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<document id="AAAA52ACD6E5908FE84FA0662554DA04" ID-CLB-Dataset="46923" ID-DOI="10.5281/zenodo.199938" ID-GBIF-Dataset="c449672a-3692-4687-9814-96dec46b4f33" ID-ISSN="1175-5326" ID-Zenodo-Dep="199938" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460350840799" checkinUser="plazi" docAuthor="Shidlovskiy, Konstantin M., Watters, Brian R. &amp; Wildekamp, Rudolf H." docDate="2010" docId="C34987E7FFBFFFB3E380F9148F51FF41" docLanguage="en" docName="zt02724p057.pdf" docOrigin="Zootaxa 2724" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nothobranchius krysanovi Shidlovskiy, Watters &amp; Wildekamp, 2010, sp. nov." docType="treatment" docVersion="11" lastPageNumber="49" masterDocId="3F70FF9FFFB7FFBFE317FF868A00FFBD" masterDocTitle="Notes on the annual killifish species Nothobranchius rachovii (Cyprinodontiformes; Nothobranchiidae) with the description of two new species" masterLastPageNumber="57" masterPageNumber="37" pageNumber="45" updateTime="1698293656746" updateUser="ExternalLinkService">
<mods:mods id="52DC89F523D8BA274A0997ED38383CA3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B4B10BA336A0EC5C5FC6215CC0305A85">Notes on the annual killifish species Nothobranchius rachovii (Cyprinodontiformes; Nothobranchiidae) with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="2AA93A9ACC657D56B302078D88909F0F" type="personal">
<mods:role id="63EC6D860BDA7F9BB9D4BC97C188D86F">
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</mods:role>
<mods:namePart id="4E91F8B7E0937DF1CE28506427268722">Shidlovskiy, Konstantin M.</mods:namePart>
</mods:name>
<mods:name id="7B9970B120EF3EFB8DC7D0C1440BC72F" type="personal">
<mods:role id="BCEE8D7F4212E01786E71AE05A863D42">
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<mods:namePart id="44BFC9BB4CE64F1E8F2B0DC966312905">Watters, Brian R.</mods:namePart>
</mods:name>
<mods:name id="34C89919FCD3E67A72826B702AF535C7" type="personal">
<mods:role id="13ABD6090564F4859856F9B8D4DC328E">
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</mods:role>
<mods:namePart id="0E1012B60E9AB9B7E5075171E243DFE3">Wildekamp, Rudolf H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1345463DE0825C744789A7F9FD766903">text</mods:typeOfResource>
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<mods:title id="577DE8BBFFE8D938FA0A7DCE8657FED7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="97B10A9EE6F7E450411ED49B27B5CD1D">
<mods:date id="36BBDE3004A26C9E920B04749D0E40BB">2010</mods:date>
<mods:detail id="365DDF2C7BE8D8E65580F20631A4A1AB" type="volume">
<mods:number id="BDC54476666BACA859D11992C036AA63">2724</mods:number>
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<mods:classification id="5C0215BE744AD1728F834A5EAA491F6F">journal article</mods:classification>
<mods:identifier id="4CF5456D46571EF7A476813AA55B719F" type="CLB-Dataset">46923</mods:identifier>
<mods:identifier id="16D5DE0F31EB28AF16895A6BFDD79404" type="DOI">10.5281/zenodo.199938</mods:identifier>
<mods:identifier id="2B9D3D92AD7A44DE56381949D5187F0D" type="GBIF-Dataset">c449672a-3692-4687-9814-96dec46b4f33</mods:identifier>
<mods:identifier id="AF41882AA6635753690A9076E25AA34F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="43AEEFA505A7D8A22FA1659E1A781C0E" type="Zenodo-Dep">199938</mods:identifier>
</mods:mods>
<treatment id="C34987E7FFBFFFB3E380F9148F51FF41" ID-DOI="http://doi.org/10.5281/zenodo.6205626" ID-GBIF-Taxon="119407862" ID-Zenodo-Dep="6205626" LSID="urn:lsid:plazi:treatment:C34987E7FFBFFFB3E380F9148F51FF41" httpUri="http://treatment.plazi.org/id/C34987E7FFBFFFB3E380F9148F51FF41" lastPageId="12" lastPageNumber="49" pageId="8" pageNumber="45">
<subSubSection id="03FA657AFFBFFFB7E380F9148B2EF972" pageId="8" pageNumber="45" type="nomenclature">
<paragraph id="4B5F36F1FFBFFFB7E380F914883DF911" blockId="8.[151,573,1682,1743]" box="[151,573,1682,1708]" pageId="8" pageNumber="45">
<heading id="1017819DFFBFFFB7E380F914883DF911" bold="true" box="[151,573,1682,1708]" fontSize="11" level="1" pageId="8" pageNumber="45" reason="1">
<emphasis id="7994EAE3FFBFFFB7E380F914883DF911" bold="true" box="[151,573,1682,1708]" pageId="8" pageNumber="45">
<taxonomicName id="8CE04D72FFBFFFB7E380F9148BD8F911" ID-CoL="47SMG" box="[151,472,1682,1708]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="krysanovi" status="sp. nov.">
<emphasis id="7994EAE3FFBFFFB7E380F9148BD8F911" bold="true" box="[151,472,1682,1708]" italics="true" pageId="8" pageNumber="45">Nothobranchius krysanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62A75798FFBFFFB7E2F7F914883DF911" box="[480,573,1682,1708]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4B5F36F1FFBFFFB7E380F9338B2EF972" blockId="8.[151,573,1682,1743]" box="[151,302,1717,1743]" pageId="8" pageNumber="45">
