418 lines
50 KiB
XML
418 lines
50 KiB
XML
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<mods:title id="92AA999D1ACE15C98D5458B60E67486A">Taxonomic review of the Oriental flower beetle Coilodera penicillata species complex (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<emphasis id="B910060EFFD7FC492E81FFAAF331FC43" bold="true" box="[151,270,902,928]" italics="true" pageId="3" pageNumber="514">Coilodera</emphasis>
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<emphasis id="B910060EFFD7FC492E81FFEBF2C1FC3E" box="[151,254,967,989]" italics="true" pageId="3" pageNumber="514">Coilodera</emphasis>
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<bibRefCitation id="EFF5A7EDFFD7FC492F12FFE4F340FC3E" author="Hope" box="[260,383,967,989]" pageId="3" pageNumber="514" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope, 1831</bibRefCitation>
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: 25
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;
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<treatmentCitation id="0AC5FC0DFFD7FC492FA6FFEBF06DFC3E" author="Arrow" box="[432,594,967,989]" page="41" pageId="3" pageNumber="514" year="1910">
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<bibRefCitation id="EFF5A7EDFFD7FC492FA6FFEBF010FC3E" author="Arrow" box="[432,559,967,989]" pageId="3" pageNumber="514" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor & Francis, London, United Kingdom, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
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: 41
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(synonym of
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<taxonomicName id="4C64A19FFFD7FC492CF7FFE4F16FFC3E" authorityName="Hoffmannsegg" authorityYear="1817" box="[737,848,968,989]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
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<emphasis id="B910060EFFD7FC492CF7FFE4F16FFC3E" box="[737,848,968,989]" italics="true" pageId="3" pageNumber="514">Macronota</emphasis>
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);
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<treatmentCitation id="0AC5FC0DFFD7FC492D71FFEBF629FC3E" author="Arrow" box="[871,1046,967,989]" page="141" pageId="3" pageNumber="514" year="1946">
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<bibRefCitation id="EFF5A7EDFFD7FC492D71FFEBF1D9FC3E" author="Arrow" box="[871,998,967,989]" pageId="3" pageNumber="514" refString="Arrow, G. J. (1946) Synonymic notes on some Oriental Cetoniine beetles, with descriptions of two new species. Proceedings of the Royal Entomological Society of London (B), 15, 140 - 144. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 01399. x" type="journal article" year="1946">Arrow 1946</bibRefCitation>
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: 141
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(as replacement name of
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<taxonomicName id="4C64A19FFFD7FC492B0BFFE4F7B3FC3E" authorityName="Hoffmannsegg" authorityYear="1817" box="[1309,1420,968,989]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
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<emphasis id="B910060EFFD7FC492B0BFFE4F7B3FC3E" box="[1309,1420,968,989]" italics="true" pageId="3" pageNumber="514">Macronota</emphasis>
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);
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<treatmentCitation id="0AC5FC0DFFD7FC492ED1FFC9F3BEFC18" author="Krikken" box="[199,385,997,1019]" page="63" pageId="3" pageNumber="514" year="1984">
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<bibRefCitation id="EFF5A7EDFFD7FC492ED1FFC9F366FC18" author="Krikken" box="[199,345,997,1019]" pageId="3" pageNumber="514" refString="Krikken, J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera." type="book" year="1984">Krikken 1984</bibRefCitation>
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: 63
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||
</treatmentCitation>
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;
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<treatmentCitation id="0AC5FC0DFFD7FC492F98FFC9F0A2FC18" author="Sakai" box="[398,669,997,1019]" page="356" pageId="3" pageNumber="514" year="1998">
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<bibRefCitation id="EFF5A7EDFFD7FC492F98FFC9F058FC18" author="Sakai" box="[398,615,997,1019]" pageId="3" pageNumber="514" refString="Sakai, K. & Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Volume 3. Mushi- Sha, Tokyo, Japan, 421 pp." type="book" year="1998">Sakai & Nagai 1998</bibRefCitation>
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: 356
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||
</treatmentCitation>
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;
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<treatmentCitation id="0AC5FC0DFFD7FC492CBCFFC9F164FC18" author="Krajcik" box="[682,859,997,1019]" page="88" pageId="3" pageNumber="514" year="1998">
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<bibRefCitation id="EFF5A7EDFFD7FC492CBCFFC9F10AFC18" author="Krajcik" box="[682,821,997,1019]" pageId="3" pageNumber="514" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, Czech Republic, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
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: 88
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||
</treatmentCitation>
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(catalogue);
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||
<treatmentCitation id="0AC5FC0DFFD7FC492DF3FFC9F6E9FC18" author="Ek-Amnuay" box="[997,1238,997,1019]" page="246" pageId="3" pageNumber="514" year="2008">
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<bibRefCitation id="EFF5A7EDFFD7FC492DF3FFC9F69FFC18" author="Ek-Amnuay" box="[997,1184,997,1019]" pageId="3" pageNumber="514" refString="Ek-Amnuay, P. (2008) Beetles of Thailand (2 nd edition). Fascinating Insects. Folume 1. Amarin Printing and Publishing Public, Bangkok, Thailand, 495 pp." type="book" year="2008">Ek-Amnuay 2008</bibRefCitation>
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: 246
