494AEFC3FFDD4FBC7" ID-Table-UUID="DF1B1DAED003FFF494AEFC3FFDD4FBC7" httpUri="http://table.plazi.org/id/DF1B1DAED003FFF494AEFC3FFDD4FBC7" pageId="6" pageNumber="7" startId="6.[136,187,1009,1029]" targetBox="[144,1410,1105,1941]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BDB4D26D003FFF494AEFC3FFDD4FBC7" blockId="6.[136,1367,1007,1079]" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF494AEFC3FFF11FBF5" bold="true" box="[136,224,1009,1029]" pageId="6" pageNumber="7">Table 2. –</emphasis>
Comparison of morphological characters between 
<taxonomicName id="4C6436A5D003FFF4969EFC3FFBCCFBF5" authority="Eggenb. &amp; Leibundg." authorityName="Eggenb. &amp; Leibundg." box="[696,1085,1007,1029]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D003FFF4969EFC3FFC8EFBF3" box="[696,895,1007,1029]" italics="true" pageId="6" pageNumber="7">Calamagrostis lonana</emphasis>
Eggenb.&amp; Leibundg.
</taxonomicName>
(own measurements) and the Central European 
<taxonomicName id="4C6436A5D003FFF4955CFBC4FDA0FBEE" authority="(Timm) Koeler" authorityName="Koeler" baseAuthorityName="Timm" box="[378,593,1033,1054]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF4955CFBC4FE3CFBED" box="[378,461,1033,1054]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
(Timm) Koeler
</taxonomicName>
(measurements retrieved from HESS et al., 1967–1972; 
<bibRefCitation id="EFF530D7D003FFF4906DFBC4FB20FBEE" author="QUINGER" box="[1099,1233,1034,1054]" pageId="6" pageNumber="7" pagination="7 - 22" refId="ref6493" refString="QUINGER &amp; B. (1987). Zur Wiederentdeckung von Calafagrostis stricta (Tiff) Koeler in Bayern. Ber. Bayer. Bot. Ges. 58: 7 - 22." type="journal article" year="1987">QUINGER, 1987</bibRefCitation>
; 
<bibRefCitation id="EFF530D7D003FFF490F1FBC4FAA2FBEE" author="CONERT" box="[1239,1363,1034,1054]" pageId="6" pageNumber="7" pagination="3" refId="ref5839" refString="CONERT &amp; H. J. (1998). Calafagrostis. In: CONERT &amp; H. J. (ed.) &amp; Ill. Fl. Mitt. - Eur. Ed. 3. OEol. 1 (3). Parey Buchverlag &amp; Berlin." type="journal article" year="1998">CONERT, 1998</bibRefCitation>
; MÜLLER &amp; al., 2021; PAROLLY &amp; 
<bibRefCitation id="EFF530D7D003FFF495BEFBEDFDECFBC7" author="J. G. ROHWER" box="[408,541,1059,1079]" pageId="6" pageNumber="7" refId="ref6418" refString="PAROLLY &amp; G. &amp; J. G. ROHWER (ed.) (2016). Schmeil-Fitschen - Die Flora Deutschlands und angrenzender Lander 96. OEerlag Quelle &amp; Meyer &amp; Wiebelsheif." type="journal volume" year="2016">ROHWER, 2016</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BDB4D26D003FFF49668FB9FFB80F865" pageId="6" pageNumber="7">
<table id="F964BF86D003000D94B6FB9FFA73F865" box="[144,1410,1105,1941]" gridcols="4" gridrows="16" pageId="6" pageNumber="7">
<tr id="35544F64D003000D94B6FB9FFA73FB95" box="[144,1410,1105,1125]" gridrow="0" pageId="6" pageNumber="7" rowspan-0="1" rowspan-1="1">
<th id="76852618D003000D9668FB9FFBF3FB95" box="[590,1026,1105,1125]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
<taxonomicName id="4C6436A5D003FFF49668FB9FFD5BFB95" box="[590,682,1105,1125]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="lonana">C. lonana</taxonomicName>
</th>
<th id="76852618D003000D9035FB9FFA73FB95" box="[1043,1410,1105,1125]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">
<taxonomicName id="4C6436A5D003FFF49035FB9FFB9FFB95" box="[1043,1134,1105,1125]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D003FFF49035FB9FFB9FFB95" bold="true" box="[1043,1134,1105,1125]" italics="true" pageId="6" pageNumber="7">C. stricta</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35544F64D003000D94B6FBB0FA73FB62" box="[144,1410,1150,1170]" gridrow="1" pageId="6" pageNumber="7">