(
<figureCitation id="D3DB2A74FFBFFFB7E388F9338AF1F972" box="[159,241,1717,1743]" captionStart="FIGURE 4" captionStartId="11.[151,257,1031,1055]" captionTargetBox="[151,1436,194,1009]" captionTargetId="figure@11.[151,1436,194,1010]" captionTargetPageId="11" captionText="FIGURE 4. Nothobranchius krysanovi sp. nov., male ~ 31 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04 - 10). Photograph: R. Wildekamp." httpUri="https://zenodo.org/record/199942/files/figure.png" pageId="8" pageNumber="45">Figs. 4</figureCitation>
&amp;
<figureCitation id="D3DB2A74FFBFFFB7E201F9338B25F972" box="[278,293,1717,1743]" captionStart="FIGURE 5" captionStartId="11.[151,256,1768,1792]" captionTargetBox="[337,1249,1138,1746]" captionTargetId="figure@11.[337,1249,1138,1747]" captionTargetPageId="11" captionText="FIGURE 5. Nothobranchius krysanovi sp. nov., female ~ 27 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04 - 10). Photograph: B. Watters." httpUri="https://zenodo.org/record/199943/files/figure.png" pageId="8" pageNumber="45">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="03FA657AFFBFFFB7E380F8868926F8AA" box="[151,806,1792,1815]" pageId="8" pageNumber="45" type="reference_group">
<paragraph id="4B5F36F1FFBFFFB7E380F8868926F8AA" blockId="8.[151,806,1792,1815]" box="[151,806,1792,1815]" pageId="8" pageNumber="45">
<emphasis id="7994EAE3FFBFFFB7E380F8868BA9F8AA" box="[151,425,1792,1815]" italics="true" pageId="8" pageNumber="45">
<taxonomicName id="8CE04D72FFBFFFB7E380F8868BA2F8AA" ID-CoL="47SNR" box="[151,418,1792,1815]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="rachovii">Nothobranchius rachovii</taxonomicName>
,
</emphasis>
non
<bibRefCitation id="2F714B00FFBFFFB7E2C9F886884CF8AA" author="Ahl" box="[478,588,1792,1815]" pageId="8" pageNumber="45" refString="Ahl, E. 1926. Neue oder selten importierte Fische, III. Blatter fur Aquarien- und Terrarienkunde, 37 (14): 346 - 348." type="journal article" year="1926">Ahl, 1926</bibRefCitation>
:
<bibRefCitation id="2F714B00FFBFFFB7E140F8868926F8AA" author="Watters" box="[599,806,1792,1815]" pageId="8" pageNumber="45" refString="Watters, B., Wildekamp, R. H. &amp; Cooper, B. J. 2009. Nothobranchius rachovii Ahl, 1926 - A historical review. Journal of the American Killifish Association, 42: 105 - 127." type="journal article" year="2009">
Watters
<emphasis id="7994EAE3FFBFFFB7E1B9F88688E4F8AA" box="[686,740,1792,1815]" italics="true" pageId="8" pageNumber="45">et al.</emphasis>
, 2009
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="03FA657AFFBFFFB6E380F8C388C3FF6A" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45" type="materials_examined">
<paragraph id="4B5F36F1FFBFFFB7E380F8C3883FF817" blockId="8.[151,1437,1861,2036]" pageId="8" pageNumber="45">
<emphasis id="7994EAE3FFBFFFB7E380F8C38B12F8E2" bold="true" box="[151,274,1861,1887]" pageId="8" pageNumber="45">
<typeStatus id="945B8853FFBFFFB7E380F8C38B0DF8E2" box="[151,269,1861,1887]" pageId="8" pageNumber="45" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="2DF1AE34FFBFFFB7E20CF8C38B73F8E2" box="[283,371,1861,1887]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="8" pageNumber="45">MRAC</collectionCode>
A4-039-P-0078, male,
<quantity id="8C189B14FFBFFFB7E198F8C388FAF8E2" box="[655,762,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="8" pageNumber="45" unit="mm" value="31.2">31.2 mm</quantity>
SL,
<collectingCountry id="33F77661FFBFFFB7E022F8C389D6F8E2" box="[821,982,1861,1887]" name="Mozambique" pageId="8" pageNumber="45">Mozambique</collectingCountry>
,
<quantity id="8C189B14FFBFFFB7E0F5F8C38E2EF8E2" box="[994,1070,1861,1887]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="45" unit="km" value="12.0">12 km</quantity>
S of Nicuadala in direction of Quelimane, Licungo River system,
<geoCoordinate id="2ED45036FFBFFFB7E13DF8EC88A9F839" box="[554,681,1898,1924]" direction="south" orientation="latitude" pageId="8" pageNumber="45" precision="92" value="-17.658333">17° 39.5' S</geoCoordinate>
,
<geoCoordinate id="2ED45036FFBFFFB7E1A3F8EC8935F839" box="[692,821,1898,1924]" direction="east" orientation="longitude" pageId="8" pageNumber="45" precision="92" value="36.858334">36° 51.5' E</geoCoordinate>
, B. Watters, B. Cooper, R. Wildekamp, J. Jordaan, J. Bornman and J. Ippel,
<date id="3F5E1031FFBFFFB7E2B6F816883BF817" box="[417,571,1936,1962]" pageId="8" pageNumber="45" value="2004-04-06">6 April 2004</date>
.