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||
</treatmentCitation>
|
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;
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||
<treatmentCitation id="0AC5FC0DFFD7FC492AF4FFC9F31FFBF9" author="Smetana" page="39" pageId="3" pageNumber="514" year="2006">
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<bibRefCitation id="EFF5A7EDFFD7FC492AF4FFC9F2C1FBF9" author="Smetana" pageId="3" pageNumber="514" refString="Smetana, A. & Smith, A. B. T. (2006) New acts and comments. Scarabaeidae: Cetoniinae. In: Lobl, I. & Smetana, A. (eds.), Catalogue of Palaearctic Coleoptera Folume 3. Apollo Books, Stenstrup, Denmark, pp. 38 - 39." type="journal article" year="2006">Smetana & Smith 2006</bibRefCitation>
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: 39
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||
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(spelling); Smetana 2006: 306;
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<treatmentCitation id="0AC5FC0DFFD7FC492C73F828F12CFBF9" author="Krajcik" box="[613,787,1028,1050]" page="53" pageId="3" pageNumber="514" year="2011">
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<bibRefCitation id="EFF5A7EDFFD7FC492C73F828F0D1FBF9" author="Krajcik" box="[613,750,1028,1050]" pageId="3" pageNumber="514" refString="Krajcik, M. (2011) Illustrated catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x. Supplement, 1 - 113." type="book chapter" year="2011">Krajčík 2011</bibRefCitation>
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: 53
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</treatmentCitation>
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;
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<treatmentCitation id="0AC5FC0DFFD7FC492D0BF828F1F6FBF9" author="Krajcik" box="[797,969,1028,1050]" page="76" pageId="3" pageNumber="514" year="2012">
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<bibRefCitation id="EFF5A7EDFFD7FC492D0BF828F198FBF9" author="Krajcik" box="[797,935,1028,1050]" pageId="3" pageNumber="514" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">Krajčík 2012</bibRefCitation>
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: 76
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</treatmentCitation>
|
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(catalogue);
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<treatmentCitation id="0AC5FC0DFFD7FC492A59F828F735FBF9" author="Bezdek" box="[1103,1290,1028,1050]" page="401" pageId="3" pageNumber="514" year="2016">
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<bibRefCitation id="EFF5A7EDFFD7FC492A59F828F6E8FBF9" author="Bezdek" box="[1103,1239,1028,1050]" pageId="3" pageNumber="514" refString="Bezdek, A. (2016) Subfamily Cetoniinae, pp. 367 - 412. In: Lobl, I. & Lobl, D. (eds), Catalogue of Palaearctic Coleoptera, Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, The Netherlands." type="journal article" year="2016">Bezděk 2016</bibRefCitation>
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: 401
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||
</treatmentCitation>
|
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.
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||
<taxonomicName id="4C64A19FFFD7FC492E81F80FF37BFBDA" authority="Hope" authorityName="Hope" box="[151,324,1059,1081]" class="Insecta" family="Scarabaeidae" genus="Coelodera" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
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<emphasis id="B910060EFFD7FC492E81F80FF33DFBDA" box="[151,258,1059,1081]" italics="true" pageId="3" pageNumber="514">Coelodera</emphasis>
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Hope
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</taxonomicName>
|
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:
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||
<treatmentCitation id="0AC5FC0DFFD7FC492F59F80FF010FBDA" author="Burmeister" box="[335,559,1059,1081]" page="320" pageId="3" pageNumber="514" year="1842">
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<bibRefCitation id="EFF5A7EDFFD7FC492F59F80FF3C2FBDA" author="Burmeister" box="[335,509,1059,1081]" pageId="3" pageNumber="514" refString="Burmeister, H. C. C. (1842) Handbuch der Entomologie, Band III. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Melitophila. T. C. F. Enslin, Berlin, Germany, 828 pp." type="book" year="1842">Burmeister 1842</bibRefCitation>
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: 320
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|
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(incorrect spelling);
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<treatmentCitation id="0AC5FC0DFFD7FC492D1EF80FF1F5FBDA" author="Harold" box="[776,970,1059,1081]" page="1297" pageId="3" pageNumber="514" year="1869">
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<bibRefCitation id="EFF5A7EDFFD7FC492D1EF80FF1B3FBDA" author="Harold" box="[776,908,1059,1081]" pageId="3" pageNumber="514" refString="Harold, E. von (1869) Scarabaeidae, Tome IV. In: Gemminger, M. & Harold, E. von (eds.), Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. E. H. Grummi, Munich, Germany, pp. 979 - 1346." type="book chapter" year="1869">Harold 1869</bibRefCitation>
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: 1297
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</treatmentCitation>
|
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(synonym of
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<taxonomicName id="4C64A19FFFD7FC492A4AF808F6F4FBDA" authorityName="Hoffmannsegg" authorityYear="1817" box="[1116,1227,1060,1081]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
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<emphasis id="B910060EFFD7FC492A4AF808F6F4FBDA" box="[1116,1227,1060,1081]" italics="true" pageId="3" pageNumber="514">Macronota</emphasis>
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);
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<treatmentCitation id="0AC5FC0DFFD7FC492AF4F80FF7A3FBDA" author="Schoch" box="[1250,1436,1059,1081]" page="203" pageId="3" pageNumber="514" year="1894">