<th id="76852618D003000D94B6FBB0FF00FB62" box="[144,241,1150,1170]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">Habit</th>
<td id="76852618D003000D952FFBB0FDC6FB62" box="[265,567,1150,1170]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952FFBB0FE42FB62" bold="true" box="[265,435,1150,1170]" pageId="6" pageNumber="7">Culm length [cm]</emphasis>
</td>
<td id="76852618D003000D9668FBB0FBF3FB62" box="[590,1026,1150,1170]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">(10–)13–18(–22)</td>
<td id="76852618D003000D9035FBB0FA73FB62" box="[1043,1410,1150,1170]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">30–100(–120)</td>
</tr>
<tr id="35544F64D003000D94B6FB64FA73FB30" box="[144,1410,1194,1216]" gridrow="2" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FFB64FDC6FB30" box="[265,567,1194,1216]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFB64FE81FB4E" bold="true" box="[268,368,1194,1214]" pageId="6" pageNumber="7">Size [mm]</emphasis>
</td>
<td id="76852618D003000D9668FB64FBF3FB30" box="[590,1026,1194,1216]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">80(–120) × 2–3(–3.5)</td>
<td id="76852618D003000D9035FB64FA73FB30" box="[1043,1410,1194,1216]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">up to 600 × (1.5–)2–4(–5)</td>
</tr>
<tr id="35544F64D003000D94B6FB19FA73FAEA" box="[144,1410,1239,1306]" gridrow="3" pageId="6" pageNumber="7">
<th id="76852618D003000D94B6FB19FF00FAEA" box="[144,241,1239,1306]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF494B5FB3CFF28FAF6" bold="true" box="[147,217,1266,1286]" pageId="6" pageNumber="7">Leaves</emphasis>
</th>
<td id="76852618D003000D952FFB19FDC6FAEA" box="[265,567,1239,1306]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFB19FE88FB1C" bold="true" box="[268,377,1239,1260]" pageId="6" pageNumber="7">Colour and</emphasis>
<emphasis id="B9109134D003FFF49558FB16FE5FFB1C" bold="true" box="[382,430,1240,1260]" pageId="6" pageNumber="7">form</emphasis>
<emphasis id="B9109134D003FFF4952AFACBFE36FAE9" bold="true" box="[268,455,1285,1305]" pageId="6" pageNumber="7">Ligule length [mm]</emphasis>
</td>
<td id="76852618D003000D9668FB19FBF3FAEA" box="[590,1026,1239,1306]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">
<taxonomicName id="4C6436A5D003FFF49668FB17FD8AFB1D" box="[590,635,1241,1261]" form="light" pageId="6" pageNumber="7" rank="form">light</taxonomicName>
green to greyish green, mostly enrolled 0.5–1
</td>
<td id="76852618D003000D9035FB19FA73FAEA" box="[1043,1410,1239,1306]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">fresh to greyish green, flat or rolled 2–3</td>
</tr>
<tr id="35544F64D003000D94B6FAFCFA73FAB8" box="[144,1410,1330,1352]" gridrow="4" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FFAFCFDC6FAB8" box="[265,567,1330,1352]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFAFCFDC6FAB6" bold="true" box="[268,567,1330,1350]" pageId="6" pageNumber="7">Ligule indument (abaxial side)</emphasis>
</td>
<td id="76852618D003000D9668FAFCFBF3FAB8" box="[590,1026,1330,1352]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">glabrous</td>
<td id="76852618D003000D9035FAFCFA73FAB8" box="[1043,1410,1330,1352]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">hairy</td>
</tr>
<tr id="35544F64D003000D94B6FAAEFA73FA85" box="[144,1410,1376,1397]" gridrow="5" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FFAAEFDC6FA85" box="[265,567,1376,1397]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFAAEFE75FA84" bold="true" box="[268,388,1376,1396]" pageId="6" pageNumber="7">Length [cm]</emphasis>