</paragraph>
<paragraph id="4B5F36F1FFBFFFB6E3D1F83388C3FF6A" blockId="8.[151,1437,1861,2036]" lastBlockId="9.[151,1437,152,2007]" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45">
<emphasis id="7994EAE3FFBFFFB7E3D1F8338B47F872" bold="true" box="[198,327,1973,1999]" pageId="8" pageNumber="45">
<typeStatus id="945B8853FFBFFFB7E3D1F8338B42F872" box="[198,322,1973,1999]" pageId="8" pageNumber="45" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="2DF1AE34FFBFFFB7E245F8338BA9F872" box="[338,425,1973,1999]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="8" pageNumber="45">MRAC</collectionCode>
A4-039-P0079-0080,
<specimenCount id="5DE6FD78FFBFFFB7E1AEF833891AF872" box="[697,794,1973,1999]" pageId="8" pageNumber="45" type="male">2 males</specimenCount>
, 24.3–26.0 mm SL, collected with
<typeStatus id="945B8853FFBFFFB7E7DBF8338F36F872" box="[1228,1334,1973,1999]" pageId="8" pageNumber="45" type="holotype">holotype</typeStatus>
;
<collectionCode id="2DF1AE34FFBFFFB7E652F8338F9CF872" box="[1349,1436,1973,1999]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="8" pageNumber="45">MRAC</collectionCode>
A4-039-P-0079, male,
<quantity id="8C189B14FFBFFFB7E2BBF85C8817F849" box="[428,535,2010,2036]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="8" pageNumber="45" unit="mm" value="28.8">28.8 mm</quantity>
SL,
<specimenCount id="5DE6FD78FFBFFFB7E145F85C88C7F849" box="[594,711,2010,2036]" pageId="8" pageNumber="45" type="female">4 females</specimenCount>
<quantity id="8C189B14FFBFFFB7E1D8F85C897DF849" box="[719,893,2010,2036]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" metricValueMax="3.02" metricValueMin="2.7199999999999998" pageId="8" pageNumber="45" unit="mm" value="28.7" valueMax="30.2" valueMin="27.2">27.2–30.2 mm</quantity>
SL,
<collectingCountry id="33F77661FFBFFFB7E0AEF85C8E5AF849" box="[953,1114,2010,2036]" name="Mozambique" pageId="8" pageNumber="45">Mozambique</collectingCountry>
,
<quantity id="8C189B14FFBFFFB7E771F85C8EB2F849" box="[1126,1202,2010,2036]" metricMagnitude="4" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="45" unit="km" value="84.0">84 km</quantity>
from Caia ferry on road to Nicuadala, Lima River system,
<geoCoordinate id="2ED45036FFBEFFB6E16DFF1E8903FF0F" box="[634,771,152,178]" direction="south" orientation="latitude" pageId="9" pageNumber="46" precision="92" value="-17.563334">17° 33.8' S</geoCoordinate>
,
<geoCoordinate id="2ED45036FFBEFFB6E006FF1E899DFF0F" box="[785,925,152,178]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="92" value="36.073334">36° 04.4' E</geoCoordinate>
, B. Watters, B. Cooper, R. Wildekamp, J. Jordaan, J. Bornman and J. Ippel,
<date id="3F5E1031FFBEFFB6E132FF3B88BFFF6A" box="[549,703,189,215]" pageId="9" pageNumber="46" value="2004-04-05">5 April 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="03FA657AFFBEFFB6E3D1FF648B67FD2A" pageId="9" pageNumber="46" type="diagnosis">
<paragraph id="4B5F36F1FFBEFFB6E3D1FF6488B6FD9A" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
<emphasis id="7994EAE3FFBEFFB6E3D1FF648B46FF41" bold="true" box="[198,326,226,252]" pageId="9" pageNumber="46">Diagnosis.</emphasis>
Morphology similar to
<taxonomicName id="8CE04D72FFBEFFB6E166FF6488F9FF46" box="[625,761,226,251]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="rachovii">
<emphasis id="7994EAE3FFBEFFB6E166FF6488F9FF46" box="[625,761,226,251]" italics="true" pageId="9" pageNumber="46">N. rachovii</emphasis>
</taxonomicName>
.