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<bibRefCitation id="EFF5A7EDFFD7FC492AF4F80FF755FBDA" author="Schoch" box="[1250,1386,1059,1081]" pageId="3" pageNumber="514" refString="Schoch, G. (1894) Ueber die Systematik der Cetoniden. Mittheilungen der Schweizerischen Entomologischen Gesellschaft, 9, 164 - 225." type="journal article" year="1894">Schoch 1894</bibRefCitation>
|
||
: 203
|
||
</treatmentCitation>
|
||
(in key);
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492F33F86DF3E1FBB4" author="Schoch" box="[293,478,1089,1111]" page="38" pageId="3" pageNumber="514" year="1895">
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||
<bibRefCitation id="EFF5A7EDFFD7FC492F33F86DF387FBB4" author="Schoch" box="[293,440,1089,1111]" pageId="3" pageNumber="514" refString="Schoch, G. (1895 a) Die Genera und Species meiner Cetoniden-Sammlung I. Teil: Tribu. Goliathidae, Gymnetidae, Madagassae, Schizorrhinidae. E. Zwingli. Zurich, Switzerland, 64 pp." type="book" year="1895" yearSuffix="a">Schoch 1895a</bibRefCitation>
|
||
: 38
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492FFCF86DF09DFBB4" author="Schoch" box="[490,674,1089,1111]" page="34" pageId="3" pageNumber="514" year="1896">
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492FFCF86DF040FBB4" author="Schoch" box="[490,639,1089,1111]" pageId="3" pageNumber="514" refString="Schoch, G. (1896 b) Catalogus systematicus Cetonidarum et Trichiidarum ad huc cognitarum. W. Junk, Zurich, Switzerland, 95 pp." type="book" year="1896" yearSuffix="b">Schoch 1896b</bibRefCitation>
|
||
: 34
|
||
</treatmentCitation>
|
||
(catalogue);
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492D3FF86DF637FBB4" author="Schenkling" box="[809,1032,1089,1111]" page="133" pageId="3" pageNumber="514" year="1921">
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492D3FF86DF1E7FBB4" author="Schenkling" box="[809,984,1089,1111]" pageId="3" pageNumber="514" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, Germany, 431 pp." type="book" year="1921">Schenkling 1921</bibRefCitation>
|
||
: 133
|
||
</treatmentCitation>
|
||
(catalogue);
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492A99F86DF704FBB4" author="Miwa" box="[1167,1339,1089,1111]" page="305" pageId="3" pageNumber="514" year="1931">
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492A99F86DF737FBB4" author="Miwa" box="[1167,1288,1089,1111]" pageId="3" pageNumber="514" refString="Miwa, Y. (1931) A systematic catalogue of Formosan Coleoptera. Report of the Department of Agriculture of the Government Research Institute, 55, 1 - 359." type="journal article" year="1931">Miwa 1931</bibRefCitation>
|
||
: 305
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492B50F86DF358FB95" author="Miwa" page="83" pageId="3" pageNumber="514" year="1939">
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492B50F86DF37DFB95" author="Miwa" pageId="3" pageNumber="514" refString="Miwa, Y. & Chujo, M. (1939) Catalogus Coleopterum Japonicorm, Pars 5, Scarabaeidae. Noda-Syobo, Taihoku, Taiwan, 94 pp." type="book" year="1939">Miwa & Chûjô 1939</bibRefCitation>
|
||
: 83
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AC5FC0DFFD7FC492F64F84DF00FFB95" author="Yu" box="[370,560,1120,1142]" page="180" pageId="3" pageNumber="514" year="1998">
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492F64F84DF3C2FB95" author="Yu" box="[370,509,1120,1142]" pageId="3" pageNumber="514" refString="Yu, C. K., Kobayashi, H. & Chu, Y. I. (1998) The Scarabaeidae of Taiwan. Mu Sheng Co., Taipei, Taiwan, 263 pp." type="book" year="1998">
|
||
Yu
|
||
<emphasis id="B910060EFFD7FC492F82F84DF3FEFB95" box="[404,449,1120,1142]" italics="true" pageId="3" pageNumber="514">et al</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
: 180
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E8997FFD7FC492E81F8ACF2E1F811" pageId="3" pageNumber="514" type="materials_examined">
|
||
<paragraph id="8BDBDA1CFFD7FC492E81F8ACF131FB76" blockId="3.[151,1436,967,1173]" box="[151,782,1151,1173]" pageId="3" pageNumber="514">
|
||
Type species:
|
||
<taxonomicName id="4C64A19FFFD7FC492F30F853F043FB76" authority="Hope, 1831" authorityName="Hope" authorityYear="1831" box="[294,636,1151,1173]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="penicillata">
|
||
<emphasis id="B910060EFFD7FC492F30F853F3C0FB76" box="[294,511,1151,1173]" italics="true" pageId="3" pageNumber="514">Coilodera penicillata</emphasis>
|
||
<bibRefCitation id="EFF5A7EDFFD7FC492C13F8ACF043FB76" author="Hope" box="[517,636,1151,1173]" pageId="3" pageNumber="514" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope, 1831</bibRefCitation>
|
||
</taxonomicName>
|
||
by monotypy.
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD7FC492E81F8EDF2E1F811" blockId="3.[151,1437,1217,2034]" pageId="3" pageNumber="514">
|
||
<emphasis id="B910060EFFD7FC492E81F8EDF14CFB39" bold="true" box="[151,883,1217,1242]" pageId="3" pageNumber="514">
|
||
Characterization of the
|
||
<taxonomicName id="4C64A19FFFD7FC492FAFF8EDF093FB39" authorityName="Hope" authorityYear="1831" box="[441,684,1217,1242]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="penicillata">
|
||
<emphasis id="B910060EFFD7FC492FAFF8EDF093FB39" bold="true" box="[441,684,1217,1242]" italics="true" pageId="3" pageNumber="514">Coilodera penicillata</emphasis>
|
||
</taxonomicName>
|
||
species complex
|
||
</emphasis>
|
||
(Male). Length:
|
||
<quantity id="4C9C77F9FFD7FC492A20F8EDF6E1FB3A" box="[1078,1246,1217,1242]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.61" metricValueMin="1.85" pageId="3" pageNumber="514" unit="mm" value="22.3" valueMax="26.1" valueMin="18.5">18.5–26.1 mm</quantity>
|
||
; width:
|
||
<quantity id="4C9C77F9FFD7FC492B2DF8EEF2F9FB1E" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.17" metricValueMin="0.88" pageId="3" pageNumber="514" unit="mm" value="10.25" valueMax="11.7" valueMin="8.8">8.8–11.7 mm</quantity>
|
||
. Body black, more-or-less fusiform, shiny; with several orange-red, orange, yellow, or lemon-yellow tomentose areas; small, round setiferous punctures evenly covered on tomentose areas. Dorsal midline of body distinctly impressed. Head, pronotum, and protibia with wine-red, red, or metallic green reflections.