</td>
<td id="76852618D003000D9668FAAEFBF3FA85" box="[590,1026,1376,1397]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">(2.4–)3.1–3.7(–4)</td>
<td id="76852618D003000D9035FAAEFA73FA85" box="[1043,1410,1376,1397]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">8–15(–20)</td>
</tr>
<tr id="35544F64D003000D94B6FA43FA73FA20" box="[144,1410,1421,1488]" gridrow="6" pageId="6" pageNumber="7">
<th id="76852618D003000D94B6FA43FF00FA20" box="[144,241,1421,1488]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF494B5FA66FF19FA4C" bold="true" box="[147,232,1448,1468]" pageId="6" pageNumber="7">Panicles</emphasis>
</th>
<td id="76852618D003000D952FFA43FDC6FA20" box="[265,567,1421,1488]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFA43FEBDFA51" bold="true" box="[268,332,1421,1441]" pageId="6" pageNumber="7">Shape</emphasis>
<emphasis id="B9109134D003FFF4952AFA74FEBEFA3E" bold="true" box="[268,335,1466,1486]" pageId="6" pageNumber="7">Colour</emphasis>
</td>
<td id="76852618D003000D9668FA43FBF3FA20" box="[590,1026,1421,1488]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">contracted but spread at anthesis grey to lilac</td>
<td id="76852618D003000D9035FA43FA73FA20" box="[1043,1410,1421,1488]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">contracted, rarely spread at anthesis grey to pale lilac</td>
</tr>
<tr id="35544F64D003000D94B6FA26FA73FA0D" box="[144,1410,1512,1533]" gridrow="7" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FFA26FDC6FA0D" box="[265,567,1512,1533]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AFA26FE2FFA0C" bold="true" box="[268,478,1512,1532]" pageId="6" pageNumber="7">Branche length [mm]</emphasis>
</td>
<td id="76852618D003000D9668FA26FBF3FA0D" box="[590,1026,1512,1533]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">1–6</td>
<td id="76852618D003000D9035FA26FA73FA0D" box="[1043,1410,1512,1533]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">15–60</td>
</tr>
<tr id="35544F64D003000D94B6F9DBFA73F9DA" box="[144,1410,1557,1578]" gridrow="8" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF9DBFDC6F9DA" box="[265,567,1557,1578]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF9DBFE7AF9D9" bold="true" box="[268,395,1557,1577]" pageId="6" pageNumber="7">Length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F9DBFBF3F9DA" box="[590,1026,1557,1578]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">2.5–3.2(–3.6)</td>
<td id="76852618D003000D9035F9DBFA73F9DA" box="[1043,1410,1557,1578]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">3–4(–4.5)</td>
</tr>
<tr id="35544F64D003000D94B6F98CFA73F9A8" box="[144,1410,1602,1624]" gridrow="9" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF98CFDC6F9A8" box="[265,567,1602,1624]" gridcol="1" gridrow="9" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF98CFE3AF9A7" bold="true" box="[268,459,1602,1623]" pageId="6" pageNumber="7">Glume length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F98CFBF3F9A8" box="[590,1026,1602,1624]" gridcol="2" gridrow="9" pageId="6" pageNumber="7">2.6–3.1</td>
<td id="76852618D003000D9035F98CFA73F9A8" box="[1043,1410,1602,1624]" gridcol="3" gridrow="9" pageId="6" pageNumber="7">3–4(–4.5)</td>
</tr>
<tr id="35544F64D003000D94B6F9BEFA73F975" box="[144,1410,1648,1669]" gridrow="10" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF9BEFDC6F975" box="[265,567,1648,1669]" gridcol="1" gridrow="10" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF9BEFE05F974" bold="true" box="[268,500,1648,1668]" pageId="6" pageNumber="7">Callus hair length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F9BEFBF3F975" box="[590,1026,1648,1669]" gridcol="2" gridrow="10" pageId="6" pageNumber="7">1–1.4, &lt;½ of lemma length</td>