<taxonomicName id="8CE04D72FFBEFFB6E01BFF648E47FF46" box="[780,1095,226,251]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="krysanovi">
<emphasis id="7994EAE3FFBEFFB6E01BFF648E47FF46" box="[780,1095,226,251]" italics="true" pageId="9" pageNumber="46">Nothobranchius krysanovi</emphasis>
</taxonomicName>
is distinguished from other species of the genus by the following combination of characters: males with bright colouration consisting of alternating light blue and orange-red oblique bars on body, dorsal and anal fins, and orange-brown caudal fin with orange subterminal and black terminal band. Female with faint brown oblique bars on posterior part of flanks. It can be distinguished from the other species of the subgenus
<taxonomicName id="8CE04D72FFBEFFB6E0F0FEFE8EA9FE2C" box="[999,1193,376,401]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="7994EAE3FFBEFFB6E0F0FEFE8EA9FE2C" box="[999,1193,376,401]" italics="true" pageId="9" pageNumber="46">Nothobranchius</emphasis>
</taxonomicName>
as follows: from
<taxonomicName id="8CE04D72FFBEFFB6E696FEFE8AE6FE0B" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="furzeri">
<emphasis id="7994EAE3FFBEFFB6E696FEFE8AE6FE0B" italics="true" pageId="9" pageNumber="46">N. furzeri</emphasis>
</taxonomicName>
by male colouration, higher number of dorsal fin rays (15–17 vs. 14 or 15); from
<taxonomicName id="8CE04D72FFBEFFB6E7ABFE1B8F35FE0B" box="[1212,1333,413,438]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="kadleci">
<emphasis id="7994EAE3FFBEFFB6E7ABFE1B8F35FE0B" box="[1212,1333,413,438]" italics="true" pageId="9" pageNumber="46">N. kadleci</emphasis>
</taxonomicName>
by male colouration and shape of the frontal region, higher number of dorsal and anal fin rays (15–17 vs. 13 or 14 and 15–17 vs. 13 or 14) and from
<taxonomicName id="8CE04D72FFBEFFB6E2EBFE6E889FFDBC" box="[508,671,488,513]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="orthonotus">
<emphasis id="7994EAE3FFBEFFB6E2EBFE6E889FFDBC" box="[508,671,488,513]" italics="true" pageId="9" pageNumber="46">N. orthonotus</emphasis>
</taxonomicName>
by male and female colouration, and lower number of scales on the mid-longitudinal series (27–29 vs. 28–33).
</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D1FDB48B67FD2A" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
<taxonomicName id="8CE04D72FFBEFFB6E3D1FDB48800FDF6" box="[198,512,562,587]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="krysanovi">
<emphasis id="7994EAE3FFBEFFB6E3D1FDB48800FDF6" box="[198,512,562,587]" italics="true" pageId="9" pageNumber="46">Nothobranchius krysanovi</emphasis>
</taxonomicName>
can also be distinguished from
<taxonomicName id="8CE04D72FFBEFFB6E09DFDB48E11FDF6" box="[906,1041,562,587]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="rachovii">
<emphasis id="7994EAE3FFBEFFB6E09DFDB48E11FDF6" box="[906,1041,562,587]" italics="true" pageId="9" pageNumber="46">N. rachovii</emphasis>
</taxonomicName>
by its higher number of diploid chromosomes (2n = 18 vs. 2n = 16); and from
<taxonomicName id="8CE04D72FFBEFFB6E1A1FDDE893BFDCC" box="[694,827,600,625]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="pienaari">
<emphasis id="7994EAE3FFBEFFB6E1A1FDDE893BFDCC" box="[694,827,600,625]" italics="true" pageId="9" pageNumber="46">N. pienaari</emphasis>
</taxonomicName>
by its much lower diploid chromosome number (2n = 18 vs. 2n = 34).
