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<document id="B6F83E1D3A5A0FC5673F5CC41CC30389" ID-DOI="10.5281/zenodo.12587419" ID-ISSN="2064-2474" ID-Zenodo-Dep="12587419" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719597512946" checkinUser="felipe" docAuthor="DellacaSa, M. &amp; DellacaSa, G." docDate="2002" docId="101D8782FFAD2A5DFE49FE76A8639E75" docLanguage="en" docName="ActaZoolAcadSciH.48.4.269-316.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 48 (4)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Ammoecius franzi DellacaSa &amp; DellacaSa 2002, comb. n." docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="EC24FFFAFFB72A47FFB4FFC1AB009B55" masterDocTitle="Systematic Redefinition Of Taxa Belonging To The Genera Ahermodontus Báguena, 1930 And Ammoecius Mulsant, 1842, With Description Of The New Genus Vladimirellus (Coleoptera: Aphodiidae)" masterLastPageNumber="316" masterPageNumber="269" pageNumber="295" updateTime="1719604724815" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="6876428F7847FD7272E006AF78C3C189" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="586BA930082844FCD4B034400BE06214">
<mods:title id="108F9386A11CCB1E88E926015E09570F">Systematic Redefinition Of Taxa Belonging To The Genera Ahermodontus Báguena, 1930 And Ammoecius Mulsant, 1842, With Description Of The New Genus Vladimirellus (Coleoptera: Aphodiidae)</mods:title>
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<mods:name id="C24113F7ABEB59C26A22B326B2808829" type="personal">
<mods:role id="2E5B64BDE33436857EF414615BD50666">
<mods:roleTerm id="9CCEE23138184B9122482D3CA7E92CBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F699D32AA95671FD5DF7A5A94556CBC">DellacaSa, M.</mods:namePart>
</mods:name>
<mods:name id="A56F778EB8F695203CC77C4273BD394B" type="personal">
<mods:role id="B7D341B5461BBDDFF1CC0F796CE3DD11">
<mods:roleTerm id="10473E2D5A8183A8AC24A41A448E8D09">Author</mods:roleTerm>
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<mods:namePart id="9F82C481F03AD0B06ACBB03A3397B34B">DellacaSa, G.</mods:namePart>
</mods:name>
<mods:typeOfResource id="01507AA30E53A5DE2D6709B9A2AC411C">text</mods:typeOfResource>
<mods:relatedItem id="CF5EC0F9D448822717A30D4F8599D2C0" type="host">
<mods:titleInfo id="BDDD7A9DBA8E299DBA5299FDB83792B2">
<mods:title id="6F5B506F394CAA6AA3C8F1342F37152A">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="777CEB153ACE3D4947477D48077CD81D">
<mods:date id="966C65013B4E064BC16859197EBDAF1F">2002</mods:date>
<mods:detail id="D1366316BC19C9E2FDCE34F6A88F25D9" type="pubDate">
<mods:number id="AD58D78BBC2E619448A9DE25431CCEEF">2003-02-28</mods:number>
</mods:detail>
<mods:detail id="803E701B3CDB1ACA64BD3FA6672C888C" type="volume">
<mods:number id="757B15AE74FC318ED5002C04EF41FAFA">48</mods:number>
</mods:detail>
<mods:detail id="A8F9B0FE39DC48462705A15859DBF8F2" type="issue">
<mods:number id="745E9CEB50D714B0B1B781D09B44F483">4</mods:number>
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<mods:start id="C53FAFCD7A4EEC91268A7A4DDC5123E5">269</mods:start>
<mods:end id="C22F504EE4A25441E554A8178F4BB7AC">316</mods:end>
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<mods:classification id="9174F22EC446870175769875AC54D350">journal article</mods:classification>
<mods:identifier id="0E68C5CF02E62BE22A07329E1B1DF6D1" type="DOI">10.5281/zenodo.12587419</mods:identifier>
<mods:identifier id="B0BDD98B57600B5E1A9F555DDA6DBCE7" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="FCA88943237ACA3FCF8A7A21FC49F2DD" type="Zenodo-Dep">12587419</mods:identifier>