<td id="76852618D003000D9035F9BEFA73F975" box="[1043,1410,1648,1669]" gridcol="3" gridrow="10" pageId="6" pageNumber="7">1.5–2.5, ½ to ¾ of lemma length</td>
</tr>
<tr id="35544F64D003000D94B6F953FA73F910" box="[144,1410,1693,1760]" gridrow="11" pageId="6" pageNumber="7">
<th id="76852618D003000D94B6F953FF00F910" box="[144,241,1693,1760]" gridcol="0" gridrow="11" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF494B5F976FF00F93C" bold="true" box="[147,241,1720,1740]" pageId="6" pageNumber="7">Spikelets</emphasis>
</th>
<td id="76852618D003000D952FF953FDC6F910" box="[265,567,1693,1760]" gridcol="1" gridrow="11" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF953FDDAF941" bold="true" box="[268,555,1693,1713]" pageId="6" pageNumber="7">Axillary process length [mm]</emphasis>
<emphasis id="B9109134D003FFF4952AF905FE25F92F" bold="true" box="[268,468,1739,1759]" pageId="6" pageNumber="7">Lemma length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F953FBF3F910" box="[590,1026,1693,1760]" gridcol="2" gridrow="11" pageId="6" pageNumber="7">0.3–0.5 2.1–2.4(–2.7)</td>
<td id="76852618D003000D9035F953FA73F910" box="[1043,1410,1693,1760]" gridcol="3" gridrow="11" pageId="6" pageNumber="7">0.8–1 2.6–3.5</td>
</tr>
<tr id="35544F64D003000D94B6F936FA73F8FD" box="[144,1410,1784,1805]" gridrow="12" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF936FDC6F8FD" box="[265,567,1784,1805]" gridcol="1" gridrow="12" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF936FE47F8FC" bold="true" box="[268,438,1784,1804]" pageId="6" pageNumber="7">Awn length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F936FBF3F8FD" box="[590,1026,1784,1805]" gridcol="2" gridrow="12" pageId="6" pageNumber="7">1.5–1.9</td>
<td id="76852618D003000D9035F936FA73F8FD" box="[1043,1410,1784,1805]" gridcol="3" gridrow="12" pageId="6" pageNumber="7">2–3</td>
</tr>
<tr id="35544F64D003000D94B6F8EBFA73F8CA" box="[144,1410,1829,1850]" gridrow="13" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF8EBFDC6F8CA" box="[265,567,1829,1850]" gridcol="1" gridrow="13" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF8EBFE33F8C9" bold="true" box="[268,450,1829,1849]" pageId="6" pageNumber="7">Palea length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F8EBFBF3F8CA" box="[590,1026,1829,1850]" gridcol="2" gridrow="13" pageId="6" pageNumber="7">1.3–1.6</td>
<td id="76852618D003000D9035F8EBFA73F8CA" box="[1043,1410,1829,1850]" gridcol="3" gridrow="13" pageId="6" pageNumber="7">1.8–2.3</td>
</tr>
<tr id="35544F64D003000D94B6F89DFA73F898" box="[144,1410,1875,1896]" gridrow="14" pageId="6" pageNumber="7" rowspan-0="1">
<td id="76852618D003000D952FF89DFDC6F898" box="[265,567,1875,1896]" gridcol="1" gridrow="14" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952AF89DFE3FF897" bold="true" box="[268,462,1875,1895]" pageId="6" pageNumber="7">Anther length [mm]</emphasis>
</td>
<td id="76852618D003000D9668F89DFBF3F898" box="[590,1026,1875,1896]" gridcol="2" gridrow="14" pageId="6" pageNumber="7">0.9–1.4</td>
<td id="76852618D003000D9035F89DFA73F898" box="[1043,1410,1875,1896]" gridcol="3" gridrow="14" pageId="6" pageNumber="7">2–2.5</td>
</tr>
<tr id="35544F64D003000D94B6F84FFA73F865" box="[144,1410,1921,1941]" gridrow="15" pageId="6" pageNumber="7">
<th id="76852618D003000D94B6F84FFF00F865" box="[144,241,1921,1941]" gridcol="0" gridrow="15" pageId="6" pageNumber="7">Cytology</th>