</paragraph>
</subSubSection>
<subSubSection id="03FA657AFFBEFFB6E3D1FD248BE7F8A1" pageId="9" pageNumber="46" type="description">
<paragraph id="4B5F36F1FFBEFFB6E3D1FD248883FC91" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
<emphasis id="7994EAE3FFBEFFB6E3D1FD248B5AFD01" bold="true" box="[198,346,674,700]" pageId="9" pageNumber="46">Description.</emphasis>
Morphometric data are presented in Table 3. Robust
<taxonomicName id="8CE04D72FFBEFFB6E0C5FD248E8AFD06" box="[978,1162,674,699]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="7994EAE3FFBEFFB6E0C5FD248E8AFD06" box="[978,1162,674,699]" italics="true" pageId="9" pageNumber="46">Nothobranchius</emphasis>
</taxonomicName>
of medium size. Snout pointed, mouth terminal, directed slightly upward. Body laterally compressed and deep. Dorsal profile concave on head, convex from nape to end of dorsal fin; more convex in older males than in younger specimens. Profile of caudal peduncle nearly straight.
</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D2FCBE8F50FC21" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">Supraorbital squamation G-type with frontal part partly covered with epidermal tissue. Frontal and central supraorbital neuromast systems fused and forming two distinct shallow grooves. Both lined with three shallow lobes at both sides of the groove. Posterior cephalic neuromast systems in two curved grooves.</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D2FC2E8B46FB8F" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">Dorsal fin rays 15–17, anal fin rays 15–17. Scales on the median longitudinal line 27–29+3 on caudal-fin base, most with shallow pit with one neuromast. Transverse row of scales above pelvic fins 11, circumpeduncular scales 12. Pelvic fins short, not reaching origin of anal fin. Pectoral fins reaching to first pelvic fin rays.</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D2FBBB8F71FB1F" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
Male: maximum size observed
<quantity id="8C189B14FFBEFFB6E15CFBBB88B8FBEA" box="[587,696,1085,1111]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="9" pageNumber="46" unit="mm" value="31.2">31.2 mm</quantity>
SL. Body laterally compressed and deep. All unpaired fins rounded, dorsal and anal fin covered with epidermal tissue. Papillae on the dorsal and anal fin rays, tips projecting through epidermis. Opercular membrane projecting from opercle, distal edge slightly wrinkled.
</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D2FB2B8863FA8A" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
Female: smaller than male, maximum size observed
<quantity id="8C189B14FFBEFFB6E03EFB2B8991FB7A" box="[809,913,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="9" pageNumber="46" unit="mm" value="30.2">30.2 mm</quantity>
SL. Body less deep than in male. Dorsal and caudal fins rounded, anal fin triangular, tip rounded, rays 3–7 longer and more rigid. Anal fin positioned more posteriorly than in male. No epidermal tissue covering dorsal and anal fins or papillae on rays. Opercular membrane not projecting from opercle.
</paragraph>
<paragraph id="4B5F36F1FFBEFFB6E3D2FAC48BE7F8A1" blockId="9.[151,1437,152,2007]" pageId="9" pageNumber="46">
<emphasis id="7994EAE3FFBEFFB6E3D2FAC48B67FAE1" bold="true" box="[197,359,1346,1372]" pageId="9" pageNumber="46">Colouration.</emphasis>
Live male (see
<figureCitation id="D3DB2A74FFBEFFB6E13DFAC48875FAE1" box="[554,629,1346,1372]" captionStart="FIGURE 4" captionStartId="11.[151,257,1031,1055]" captionTargetBox="[151,1436,194,1009]" captionTargetId="figure@11.[151,1436,194,1010]" captionTargetPageId="11" captionText="FIGURE 4. Nothobranchius krysanovi sp. nov., male ~ 31 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04 - 10). Photograph: R. Wildekamp." httpUri="https://zenodo.org/record/199942/files/figure.png" pageId="9" pageNumber="46">Fig. 4</figureCitation>
): body colour iridescent light-blue. Scales with distinct orange-red margins. Margin of some scales on posterior part of body and caudal peduncle more distinct, forming oblique bars, lower end forward. Snout and head carmine-red to orange-red, throat orange-red. Back anterior to dorsal fin carmine-red or brown-red. Operculum with 2 or 3 orange-red oblique stripes. Projecting part of branchiostegal membrane red to orange-red. Iris yellow-bronze to gold with incomplete dark vertical bar. Caudal fin orange-brown in the inner three quarters followed by a lighter orange subterminal band and black margin. Dorsal fin bright blue with red or red-brown spots, larger in proximal part and smaller distally. Spots may merge into oblong markings across fin. Dorsal fin with narrow blue-white margin. Anal fin similar to dorsal fin with lighter iridescent blue distal part lacking spots. Ventral fins bright blue with dark red proximal spots. Pectoral fins transparent translucent red with wide blue-white margin. Iris golden with dark vertical bar. Live female (see
<figureCitation id="D3DB2A74FFBEFFB6E27DF93E8BB5F96F" box="[362,437,1720,1746]" captionStart="FIGURE 5" captionStartId="11.[151,256,1768,1792]" captionTargetBox="[337,1249,1138,1746]" captionTargetId="figure@11.[337,1249,1138,1747]" captionTargetPageId="11" captionText="FIGURE 5. Nothobranchius krysanovi sp. nov., female ~ 27 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04 - 10). Photograph: B. Watters." httpUri="https://zenodo.org/record/199943/files/figure.png" pageId="9" pageNumber="46">Fig. 5</figureCitation>
): body colour grey-brown, darker on the back to silver on abdomen. Several faint brown oblique bars, lower end to front, on posterior part of flanks. All fins colourless. Iris bronze or gold with incomplete vertical dark bar.