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<treatment id="101D8782FFAD2A5DFE49FE76A8639E75" LSID="urn:lsid:plazi:treatment:101D8782FFAD2A5DFE49FE76A8639E75" httpUri="http://treatment.plazi.org/id/101D8782FFAD2A5DFE49FE76A8639E75" lastPageNumber="295" pageId="26" pageNumber="295">
<subSubSection id="D0AE651FFFAD2A5DFE49FE76AF369A84" box="[509,1078,439,465]" pageId="26" pageNumber="295" type="nomenclature">
<paragraph id="980B3694FFAD2A5DFE49FE76AF369A84" blockId="26.[509,1078,439,465]" box="[509,1078,439,465]" pageId="26" pageNumber="295">
<heading id="C34381F8FFAD2A5DFE49FE76AF369A84" box="[509,1078,439,465]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="295" reason="6">
<taxonomicName id="5FB44D17FFAD2A5DFE49FE76A8BF9A84" ID-CoL="CTN4" authority="(PETROVITZ, 1964)" authorityName="DellacaSa &amp; DellacaSa" authorityYear="2002" baseAuthorityName="PETROVITZ" baseAuthorityYear="1964" box="[509,959,439,465]" class="Insecta" family="Scarabaeidae" genus="Ammoecius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="295" phylum="Arthropoda" rank="species" species="franzi" status="comb. nov.">
<emphasis id="AAC0EA86FFAD2A5DFE49FE76A9D09A84" box="[509,720,439,465]" italics="true" pageId="26" pageNumber="295">Ammoecius franzi</emphasis>
(
<bibRefCitation id="FC254B65FFAD2A5DFD54FE76A8B79A84" author="PETROVITZ, R." box="[736,951,439,465]" pageId="26" pageNumber="295" pagination="171 - 207" refId="ref19722" refString="PETROVITZ, R. (1964) Neue afrikanische und australische Aphodiidae. Reichenbachia 4: 171 - 207." type="journal article" year="1964">
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, 1964
</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="AAC0EA86FFAD2A5DFC7FFE76AF369A84" bold="true" box="[971,1078,439,465]" pageId="26" pageNumber="295">
<taxonomicNameLabel id="B1F357FDFFAD2A5DFC7FFE76AF369A84" box="[971,1078,439,465]" pageId="26" pageNumber="295" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D0AE651FFFAD2A5DFD0CFE1BA87A9AA1" box="[696,890,474,500]" pageId="26" pageNumber="295" type="description">
<paragraph id="980B3694FFAD2A5DFD0CFE1BA87A9AA1" blockId="26.[696,890,474,500]" box="[696,890,474,500]" pageId="26" pageNumber="295">
<heading id="C34381F8FFAD2A5DFD0CFE1BA87A9AA1" box="[696,890,474,500]" centered="true" fontSize="11" level="3" pageId="26" pageNumber="295" reason="3">
(
<figureCitation id="008F2A11FFAD2A5DFD74FE1BA81C9AA1" box="[704,796,474,500]" captionStart="Figs 65–68" captionStartId="27.[317,360,1715,1737]" captionTargetBox="[411,1200,597,1670]" captionTargetId="figure-72@27.[370,1216,589,1686]" captionTargetPageId="27" captionText="Figs 65–68. Habitus of: 65 = Ammoecius numidicus MULSANT, 1851 (male, length 4.5 mm, Algeria: Mahouna), 66 = A. franzi (PETROVITZ, 1964) (male, length 5.5 mm, Morocco: Oukaïmeden), 67 = A. amplicollis (PEYERIMHOFF, 1939) (male, length 5.0 mm, Morocco: Oukaïmeden), 68 = A. meur-" figureDoi="http://doi.org/10.5281/zenodo.12587442" httpUri="https://zenodo.org/record/12587442/files/figure.png" pageId="26" pageNumber="295">Figs 66</figureCitation>
,