<td id="76852618D003000D952FF84FFDC6F865" box="[265,567,1921,1941]" gridcol="1" gridrow="15" pageId="6" pageNumber="7">
<emphasis id="B9109134D003FFF4952FF84FFE11F865" bold="true" box="[265,480,1921,1941]" pageId="6" pageNumber="7">Chromosome number</emphasis>
</td>
<td id="76852618D003000D9668F84FFBF3F865" box="[590,1026,1921,1941]" gridcol="2" gridrow="15" pageId="6" pageNumber="7">
2 
<emphasis id="B9109134D003FFF4967CF84FFD97F865" box="[602,614,1921,1941]" italics="true" pageId="6" pageNumber="7">n</emphasis>
= 28
</td>
<td id="76852618D003000D9035F84FFA73F865" box="[1043,1410,1921,1941]" gridcol="3" gridrow="15" pageId="6" pageNumber="7">
<geoCoordinate id="EE502BE1D003FFF49035F84FFBDAF865" box="[1043,1067,1921,1941]" degrees="28" direction="north" minutes="2" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="28.033333">
2 
<emphasis id="B9109134D003FFF49039F84FFBDAF865" box="[1055,1067,1921,1941]" italics="true" pageId="6" pageNumber="7">n</emphasis>
</geoCoordinate>
= c. 70
</td>
</tr>
</table>
</paragraph>
<caption id="DF1B1DAED002FFF59457FB66FABAFB4C" ID-DOI="http://doi.org/10.5281/zenodo.8110239" ID-Zenodo-Dep="8110239" box="[113,1355,1192,1212]" httpUri="https://zenodo.org/record/8110239/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,142,1192,1212]" targetBox="[113,1451,276,1168]" targetPageId="7" targetType="figure">
<paragraph id="8BDB4D26D002FFF59457FB66FABAFB4C" blockId="7.[113,1355,1192,1237]" box="[113,1355,1192,1212]" pageId="7" pageNumber="8">
<emphasis id="B9109134D002FFF59457FB66FF46FB4C" bold="true" box="[113,183,1192,1212]" pageId="7" pageNumber="8">Fig. 4. –</emphasis>
Alpine alluvial plain of Lona, with fens, alluvial and spring vegetation patches on both sides of the meandering Torrent de Lona.
</paragraph>
</caption>
<paragraph id="8BDB4D26D002FFF59457FB0FFEC6FB25" blockId="7.[113,1355,1192,1237]" box="[113,311,1217,1237]" pageId="7" pageNumber="8">[Photo: S. Eggenberg]</paragraph>
<paragraph id="8BDB4D26D002FFF59457FAC2FDC9F9B4" blockId="7.[113,764,1292,1956]" pageId="7" pageNumber="8">
Alps. Despite the tiny axillary process, SAARELA et al. (2017) place other tetraploid forfs of 
<taxonomicName id="4C6436A5D002FFF595EBFAE3FDDFFAB4" box="[461,558,1325,1348]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D002FFF595EBFAE3FDDFFAB4" box="[461,558,1325,1348]" italics="true" pageId="7" pageNumber="8">C. stricta</emphasis>
</taxonomicName>
aggregate close to 
<taxonomicName id="4C6436A5D002FFF59457FA83FE7DFA94" authority="(Weber) Roth" authorityName="Roth" baseAuthorityName="Weber" box="[113,396,1356,1380]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis id="B9109134D002FFF59457FA83FF1DFA94" box="[113,236,1357,1380]" italics="true" pageId="7" pageNumber="8">C. canescens</emphasis>
(Weber) Roth
</taxonomicName>
and 
<taxonomicName id="4C6436A5D002FFF595E5FA83FD19FA94" authority="(Chaix) J. F. Gfel." authorityName="J. F. Gfel." baseAuthorityName="Chaix" box="[451,744,1356,1380]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="B9109134D002FFF595E5FA83FDD7FA94" box="[451,550,1356,1380]" italics="true" pageId="7" pageNumber="8">C. villosa</emphasis>
(Chaix) J.F. Gfel.
</taxonomicName>
, i.e., to species that do not possess an axial process. However, the subspecies that often shows sfall growth, 
<taxonomicName id="4C6436A5D002FFF5967CFA43FF06FA34" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="subSpecies" species="stricta" subSpecies="groenlandica">
<emphasis id="B9109134D002FFF5967CFA43FD46FA54" box="[602,695,1421,1444]" italics="true" pageId="7" pageNumber="8">C. stricta</emphasis>
subsp. 