</paragraph>
</subSubSection>
<subSubSection id="03FA657AFFBEFFB4E3D1F8A18844F865" lastPageId="11" lastPageNumber="48" pageId="9" pageNumber="46" type="distribution">
<paragraph id="4B5F36F1FFBEFFB4E3D1F8A18844F865" blockId="9.[151,1437,152,2007]" lastBlockId="11.[151,1436,1870,2008]" lastPageId="11" lastPageNumber="48" pageId="9" pageNumber="46">
<emphasis id="7994EAE3FFBEFFB6E3D1F8A18805F8FC" bold="true" box="[198,517,1831,1857]" pageId="9" pageNumber="46">Distribution and habitat.</emphasis>
<taxonomicName id="8CE04D72FFBEFFB6E107F8AE8949F8FC" box="[528,841,1832,1857]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="krysanovi">
<emphasis id="7994EAE3FFBEFFB6E107F8AE8949F8FC" box="[528,841,1832,1857]" italics="true" pageId="9" pageNumber="46">Nothobranchius krysanovi</emphasis>
</taxonomicName>
is known from ephemeral pools and swamps on floodplains in coastal lowlands of east-central
<collectingCountry id="33F77661FFBEFFB6E1AAF8CB8958F8DA" box="[701,856,1869,1895]" name="Mozambique" pageId="9" pageNumber="46">Mozambique</collectingCountry>
north of the lower Zambezi River (see
<figureCitation id="D3DB2A74FFBEFFB6E63BF8CB8F72F8DA" box="[1324,1394,1869,1895]" captionStart="FIGURE 3" captionStartId="10.[151,255,1892,1916]" captionTargetBox="[176,1406,196,1876]" captionTargetId="figure@10.[174,1406,194,1880]" captionTargetPageId="10" captionText="FIGURE 3. Distribution of Nothobranchius rachovii (diamond symbol), N. krysanovi sp. nov. (triangle symbol) and N. pienaari sp. nov. (square symbol). Red-filled symbols represent type localities. Note that individual symbols may represent multiple sites where such sites are in close proximity to one another. Question marks beside two of the diamond symbols indicate sites of populations of unconfirmed affinity." httpUri="https://zenodo.org/record/199941/files/figure.png" pageId="9" pageNumber="46">Fig. 3</figureCitation>
). It is presently known only from the basin of Kwa-Kwa River and its tributaries. Water level in the habitats is subject to seasonal changes and generally the water disappears completely during the dry season. Grass vegetation is usually found near the habitat margins. Much of the swamps may be utilised by local inhabitants for the cultivation of rice. Aquatic vegetation may comprise
<taxonomicName id="8CE04D72FFBCFFB4E04BF8C889CBF8DA" box="[860,971,1870,1895]" class="Magnoliopsida" family="Nymphaeaceae" genus="Nymphea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Nymphaeales" pageId="11" pageNumber="48" phylum="Magnoliophyta" rank="genus">
<emphasis id="7994EAE3FFBCFFB4E04BF8C889CBF8DA" box="[860,971,1870,1895]" italics="true" pageId="11" pageNumber="48">Nymphea</emphasis>
</taxonomicName>
,
<taxonomicName id="8CE04D72FFBCFFB4E0CDF8C88E2BF8DA" box="[986,1067,1870,1895]" class="Liliopsida" family="Hydrocharitaceae" genus="Ottelia" kingdom="Plantae" order="Alismatales" pageId="11" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="7994EAE3FFBCFFB4E0CDF8C88E2BF8DA" box="[986,1067,1870,1895]" italics="true" pageId="11" pageNumber="48">Ottelia</emphasis>
</taxonomicName>
,
<taxonomicName id="8CE04D72FFBCFFB4E72DF8C88EDEF8DA" box="[1082,1246,1870,1895]" class="Liliopsida" family="Hydrocharitaceae" genus="Lagarosiphon" kingdom="Plantae" order="Alismatales" pageId="11" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="7994EAE3FFBCFFB4E72DF8C88EDEF8DA" box="[1082,1246,1870,1895]" italics="true" pageId="11" pageNumber="48">Lagarosiphon</emphasis>
</taxonomicName>
and
<taxonomicName id="8CE04D72FFBCFFB4E601F8C88F9BF8DA" box="[1302,1435,1870,1895]" class="Magnoliopsida" family="Lentibulariaceae" genus="Utricularia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="7994EAE3FFBCFFB4E601F8C88F9BF8DA" box="[1302,1435,1870,1895]" italics="true" pageId="11" pageNumber="48">Utricularia</emphasis>
</taxonomicName>
species. Sympatric fish species observed are
<taxonomicName id="8CE04D72FFBCFFB4E1DFF8F58973F831" box="[712,883,1907,1932]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="orthonotus">
<emphasis id="7994EAE3FFBCFFB4E1DFF8F58973F831" box="[712,883,1907,1932]" italics="true" pageId="11" pageNumber="48">N. orthonotus</emphasis>