|
||
<emphasis id="B910060EFFD7FC492B14F901F77DFAA5" bold="true" box="[1282,1346,1325,1350]" pageId="3" pageNumber="514">Head</emphasis>
|
||
: Dorsal surface densely microsculptured; anterior margin of clypeus bisinuate, not raised; midline of frons slightly convex; frontolateral area clad with tomentum and sparse, long setae. Eye canthus extending laterally, narrow, long. Antenna brown, simple. Gula rounded, with a few short setae on sides.
|
||
<emphasis id="B910060EFFD7FC492DAEF9B5F605FA51" bold="true" box="[952,1082,1433,1458]" pageId="3" pageNumber="514">Pronotum:</emphasis>
|
||
Widest at base; pronotal ridges strongly raised; lateral margin curved and posterior margin distinctly protruding; except margin and pronotal ridges, other areas clad with tomentum and setiferous punctures. Posterior margin of the median tomentose area of pronotum notched or curved.
|
||
<emphasis id="B910060EFFD7FC492FE3FA29F049F9FD" bold="true" box="[501,630,1541,1566]" pageId="3" pageNumber="514">Scutellum:</emphasis>
|
||
Longitudinal triangle in shape, with or without tomentum.
|
||
<emphasis id="B910060EFFD7FC492B21FA29F7A3F9FD" bold="true" box="[1335,1436,1541,1566]" pageId="3" pageNumber="514">Elytron:</emphasis>
|
||
Prediscal area, humeral umbone and costa almost glabrous; lateral and distal declivities with short, sinuous striolae and sparse, long setae; mediodiscal and postdiscal areas of each elytron with 4 grooves, clad with sparse, long setae.
|
||
<emphasis id="B910060EFFD7FC492ECBFA5DF10FF969" bold="true" box="[221,816,1649,1674]" pageId="3" pageNumber="514">Mesepimeron, mesepisternum, and metepimeron:</emphasis>
|
||
Mesepimeron and mesepisternum clad with tomentum and setiferous punctures; metepimeron glabrous.
|
||
<emphasis id="B910060EFFD7FC492CCFFAB9F176F94D" bold="true" box="[729,841,1685,1710]" pageId="3" pageNumber="514">Sternum:</emphasis>
|
||
Preprosternum and mesosternum clad with sparse setae. Preprosternum without distinct preprosternal apophysis. Mesometasternal process glabrous, very short, apex nearly straight or slightly pointed; mesometasternal suture depressed or not depressed, with or without long setae. Middle portion of metasternum glabrous; anterior portion clad with tomentum and setiferous punctures; posterior portion with round or sinuous setiferous punctures.
|
||
<emphasis id="B910060EFFD7FC492CC9FB09F164F8DD" bold="true" box="[735,859,1829,1854]" pageId="3" pageNumber="514">Pygidium:</emphasis>
|
||
A large, round, tomentose macula in middle; setae dense, very long.
|
||
<emphasis id="B910060EFFD7FC492F4DFB65F3E6F881" bold="true" box="[347,473,1865,1890]" pageId="3" pageNumber="514">Abdomen:</emphasis>
|
||
Six sternites visible; abdominal sides exposed in dorsal view. Middle portion glabrous or with sparse setae. Sternite 2–6 (or
|
||
<quantity id="4C9C77F9FFD7FC492C47FB41F040F865" box="[593,639,1901,1926]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="514" unit="in" value="5.0">5 in</quantity>
|
||
some individuals) with a large macula and long setae on each side. Sternites 5–6 with sparse, long setae.
|
||
<emphasis id="B910060EFFD7FC492C57FBBDF0BDF849" bold="true" box="[577,642,1937,1962]" pageId="3" pageNumber="514">Legs:</emphasis>
|
||
Slender, with sparse, long setae; protibia with 2 or 3 large teeth along outer margin; metatibia with 3 teeth on apex.