<emphasis id="B9109134D002FFF59457FA62FF06FA34" box="[113,247,1452,1476]" italics="true" pageId="7" pageNumber="8">groenlandica</emphasis>
</taxonomicName>
, lies outside this clade (SAARELA et al., 2017), which ifplies a clearer delifitation frof 
<taxonomicName id="4C6436A5D002FFF5961BFA03FD6AFA14" box="[573,667,1485,1508]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D002FFF5961BFA03FD6AFA14" box="[573,667,1485,1508]" italics="true" pageId="7" pageNumber="8">C. stricta</emphasis>
</taxonomicName>
. Further genetic studies should better resolve the 
<taxonomicName id="4C6436A5D002FFF5960BFA23FD7CF9F4" box="[557,653,1517,1540]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D002FFF5960BFA23FD7CF9F4" box="[557,653,1517,1540]" italics="true" pageId="7" pageNumber="8">C. stricta</emphasis>
</taxonomicName>
aggregate and provide fore clues to the taxonofic position of the new species found in the alluvial plain of Lona.
</paragraph>
<paragraph id="8BDB4D26D002FFF594B1F982FF4DF8D4" blockId="7.[113,764,1292,1956]" pageId="7" pageNumber="8">The flower organs develop very late. Anthers and ovaries with feathery stigfas could be observed frof the end of Septefber, but no fertile pollen was observed. Conversely, the species shows a strong growth of underground runners. In cultivation, the species shows especially strong vegetative growth and forfs dense, vigorous fats within one growing season.</paragraph>
<paragraph id="8BDB4D26D002FFF594B1F8E2FAB5FA54" blockId="7.[113,764,1292,1956]" lastBlockId="7.[801,1452,1292,1444]" pageId="7" pageNumber="8">
The 2 
<emphasis id="B9109134D002FFF594F4F8E3FF11F8B4" box="[210,224,1837,1860]" italics="true" pageId="7" pageNumber="8">n</emphasis>
-genofe size estifates obtained by flow cytofetry varied between 7.33 pg and 8.10 pg of DNA for safples of 
<taxonomicName id="4C6436A5D002FFF59457F8A2FEA1F874" box="[113,336,1900,1924]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="lonana">
<emphasis id="B9109134D002FFF59457F8A2FEA1F874" box="[113,336,1900,1924]" italics="true" pageId="7" pageNumber="8">Calamagrostis lonana</emphasis>
</taxonomicName>
, perfectly fatching estifates for tetraploid 
<taxonomicName id="4C6436A5D002FFF59496F842FEB0F854" box="[176,321,1932,1956]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="B9109134D002FFF59496F842FEB0F854" box="[176,321,1932,1956]" italics="true" pageId="7" pageNumber="8">Calamagrostis</emphasis>
</taxonomicName>
taxa (2 
<emphasis id="B9109134D002FFF595B7F843FE6EF854" box="[401,415,1933,1956]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= 28) published in the literature (SMARDA et al., 2019; ZONNEFELD, 2019). In contrast, all other safples of presufably 
<taxonomicName id="4C6436A5D002FFF5903FFAE3FB87FAB4" box="[1049,1142,1325,1348]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B9109134D002FFF5903FFAE3FB87FAB4" box="[1049,1142,1325,1348]" italics="true" pageId="7" pageNumber="8">C. stricta</emphasis>
</taxonomicName>
collected across nearby locations in 
<collectingCountry id="F3730DB6D002FFF59750FA83FC4CFA94" box="[886,957,1357,1380]" name="France" pageId="7" pageNumber="8">France</collectingCountry>
and Gerfany presented genofe size estifates ranging frof 14.39 pg to 16.50 pg, rather fatching decaploid taxa of 
<taxonomicName id="4C6436A5D002FFF59749FA42FC0EFA54" box="[879,1023,1420,1444]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="B9109134D002FFF59749FA42FC0EFA54" box="[879,1023,1420,1444]" italics="true" pageId="7" pageNumber="8">Calamagrostis</emphasis>
</taxonomicName>
with 2 
<emphasis id="B9109134D002FFF5906CFA43FBA9FA54" box="[1098,1112,1421,1444]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= c. 70 chrofosofes.
</paragraph>
</subSubSection>
</treatment>
</document>