</taxonomicName>
,
<taxonomicName id="8CE04D72FFBCFFB4E093F8F58E6DF831" box="[900,1133,1907,1932]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="gariepinus">
<emphasis id="7994EAE3FFBCFFB4E093F8F58E6DF831" box="[900,1133,1907,1932]" italics="true" pageId="11" pageNumber="48">Clarias gariepinus</emphasis>
</taxonomicName>
,
<taxonomicName id="8CE04D72FFBCFFB4E769F8F58F95F831" box="[1150,1429,1907,1932]" class="Actinopterygii" family="Anabantidae" genus="Ctenopoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="multispine">
<emphasis id="7994EAE3FFBCFFB4E769F8F58F95F831" box="[1150,1429,1907,1932]" italics="true" pageId="11" pageNumber="48">Ctenopoma multispine</emphasis>
</taxonomicName>
,
<taxonomicName id="8CE04D72FFBCFFB4E380F81F884FF80F" authority="Peters, 1844" authorityName="Peters" authorityYear="1844" box="[151,591,1944,1970]" class="Sarcopterygii" family="Protopteridae" genus="Protopterus" kingdom="Animalia" order="Lepidosireniformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="amphibius">
<emphasis id="7994EAE3FFBCFFB4E380F81F8BA3F80F" box="[151,419,1945,1970]" italics="true" pageId="11" pageNumber="48">Protopterus amphibius</emphasis>
(Peters, 1844)
</taxonomicName>
and unidentified
<taxonomicName id="8CE04D72FFBCFFB4E00AF81F8972F80F" box="[797,882,1945,1970]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="48" phylum="Chordata" rank="genus">
<emphasis id="7994EAE3FFBCFFB4E00AF81F8972F80F" box="[797,882,1945,1970]" italics="true" pageId="11" pageNumber="48">Barbus</emphasis>
</taxonomicName>
and cichlid species.
<taxonomicName id="8CE04D72FFBCFFB4E771F81F8F9CF80F" box="[1126,1436,1945,1970]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="krysanovi">
<emphasis id="7994EAE3FFBCFFB4E771F81F8F9CF80F" box="[1126,1436,1945,1970]" italics="true" pageId="11" pageNumber="48">Nothobranchius krysanovi</emphasis>
</taxonomicName>
has an annual mode of reproduction.
</paragraph>
</subSubSection>
<caption id="1F9F6679FFBDFFB5E380F8E28927F861" httpUri="https://zenodo.org/record/199941/files/figure.png" pageId="10" pageNumber="47" targetBox="[176,1406,196,1876]" targetPageId="10">
<paragraph id="4B5F36F1FFBDFFB5E380F8E28927F861" blockId="10.[151,1437,1892,2012]" pageId="10" pageNumber="47">
<emphasis id="7994EAE3FFBDFFB5E380F8E28B19F8C1" bold="true" box="[151,281,1892,1916]" pageId="10" pageNumber="47">FIGURE 3.</emphasis>
Distribution of
<taxonomicName id="8CE04D72FFBDFFB5E2D1F8E388D2F8C1" box="[454,722,1893,1916]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="rachovii">
<emphasis id="7994EAE3FFBDFFB5E2D1F8E388D2F8C1" box="[454,722,1893,1916]" italics="true" pageId="10" pageNumber="47">Nothobranchius rachovii</emphasis>
</taxonomicName>
(diamond symbol),
<taxonomicName id="8CE04D72FFBDFFB5E0BCF8E38E33F8C1" box="[939,1075,1893,1916]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="krysanovi" status="sp. nov.">
<emphasis id="7994EAE3FFBDFFB5E0BCF8E38E33F8C1" box="[939,1075,1893,1916]" italics="true" pageId="10" pageNumber="47">N. krysanovi</emphasis>
</taxonomicName>
<emphasis id="7994EAE3FFBDFFB5E72EF8E28E8FF8C1" bold="true" box="[1081,1167,1892,1916]" pageId="10" pageNumber="47">
<taxonomicNameLabel id="62A75798FFBDFFB5E72EF8E28E8FF8C1" box="[1081,1167,1892,1916]" pageId="10" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(triangle symbol) and
<taxonomicName id="8CE04D72FFBDFFB5E693F8E38AF6F821" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="pienaari" status="sp. nov.">
<emphasis id="7994EAE3FFBDFFB5E693F8E38AF6F821" italics="true" pageId="10" pageNumber="47">N. pienaari</emphasis>
</taxonomicName>
<emphasis id="7994EAE3FFBDFFB5E3EAF8028B58F821" bold="true" box="[253,344,1924,1948]" pageId="10" pageNumber="47">
<taxonomicNameLabel id="62A75798FFBDFFB5E3EAF8028B58F821" box="[253,344,1924,1948]" pageId="10" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(square symbol). Red-filled symbols represent type localities. Note that individual symbols may represent multiple sites where such sites are in close proximity to one another. Question marks beside two of the diamond symbols indicate sites of populations of unconfirmed affinity.