|
||
<emphasis id="B910060EFFD7FC492CBBFB99F171F82D" bold="true" box="[685,846,1973,1998]" pageId="3" pageNumber="514">Male genital:</emphasis>
|
||
Parameres annular in frontal view; apex barbed in shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E8997FFD0FC4E2ED1FCBBF01CFF1B" pageId="4" pageNumber="515" type="description">
|
||
<paragraph id="8BDBDA1CFFD0FC4E2ED1FCBBF01CFF1B" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="515">
|
||
<emphasis id="B910060EFFD0FC4E2ED1FCBBF387FF53" bold="true" box="[199,440,151,176]" pageId="4" pageNumber="515">Sexual dimorphism.</emphasis>
|
||
Female similar to male, but lateral margin of pronotum slightly angular, protibia a little wider, abdomen convex with sixth visible abdominal segment more exposed. Outer tooth of metatibia always longer than ventral tooth in female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E8997FFD0FC4E2ED1FD28F32CFDFB" pageId="4" pageNumber="515" type="distribution">
|
||
<paragraph id="8BDBDA1CFFD0FC4E2ED1FD28F170FEFE" blockId="4.[151,1437,151,1221]" box="[199,847,260,285]" pageId="4" pageNumber="515">
|
||
<emphasis id="B910060EFFD0FC4E2ED1FD28F35FFEFE" bold="true" box="[199,352,260,285]" pageId="4" pageNumber="515">Distribution.</emphasis>
|
||
Oriental region (excluding the
|
||
<collectingCountry id="F3739A8CFFD0FC4E2CD6FD28F17CFEFE" box="[704,835,260,285]" name="Philippines" pageId="4" pageNumber="515">Philippines</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD0FC4E2ED1FD0BF32CFDFB" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="515">
|
||
<emphasis id="B910060EFFD0FC4E2ED1FD0BF3B9FEA3" bold="true" box="[199,390,295,320]" pageId="4" pageNumber="515">Commensalism.</emphasis>
|
||
The larvae and fresh adults of five
|
||
<taxonomicName id="4C64A19FFFD0FC4E2D18FD0BF1BFFEA3" authorityName="Hope" authorityYear="1831" box="[782,896,295,320]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2D18FD0BF1BFFEA3" box="[782,896,295,320]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
species were discovered in the galleries of some wood-feeding cockroaches (Blattaria:
|
||
<taxonomicName id="4C64A19FFFD0FC4E2C4DFD60F0E6FE86" box="[603,729,332,357]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="515" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C64A19FFFD0FC4E2CFCFD60F145FE86" box="[746,890,332,357]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="515" phylum="Arthropoda" rank="subFamily" subFamily="Panesthiinae">Panesthiinae</taxonomicName>
|
||
) in rotten wood, and they were regarded as facultative commensal species (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2C1BFD5CF0EDFE6B" author="Kon" box="[525,722,367,392]" pageId="4" pageNumber="515" refString="Kon, M., Johki, Y. & Sakai, K. (2002) The cetoniine beetle, Coilodera diardi (Coleoptera, Scarabaeidae, Cetoniinae), collected from a gallery of the wood-feeding cockroach, Salganea aequaliterspinosa (Blattaria, Blaberidae, Panesthiinae). Kogane, 3, 27 - 30." type="journal article" year="2002">
|
||
Kon
|
||
<emphasis id="B910060EFFD0FC4E2C5AFD5DF0B5FE6B" box="[588,650,367,392]" italics="true" pageId="4" pageNumber="515">et al.</emphasis>
|
||
2002
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2CF5FD5CF121FE6B" author="Kon" box="[739,798,368,392]" pageId="4" pageNumber="515" refString="Kon, M., Maekawa, K., Sakai, K. & Araya, K. (2003) The cetoniine beetle, Coilodera penicillata (Coleoptera, Scarabaeidae, Cetoniinae), collected from a gallery of the wood-feeding subsocial cockroach, Salganea raggei (Blattaria, Blaberidae, Panesthiinae). Kogane, 4, 41 - 44." type="journal article" year="2003">2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D39FD5CF155FE6B" author="Kon" box="[815,874,368,392]" pageId="4" pageNumber="515" refString="Kon, M., Maekawa, K., Sakai, K., Kusakabe, Y. & Araya, K. (2004) The cetoniine beetles, Coilodera penicillata and C. miksici (Coleoptera, Scarabaeidae, Cetoniinae), collected from the galleries of wood-feeding cockroaches (Blattaria, Blaberidae, Panesthiinae, Salganea) in Myanmar. Kogane, 5, 16 - 20." type="journal article" year="2004">2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D6AFD5CF63CFE6B" author="Ueno" box="[892,1027,367,392]" pageId="4" pageNumber="515" refString="Ueno, T. (2015) Larval habitats of two Coilodera species (Coleoptera: Scarabaeidae: Cetoniinae) associated with wood feeding cockroaches. Journal of Tropical Asian Entomology, 4, 28 - 32." type="journal article" year="2015">Ueno 2015</bibRefCitation>
|
||
). This view is coincided with our observation in field and artificial breeding, and we consider
|
||
<taxonomicName id="4C64A19FFFD0FC4E2D4FFDB8F1F4FE4E" authorityName="Hope" authorityYear="1831" box="[857,971,404,429]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2D4FFDB8F1F4FE4E" box="[857,971,404,429]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
spp. occasionally as inquilines of some xylophagous insects. It is presumed that
|
||
<taxonomicName id="4C64A19FFFD0FC4E2C70FD9BF0E7FE33" authorityName="Hope" authorityYear="1831" box="[614,728,439,464]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2C70FD9BF0E7FE33" box="[614,728,439,464]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
larvae can feed only on soft, highly rotten wood that decayed naturally or chewed by other wood-feeding insects in field, such as cockroaches and the larvae of
|
||