<figureCitation id="008F2A11FFAD2A5DFC9CFE1BA8729AA1" box="[808,882,474,500]" captionStart="Figs 85–95" captionStartId="30.[317,360,1740,1762]" captionTargetBox="[402,1198,428,1710]" captionTargetId="figure-18@30.[384,1202,422,1712]" captionTargetPageId="30" captionText="Figs 85–95. Epipharynx (85, 89, 93), apex of corypha (lateral view) (86, 90) and aedeagus (dorsal and lateral view) (87–88, 91–92, 94–95) of: 85–88 = Ammoecius franzi (PETROVITZ, 1964), 89–92 = A. numidicus MULSANT, 1851, 93–95 = A. incultus (PETROVITZ, 1961)" figureDoi="http://doi.org/10.5281/zenodo.12587450" httpUri="https://zenodo.org/record/12587450/files/figure.png" pageId="26" pageNumber="295">85–88</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D0AE651FFFAD2A5DFE89FDDFA8459905" pageId="26" pageNumber="295" type="reference_group">
<paragraph id="980B3694FFAD2A5DFE89FDDFA8459905" blockId="26.[317,1270,542,592]" pageId="26" pageNumber="295">
<treatmentCitationGroup id="B8A411BAFFAD2A5DFE89FDDFA8459905" pageId="26" pageNumber="295">
<treatmentCitation id="19151085FFAD2A5DFE89FDDFA84D9961" author="PETROVITZ, R." box="[317,845,542,564]" page="185" pageId="26" pageNumber="295" year="1964">
<taxonomicName id="5FB44D17FFAD2A5DFE89FDDFA84D9961" ID-CoL="FG2D" authority="PETROVITZ, 1964: 185" authorityName="PETROVITZ" authorityPageNumber="185" authorityYear="1964" box="[317,845,542,564]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="295" phylum="Arthropoda" rank="species" species="franzi" subGenus="Ammoecius">
<emphasis id="AAC0EA86FFAD2A5DFE89FDDFA9609961" box="[317,608,542,564]" italics="true" pageId="26" pageNumber="295">Aphodius (Ammoecius) franzi</emphasis>
<bibRefCitation id="FC254B65FFAD2A5DFDDCFDDFA84D9961" author="PETROVITZ, R." box="[616,845,542,564]" pageId="26" pageNumber="295" pagination="171 - 207" refId="ref19722" refString="PETROVITZ, R. (1964) Neue afrikanische und australische Aphodiidae. Reichenbachia 4: 171 - 207." type="journal article" year="1964">
<smallCapsWord id="9EEDA048FFAD2A5DFDDCFDDFA9D89967" baselines="558,558" box="[616,728,542,564]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Petrovitz" pageId="26" pageNumber="295">PETROVITZ</smallCapsWord>
, 1964: 185
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="19151085FFAD2A5DFCEDFDDFAF349961" author="PETROVITZ, R." box="[857,1076,542,564]" page="230" pageId="26" pageNumber="295" year="1968">
<bibRefCitation id="FC254B65FFAD2A5DFCEDFDDFAF349961" author="PETROVITZ, R." box="[857,1076,542,564]" pageId="26" pageNumber="295" pagination="227 - 235" refId="ref19742" refString="PETROVITZ, R. (1968) Scarabaeidae aus Marokko. Bull. Soc. Sci. Nat. Physiq. Maroc 47: 227 - 235." type="journal article" year="1968">
<smallCapsWord id="9EEDA048FFAD2A5DFCEDFDDFA8C89967" baselines="558,558" box="[857,968,542,564]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Petrovitz" pageId="26" pageNumber="295">PETROVITZ</smallCapsWord>
1968: 230
</bibRefCitation>
</treatmentCitation>
(male description);
<treatmentCitation id="19151085FFAD2A5DFECAFDFBA9339905" author="BARAUD, J." box="[382,563,570,592]" page="62" pageId="26" pageNumber="295" year="1971">
<bibRefCitation id="FC254B65FFAD2A5DFECAFDFBA9339905" author="BARAUD, J." box="[382,563,570,592]" pageId="26" pageNumber="295" pagination="63 - 71" refId="ref18511" refString="BARAUD, J. (1971) Revision des Aphodius palearctiques du sous-genre Ammoecius Muls. Bull. Soc. Ent. France 76: 63 - 71." type="journal article" year="1971">
<smallCapsWord id="9EEDA048FFAD2A5DFECAFDFBAAD5991A" baselines="586,587" box="[382,469,570,592]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Baraud" pageId="26" pageNumber="295">BARAUD</smallCapsWord>
1971: 62
</bibRefCitation>
</treatmentCitation>
;
<smallCapsWord id="9EEDA048FFAD2A5DFD89FDFBA9B6991A" baselines="586,587" box="[573,694,570,592]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Dellacasa" pageId="26" pageNumber="295">DELLACASA</smallCapsWord>