</paragraph>
</caption>
<caption id="1F9F6679FFBCFFB4E380FB818EECFB82" httpUri="https://zenodo.org/record/199942/files/figure.png" pageId="11" pageNumber="48" targetBox="[151,1436,194,1009]" targetPageId="11">
<paragraph id="4B5F36F1FFBCFFB4E380FB818EECFB82" blockId="11.[151,1436,1031,1087]" pageId="11" pageNumber="48">
<emphasis id="7994EAE3FFBCFFB4E380FB818B1EFBA2" bold="true" box="[151,286,1031,1055]" pageId="11" pageNumber="48">FIGURE 4.</emphasis>
<taxonomicName id="8CE04D72FFBCFFB4E232FB8E884BFBA2" box="[293,587,1032,1055]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="krysanovi" status="sp. nov.">
<emphasis id="7994EAE3FFBCFFB4E232FB8E884BFBA2" box="[293,587,1032,1055]" italics="true" pageId="11" pageNumber="48">Nothobranchius krysanovi</emphasis>
</taxonomicName>
<emphasis id="7994EAE3FFBCFFB4E144FB8188ACFBA2" bold="true" box="[595,684,1031,1055]" pageId="11" pageNumber="48">
<taxonomicNameLabel id="62A75798FFBCFFB4E144FB8188ACFBA2" box="[595,684,1031,1055]" pageId="11" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male ~31 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04-10). Photograph: R. Wildekamp.
</paragraph>
</caption>
<caption id="1F9F6679FFBCFFB4E380F96E8EC2F8A2" httpUri="https://zenodo.org/record/199943/files/figure.png" pageId="11" pageNumber="48" targetBox="[337,1249,1138,1746]" targetPageId="11">
<paragraph id="4B5F36F1FFBCFFB4E380F96E8EC2F8A2" blockId="11.[151,1436,1768,1823]" pageId="11" pageNumber="48">
<emphasis id="7994EAE3FFBCFFB4E380F96E8B1DF8BD" bold="true" box="[151,285,1768,1792]" pageId="11" pageNumber="48">FIGURE 5.</emphasis>
<taxonomicName id="8CE04D72FFBCFFB4E233F96F8844F8BD" box="[292,580,1769,1792]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="krysanovi" status="sp. nov.">
<emphasis id="7994EAE3FFBCFFB4E233F96F8844F8BD" box="[292,580,1769,1792]" italics="true" pageId="11" pageNumber="48">Nothobranchius krysanovi</emphasis>
</taxonomicName>
<emphasis id="7994EAE3FFBCFFB4E15BF96E88A3F8BD" bold="true" box="[588,675,1768,1792]" pageId="11" pageNumber="48">
<taxonomicNameLabel id="62A75798FFBCFFB4E15BF96E88A3F8BD" box="[588,675,1768,1792]" pageId="11" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female ~27 mm SL, topotype, not preserved; Mozambique: south of Nicuadala in the direction of Quelimane, Licungo River system (MOZ 04-10). Photograph: B. Watters.
</paragraph>
</caption>
<subSubSection id="03FA657AFFBBFFB3E3D1FF118F51FF41" pageId="12" pageNumber="49" type="etymology">
<paragraph id="4B5F36F1FFBBFFB3E3D1FF118F51FF41" blockId="12.[151,1436,151,252]" pageId="12" pageNumber="49">
<emphasis id="7994EAE3FFBBFFB3E3D1FF118B53FF0C" bold="true" box="[198,339,151,177]" pageId="12" pageNumber="49">Etymology.</emphasis>
The species is named in honour of Prof. Eugeny Y. Krysanov, a scientist at the Russian Academy of Science, who carried out cytological studies on
<taxonomicName id="8CE04D72FFBBFFB3E076FF3B8E1EFF6B" box="[865,1054,189,214]" class="Actinopterygii" family="Nothobranchiidae" genus="Nothobranchius" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="7994EAE3FFBBFFB3E076FF3B8E1EFF6B" box="[865,1054,189,214]" italics="true" pageId="12" pageNumber="49">Nothobranchius</emphasis>
</taxonomicName>
species, as well as the effects of radiation on their chromosomes in the Chernobyl area. The species name is pronounced “kree-sa-nofi”.
</paragraph>
</subSubSection>
</treatment>
</document>
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