<taxonomicName id="4C64A19FFFD0FC4E2AE1FDF0F753FE16" authorityName="Leach" authorityYear="1815" box="[1271,1388,476,501]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C64A19FFFD0FC4E2E81FDD3F330FDFB" box="[151,271,511,536]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E8997FFD0FC4E2ED1FE08F2D6FB26" pageId="4" pageNumber="515" type="description">
|
||
<paragraph id="8BDBDA1CFFD0FC4E2ED1FE08F35FFD13" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="515">
|
||
<emphasis id="B910060EFFD0FC4E2ED1FE08F348FDDE" bold="true" box="[199,375,548,573]" pageId="4" pageNumber="515">Nomenclature.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C64A19FFFD0FC4E2FEAFE08F051FDDE" authorityName="Hope" authorityYear="1831" box="[508,622,548,573]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2FEAFE08F051FDDE" box="[508,622,548,573]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
was not characterized by
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D81FE09F619FDDF" author="Hope" box="[919,1062,548,572]" pageId="4" pageNumber="515" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope (1831)</bibRefCitation>
|
||
and only brief description of the
|
||
<typeStatus id="54DF64BEFFD0FC4E2E81FE64F2F8FD83" box="[151,199,584,608]" pageId="4" pageNumber="515">type</typeStatus>
|
||
species was provided. The first generic description was given by
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2DCEFE6BF693FD83" author="Burmeister" box="[984,1196,583,608]" pageId="4" pageNumber="515" refString="Burmeister, H. C. C. (1842) Handbuch der Entomologie, Band III. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Melitophila. T. C. F. Enslin, Berlin, Germany, 828 pp." type="book" year="1842">Burmeister (1842)</bibRefCitation>
|
||
, but he changed the generic name spelling as
|
||
<taxonomicName id="4C64A19FFFD0FC4E2FD4FE40F007FD66" box="[450,568,620,645]" class="Insecta" family="Scarabaeidae" genus="Coelodera" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2FD4FE40F007FD66" box="[450,568,620,645]" italics="true" pageId="4" pageNumber="515">Coelodera</emphasis>
|
||
</taxonomicName>
|
||
, an incorrect subsequent spelling (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2DC7FE40F6D4FD66" author="Smetana" box="[977,1259,620,645]" pageId="4" pageNumber="515" refString="Smetana, A. & Smith, A. B. T. (2006) New acts and comments. Scarabaeidae: Cetoniinae. In: Lobl, I. & Smetana, A. (eds.), Catalogue of Palaearctic Coleoptera Folume 3. Apollo Books, Stenstrup, Denmark, pp. 38 - 39." type="journal article" year="2006">Smetana & Smith 2006</bibRefCitation>
|
||
). The incorrect spelling was followed by many authors (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2C4BFEA3F13DFD4B" author="Schoch" box="[605,770,655,680]" pageId="4" pageNumber="515" refString="Schoch, G. (1896 b) Catalogus systematicus Cetonidarum et Trichiidarum ad huc cognitarum. W. Junk, Zurich, Switzerland, 95 pp." type="book" year="1896" yearSuffix="b">Schoch 1896b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D1BFEBCF1A8FD4B" author="Moser" box="[781,919,656,680]" pageId="4" pageNumber="515" refString="Moser, J. (1902) Neue Cetoniden-Arten aus Tonkin, gesammelt von H. Fruhstorfer. Berliner entomologische Zeitschrift, 46, 525 - 538." type="journal article" year="1902">Moser 1902</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2DB4FEBCF60EFD4B" author="Janson" box="[930,1073,656,680]" pageId="4" pageNumber="515" refString="Janson, O. E. (1917) Additions to the knowledge of the Cetoniidae of British India. Transactions of the Entomological Society of London, 65, 102 - 111. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01404. x" type="journal article" year="1917">Janson 1917</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2A2AFEA3F6C3FD4B" author="Schenkling" box="[1084,1276,655,680]" pageId="4" pageNumber="515" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, Germany, 431 pp." type="book" year="1921">Schenkling 1921</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2B1EFEA3F7A7FD4B" author="Miksic" box="[1288,1432,655,680]" pageId="4" pageNumber="515" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, Bosnia and Herzegovina, 444 pp." type="book" year="1976">Mikšić 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2E81FE98F3BBFD2F" author="Miwa" box="[151,388,692,717]" pageId="4" pageNumber="515" refString="Miwa, Y. & Chujo, M. (1939) Catalogus Coleopterum Japonicorm, Pars 5, Scarabaeidae. Noda-Syobo, Taihoku, Taiwan, 94 pp." type="book" year="1939">Miwa & Chûjô 1939</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2F86FE98F018FD2E" author="Paulian" box="[400,551,692,717]" pageId="4" pageNumber="515" refString="Paulian, R. (1960) Coleopteres Scarabeides de L'Indochine (Rutelines et Cetonines) (Suite). Annales de la Societe entomologique de France, 129, 1 - 87." type="journal article" year="1960">Paulian 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2C24FE98F0EFFD2E" author="Antoine" box="[562,720,692,717]" pageId="4" pageNumber="515" refString="Antoine, P. (1986) Sur la tribu Taenioderini: remarques taxonomiques et description d'especes nouvelles (Coleoptera, Cetoniidae). Bulletin de la Societe Sciences Nat, 49, 9 - 20." type="journal article" year="1986">Antoine 1986</bibRefCitation>
|
||