M. 1988: 131.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D0AE651FFFAD2A5DFECAFDB5A8639E75" pageId="26" pageNumber="295" type="materials_examined">
<paragraph id="980B3694FFAD2A5DFECAFDB5AF1299DF" blockId="26.[382,1042,628,679]" box="[382,1042,628,650]" pageId="26" pageNumber="295">
<materialsCitation id="28DC3CC9FFAD2A5DFECAFDB5AF1299DF" box="[382,1042,628,650]" country="Morocco" location="Circle de Jaffar" pageId="26" pageNumber="295" specimenCount="1" typeStatus="holotype">
<typeStatus id="470F8836FFAD2A5DFECAFDB5AAAF99DF" box="[382,431,628,650]" pageId="26" pageNumber="295">Type</typeStatus>
locality: “Hoher Atlas, Ostseite,
<location id="9D6B604FFFAD2A5DFD42FDB5A89399DF" LSID="urn:lsid:plazi:treatment:101D8782FFAD2A5DFE49FE76A8639E75:9D6B604FFFAD2A5DFD42FDB5A89399DF" box="[758,915,628,650]" country="Morocco" name="Circle de Jaffar" pageId="26" pageNumber="295">Circle de Jaffar</location>
”. [
<collectingCountry id="E0A37604FFAD2A5DFC1FFDB5AF0999DF" box="[939,1033,628,650]" name="Morocco" pageId="26" pageNumber="295">Morocco</collectingCountry>
.]
</materialsCitation>
</paragraph>
<paragraph id="980B3694FFAD2A5DFECAFD50A83899F2" blockId="26.[382,1042,628,679]" box="[382,824,657,679]" pageId="26" pageNumber="295">
<typeStatus id="470F8836FFAD2A5DFECAFD50AAAF99F2" box="[382,431,657,679]" pageId="26" pageNumber="295">Type</typeStatus>
depository:
<smallCapsWord id="9EEDA048FFAD2A5DFD9EFD50A96B99F0" baselines="673,673" box="[554,619,657,679]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Franz" pageId="26" pageNumber="295">FRANZ</smallCapsWord>
H. collection, Wien.
</paragraph>
<paragraph id="980B3694FFAD2A5DFECAFD0AA8779FB2" blockId="26.[317,1270,714,1255]" pageId="26" pageNumber="295">
<emphasis id="AAC0EA86FFAD2A5DFECAFD0AAAEF99B5" box="[382,495,715,736]" italics="true" pageId="26" pageNumber="295">Description</emphasis>
– Length 5.0–6.0 mm. Oblong, strongly convex, shiny, glabrous (
<figureCitation id="008F2A11FFAD2A5DFBD4FD0BAFA799B5" box="[1120,1191,714,736]" captionStart="Figs 65–68" captionStartId="27.[317,360,1715,1737]" captionTargetBox="[411,1200,597,1670]" captionTargetId="figure-72@27.[370,1216,589,1686]" captionTargetPageId="27" captionText="Figs 65–68. Habitus of: 65 = Ammoecius numidicus MULSANT, 1851 (male, length 4.5 mm, Algeria: Mahouna), 66 = A. franzi (PETROVITZ, 1964) (male, length 5.5 mm, Morocco: Oukaïmeden), 67 = A. amplicollis (PEYERIMHOFF, 1939) (male, length 5.0 mm, Morocco: Oukaïmeden), 68 = A. meur-" figureDoi="http://doi.org/10.5281/zenodo.12587442" httpUri="https://zenodo.org/record/12587442/files/figure.png" pageId="26" pageNumber="295">Fig. 66</figureCitation>
). Black; clypeal margin and legs brownish; antennal club dark testaceous. Head with cupuliform epistoma finely, distinctly and rather densely punctured; punctation somewhat denser laterally and almost irregular granulo-rugose in front of the rather short and moderately raised anterior transverse carina; clypeus subtriangularly sinuate at middle, subdenticulate at sides, latter distinctly bordered and regularlyarcuatetogenaethatarefeeblybutdistinctlyauriculate,obtuselyrounded,elongatelyciliateand protruding more than eyes; frontal suture distinct, finely impressed; frons finely and subregularly punctured, punctures somewhat sparser and coarser than those on epistoma; epipharynx:
<figureCitation id="008F2A11FFAD2A5DFB1DFC55AFF198FF" box="[1193,1265,916,938]" captionStart="Figs 85–95" captionStartId="30.[317,360,1740,1762]" captionTargetBox="[402,1198,428,1710]" captionTargetId="figure-18@30.[384,1202,422,1712]" captionTargetPageId="30" captionText="Figs 85–95. Epipharynx (85, 89, 93), apex of corypha (lateral view) (86, 90) and aedeagus (dorsal and lateral view) (87–88, 91–92, 94–95) of: 85–88 = Ammoecius franzi (PETROVITZ, 1964), 89–92 = A. numidicus MULSANT, 1851, 93–95 = A. incultus (PETROVITZ, 1961)" figureDoi="http://doi.org/10.5281/zenodo.12587450" httpUri="https://zenodo.org/record/12587450/files/figure.png" pageId="26" pageNumber="295">Fig. 85</figureCitation>
; apex of corypha:
<figureCitation id="008F2A11FFAD2A5DFE5BFC70A93A9892" box="[495,570,945,967]" captionStart="Figs 85–95" captionStartId="30.[317,360,1740,1762]" captionTargetBox="[402,1198,428,1710]" captionTargetId="figure-18@30.[384,1202,422,1712]" captionTargetPageId="30" captionText="Figs 85–95. Epipharynx (85, 89, 93), apex of corypha (lateral view) (86, 90) and aedeagus (dorsal and lateral view) (87–88, 91–92, 94–95) of: 85–88 = Ammoecius franzi (PETROVITZ, 1964), 89–92 = A. numidicus MULSANT, 1851, 93–95 = A. incultus (PETROVITZ, 1961)" figureDoi="http://doi.org/10.5281/zenodo.12587450" httpUri="https://zenodo.org/record/12587450/files/figure.png" pageId="26" pageNumber="295">Fig. 86</figureCitation>
. Pronotum moderately transverse, strongly convex, doubly punctured; larger punctures irregularly and sparsely scattered only on basal third of disc; much smaller ones regularly and densely sparse throughout, punctation slightly denser on sides; latter feebly arcuate, distinctly bordered; anterior margin finely bordered; hind angles obtusely rounded; base feebly bisinuate, distinctly bordered. Scutellum flat, finely and sparsely punctured near base. Elytra oval-elongate, strongly convex, not denticulate at shoulder; striae fine, faintly crenulate; interstices nearly flat, sparsely and almost imperceptibly punctured. Hind tibiae with superior apical spur distinctly longer than first tarsal segment; latter somewhat longer than following three combined. Male: inferior apical spur of middle tibiae shortened, apically truncate and inwardly abruptly bent; metasternal plate incavate; aedeagus:
<figureCitation id="008F2A11FFAD2A5DFD9AFB75A99C9F9F" box="[558,668,1204,1226]" captionStart="Figs 85–95" captionStartId="30.[317,360,1740,1762]" captionTargetBox="[402,1198,428,1710]" captionTargetId="figure-18@30.[384,1202,422,1712]" captionTargetPageId="30" captionText="Figs 85–95. Epipharynx (85, 89, 93), apex of corypha (lateral view) (86, 90) and aedeagus (dorsal and lateral view) (87–88, 91–92, 94–95) of: 85–88 = Ammoecius franzi (PETROVITZ, 1964), 89–92 = A. numidicus MULSANT, 1851, 93–95 = A. incultus (PETROVITZ, 1961)" figureDoi="http://doi.org/10.5281/zenodo.12587450" httpUri="https://zenodo.org/record/12587450/files/figure.png" pageId="26" pageNumber="295">Figs 87–88</figureCitation>
. Female: inferior apical spur of middle tibiae elongate and regularly acuminate toward apex; metasternal plate nearly flat.
</paragraph>
<paragraph id="980B3694FFAD2A5DFECAFACBA8639E75" blockId="26.[382,867,1290,1312]" box="[382,867,1290,1312]" pageId="26" pageNumber="295">
Distribution:
<collectingCountry id="E0A37604FFAD2A5DFDB4FACBA9579E75" box="[512,599,1290,1312]" name="Morocco" pageId="26" pageNumber="295">Morocco</collectingCountry>
(Haut Atlas, Moyen Atlas)
</paragraph>
</subSubSection>
</treatment>
</document>
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