), some attributing authorship to Bumeister (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2AA9FE98F76EFD2F" author="Miksic" box="[1215,1361,692,717]" pageId="4" pageNumber="515" refString="Miksic, R. (1972). Revision der Gattung Coelodera Burmeister (73. Beitrag zur Kenntnis der Scarabaeiden). Revue suisse de Zoologie, 79, 675 - 707." type="journal article" year="1972">Mikšić 1972</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2B4AFE98F7A8FD2E" author="Miksic" box="[1372,1431,692,717]" pageId="4" pageNumber="515" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, Bosnia and Herzegovina, 444 pp." type="book" year="1976">1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2E81FEF4F2C1FD13" author="Ma" box="[151,254,727,752]" pageId="4" pageNumber="515" refString="Ma, W. Z. (1995) Economic Insect Fauna of China, Folume 46 Coleoptera: Cetoniidae, Trichiidae, and Falgidae. Science and Technology Press, Beijing, China, 210 pp. [In Chinese]" type="book" year="1995">Ma 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2F1FFEFBF36BFD13" author="Ma" box="[265,340,727,752]" pageId="4" pageNumber="515" refString="Ma, W. Z. (2002 b) Coleoptera: Cetoniidae. In: Huang, B. - K. (ed.), Fauna of Insects in Fujian Province of China, Folume 6. Fujian Science and Technology Press, Fuzhou, China, pp. 364 - 382. [In Chinese]" type="journal article" year="2002" yearSuffix="b">2002b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDBDA1CFFD0FC4E2ED1FED0F2D6FB26" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="515">
|
||
Considering their ambiguous definition,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2CD4FED0F156FCF7" author="Arrow" box="[706,873,764,788]" pageId="4" pageNumber="515" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor & Francis, London, United Kingdom, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
|
||
combined the genus
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A9DFED0F6C2FCF6" authorityName="Hope" authorityYear="1831" box="[1163,1277,764,789]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2A9DFED0F6C2FCF6" box="[1163,1277,764,789]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
and several Taenioderina genera according to their uncovered abdominal sides, and used the earliest name
|
||
<taxonomicName id="4C64A19FFFD0FC4E2B36FF0DF3B9FCBF" authority="Hoffmannsegg, 1817" authorityName="Hoffmannsegg" authorityYear="1817" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2B36FF0DF7A4FCDB" box="[1312,1435,801,824]" italics="true" pageId="4" pageNumber="515">Macronota</emphasis>
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2E81FF68F3B9FCBF" author="Hoffmannsegg" box="[151,390,836,861]" pageId="4" pageNumber="515" refString="Hoffmannsegg, J. C. (1817) Entomologische Bemerkungen bei Gelegenheit Abhandlungen der Amerikanischer Insekten. Zoologisches Magazin, 1, 8 - 56." type="journal article" year="1817">Hoffmannsegg, 1817</bibRefCitation>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C64A19FFFD0FC4E2FDBFF69F160FCBF" authority="Gory & Percheron, 1833" authorityName="Gory & Percheron" authorityYear="1833" box="[461,863,836,861]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="diardi">
|
||
<emphasis id="B910060EFFD0FC4E2FDBFF69F00AFCBE" box="[461,565,836,861]" italics="true" pageId="4" pageNumber="515">C. diardi</emphasis>
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2C29FF68F160FCBF" author="Gory" box="[575,863,836,861]" pageId="4" pageNumber="515" refString="Gory, M. H. & Percheron, M. A. (1833) Monographie des Cetoines et genres voisins, formant dans les familles naturelles de Latreille, le division des Scarabees Melitophiles. J. B. Baillieres, Paris, France, 410 pp." type="book" year="1833">Gory & Percheron, 1833</bibRefCitation>
|
||
</taxonomicName>
|
||
erroneously cited as the
|
||
<typeStatus id="54DF64BEFFD0FC4E2A91FF69F688FCBE" box="[1159,1207,837,861]" pageId="4" pageNumber="515">type</typeStatus>
|
||
species. The genus
|
||
<taxonomicName id="4C64A19FFFD0FC4E2E81FF45F32CFC63" authorityName="Hoffmannsegg" authorityYear="1817" box="[151,275,873,896]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2E81FF45F32CFC63" box="[151,275,873,896]" italics="true" pageId="4" pageNumber="515">Macronota</emphasis>
|
||
</taxonomicName>
|
||
was erected without any species (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2CB4FF4BF1B2FC63" author="Hoffmannsegg" box="[674,909,871,896]" pageId="4" pageNumber="515" refString="Hoffmannsegg, J. C. (1817) Entomologische Bemerkungen bei Gelegenheit Abhandlungen der Amerikanischer Insekten. Zoologisches Magazin, 1, 8 - 56." type="journal article" year="1817">Hoffmannsegg 1817</bibRefCitation>
|
||
) until
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2DCBFF4BF694FC63" author="Wiedmann" box="[989,1195,871,896]" pageId="4" pageNumber="515" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen, und dem Vorgebirge der guten Hoffnung. Zoologische Magazin, 2, 1 - 133." type="journal article" year="1823">Wiedmann (1823)</bibRefCitation>
|
||
described three new species, hence Arrow’s designation is not valid, and the
|
||
<typeStatus id="54DF64BEFFD0FC4E2D00FFA1F179FC46" box="[790,838,909,933]" pageId="4" pageNumber="515">type</typeStatus>
|
||
species can only be one of the three species (Article 67.2, International Commission on Zoological Nomenclature 1999). Unfortunately, two of them were already transferred to
|
||
<taxonomicName id="4C64A19FFFD0FC4E2F2BFFF8F39DFC0E" authorityName="Burmeister" authorityYear="1842" box="[317,418,980,1005]" class="Insecta" family="Cetoniidae" genus="Clinteria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2F2BFFF8F39DFC0E" box="[317,418,980,1005]" italics="true" pageId="4" pageNumber="515">Clinteria</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2FC0FFF8F094FC0F" author="Burmeister" box="[470,683,980,1005]" pageId="4" pageNumber="515" refString="Burmeister, H. C. C. (1842) Handbuch der Entomologie, Band III. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Melitophila. T. C. F. Enslin, Berlin, Germany, 828 pp." type="book" year="1842">Burmeister (1842)</bibRefCitation>
|
||
, and the remaining species,
|
||
<taxonomicName id="4C64A19FFFD0FC4E2DE8FFF9F691FC0E" box="[1022,1198,980,1005]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="anthracinus">
|
||
<emphasis id="B910060EFFD0FC4E2DE8FFF9F691FC0E" box="[1022,1198,980,1005]" italics="true" pageId="4" pageNumber="515">M. anthracinus</emphasis>
|
||
</taxonomicName>
|
||
, is conspecific with
|
||
<taxonomicName id="4C64A19FFFD0FC4E2E81FFDBF068FBF3" authority="(Frolich, 1792)" baseAuthorityName="Frolich" baseAuthorityYear="1792" box="[151,599,1015,1040]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="nigritus">
|
||
<emphasis id="B910060EFFD0FC4E2E81FFDBF39BFBF3" box="[151,420,1015,1040]" italics="true" pageId="4" pageNumber="515">Thaumastopeus nigritus</emphasis>
|
||
(Frölich, 1792)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C64A19FFFD0FC4E2C74FFD5F0E1FBF3" authorityName="Hoffmannsegg" authorityYear="1817" box="[610,734,1017,1040]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2C74FFD5F0E1FBF3" box="[610,734,1017,1040]" italics="true" pageId="4" pageNumber="515">Macronota</emphasis>
|
||
</taxonomicName>
|
||
thus became a synonym of
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A02FFDBF6FBFBF3" authorityName="Kraatz" authorityYear="1885" box="[1044,1220,1015,1040]" class="Insecta" family="Cetoniidae" genus="Thaumastopeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2A02FFDBF6FBFBF3" box="[1044,1220,1015,1040]" italics="true" pageId="4" pageNumber="515">Thaumastopeus</emphasis>
|
||
</taxonomicName>
|
||
, leaving
|
||
<taxonomicName id="4C64A19FFFD0FC4E2B3CFFDBF7A3FBF3" authorityName="Hope" authorityYear="1831" box="[1322,1436,1015,1040]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2B3CFFDBF7A3FBF3" box="[1322,1436,1015,1040]" italics="true" pageId="4" pageNumber="515">Coilodera</emphasis>
|
||
</taxonomicName>
|
||
as the oldest available name for this genus (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2C89F830F179FBD7" author="Kuntzen" box="[671,838,1052,1076]" pageId="4" pageNumber="515" refString="Kuntzen, H. (1929) Cetoniienstudien (Asiatische Clinterien, I Teil). Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, pp. 8 - 10, pp. 166 - 188." type="book chapter" year="1929">Kuntzen 1929</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D40F830F1D9FBD6" author="Arrow" box="[854,998,1052,1077]" pageId="4" pageNumber="515" refString="Arrow, G. J. (1946) Synonymic notes on some Oriental Cetoniine beetles, with descriptions of two new species. Proceedings of the Royal Entomological Society of London (B), 15, 140 - 144. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 01399. x" type="journal article" year="1946">Arrow 1946</bibRefCitation>
|
||
). The genus
|
||
<taxonomicName id="4C64A19FFFD0FC4E2A96F831F6C4FBD7" authorityName="Hoffmannsegg" authorityYear="1817" box="[1152,1275,1053,1076]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910060EFFD0FC4E2A96F831F6C4FBD7" box="[1152,1275,1053,1076]" italics="true" pageId="4" pageNumber="515">Macronota</emphasis>
|
||
</taxonomicName>
|
||
was not well defined, and Arrow’s combination produced several secondary homonyms that were originally placed in different genera, so this controversial treatment was only followed by few authors (
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2DF4F848F644FB9E" author="Paulian" box="[994,1147,1124,1149]" pageId="4" pageNumber="515" refString="Paulian, R. (1960) Coleopteres Scarabeides de L'Indochine (Rutelines et Cetonines) (Suite). Annales de la Societe entomologique de France, 129, 1 - 87." type="journal article" year="1960">Paulian 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2A9EF849F7A8FB9E" author="De" box="[1160,1431,1124,1149]" pageId="4" pageNumber="515" refString="De Lisle, M. O. & Chujo, M. (1964) Coleoptera from Southeast Asia (III) family Scarabaeidae (3), pp. 190 - 193. In: Kira, T. and Umesao, T. (eds.) Nature and Life in Southeast Asia Folume 3. Fauna and Flora Research Society, Kyoto, Japan." type="journal article" year="1964">De Lisle & Chûjô 1964</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2E81F8A5F325FB43" author="Ruter" box="[151,282,1160,1184]" pageId="4" pageNumber="515" refString="Ruter, G. (1972) Contribution a l'etude des Cetonides (Coleoptera: Cetonidae). Annales de la Societe entomologique de France (new series), 8, 55 - 71." type="journal article" year="1972">Ruter 1972</bibRefCitation>
|
||
), and most followed the subdivided arrangement in
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2D63F8ABF614FB43" author="Schoch" box="[885,1067,1159,1184]" pageId="4" pageNumber="515" refString="Schoch, G. (1896 b) Catalogus systematicus Cetonidarum et Trichiidarum ad huc cognitarum. W. Junk, Zurich, Switzerland, 95 pp." type="book" year="1896" yearSuffix="b">Schoch (1896b)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2A2CF8ABF731FB43" author="Schenkling" box="[1082,1294,1159,1184]" pageId="4" pageNumber="515" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, Germany, 431 pp." type="book" year="1921">Schenkling (1921)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFF5A7EDFFD0FC4E2B58F8ABF2DBFB26" author="Miksic" pageId="4" pageNumber="515" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, Bosnia and Herzegovina, 444 pp." type="book" year="1976">Mikšić (1976)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |