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<document id="C57F24223294DB374B2ABDE4C4CF6C4B" ID-DOI="10.5281/zenodo.12587419" ID-ISSN="2064-2474" ID-Zenodo-Dep="12587419" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719597512946" checkinUser="felipe" docAuthor="DellacaSa, M. &amp; DellacaSa, G." docDate="2002" docId="101D8782FFA32A53FDF4FE76AFD59E2B" docLanguage="en" docName="ActaZoolAcadSciH.48.4.269-316.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 48 (4)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Ammoecius dentatus SCHMIDT 1908" docType="treatment" docVersion="2" lastPageNumber="289" masterDocId="EC24FFFAFFB72A47FFB4FFC1AB009B55" masterDocTitle="Systematic Redefinition Of Taxa Belonging To The Genera Ahermodontus Báguena, 1930 And Ammoecius Mulsant, 1842, With Description Of The New Genus Vladimirellus (Coleoptera: Aphodiidae)" masterLastPageNumber="316" masterPageNumber="269" pageNumber="289" updateTime="1719604724815" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="7FD35987CD0C51D19A2017B4FAFE2D68">Systematic Redefinition Of Taxa Belonging To The Genera Ahermodontus Báguena, 1930 And Ammoecius Mulsant, 1842, With Description Of The New Genus Vladimirellus (Coleoptera: Aphodiidae)</mods:title>
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<taxonomicName id="5FB44D17FFA32A53FDF4FE76A8F39A84" ID-CoL="CTMV" authority="SCHMIDT, 1908" authorityName="SCHMIDT" authorityYear="1908" box="[576,1011,439,465]" class="Insecta" family="Scarabaeidae" genus="Ammoecius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="AAC0EA86FFA32A53FDF4FE76A8319A84" box="[576,817,439,465]" italics="true" pageId="20" pageNumber="289">Ammoecius dentatus</emphasis>
<bibRefCitation id="FC254B65FFA32A53FC8DFE76A8F39A84" author="SCHMIDT, A." box="[825,1011,439,465]" pageId="20" pageNumber="289" refId="ref19947" refString="SCHMIDT, A. (1908) Ammoecius dentatus n. sp. Dtsch ent. Zeitschrift: 504." type="book" year="1908">
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, 1908
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<paragraph id="980B3694FFA32A53FD0CFE1BA87A9AA1" blockId="20.[696,890,474,500]" box="[696,890,474,500]" pageId="20" pageNumber="289">
<heading id="C34381F8FFA32A53FD0CFE1BA87A9AA1" box="[696,890,474,500]" centered="true" fontSize="11" level="3" pageId="20" pageNumber="289" reason="3">
(
<figureCitation id="008F2A11FFA32A53FD74FE1BA81C9AA1" box="[704,796,474,500]" captionStart="Figs 4952" captionStartId="21.[317,360,1692,1714]" captionTargetBox="[367,1215,440,1647]" captionTargetId="figure-16@21.[362,1224,422,1663]" captionTargetPageId="21" captionText="Figs 4952. Habitus of: 49 = Ammoecius muchei (PETROVITZ, 1962) (male, length 6.0 mm, Turkey: Vil. Kastamonu, Kastamonu env.), 50 = A. satanas (CARPANETO, 1976) (male, length 7.0 mm, Turkey: Vil. Antalya, 40 km S of Akseki, 3 km S Fersin), 51 = A. felscheanus REITTER, 1904 (male, length 3.5 mm, Algeria: Oran prov., Oued-Imbert), 52 = A. dentatus SCHMIDT, 1908 (male, length" figureDoi="http://doi.org/10.5281/zenodo.12587438" httpUri="https://zenodo.org/record/12587438/files/figure.png" pageId="20" pageNumber="289">Figs 52</figureCitation>
,
<figureCitation id="008F2A11FFA32A53FC9CFE1BA8729AA1" box="[808,882,474,500]" captionStart="Figs 6980" captionStartId="28.[317,360,1745,1767]" captionTargetBox="[393,1195,437,1707]" captionTargetId="figure-18@28.[388,1198,425,1717]" captionTargetPageId="28" captionText="Figs 6980. Epipharynx (69, 73, 77), apex of corypha (lateral view) (70, 74, 78) and aedeagus (dorsal and lateral view) (7172, 7576, 7980) of: 6972 = Ammoecius amplicollis (PEYERIMHOFF, 1939)," figureDoi="http://doi.org/10.5281/zenodo.12587446" httpUri="https://zenodo.org/record/12587446/files/figure.png" pageId="20" pageNumber="289">7780</figureCitation>
)
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<taxonomicName id="5FB44D17FFA32A53FE89FDDFA9D59961" ID-CoL="CTMV" authority="SCHMIDT, 1908: 504" authorityName="SCHMIDT" authorityPageNumber="504" authorityYear="1908" box="[317,725,542,564]" class="Insecta" family="Scarabaeidae" genus="Ammoecius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="AAC0EA86FFA32A53FE89FDDFA9039961" box="[317,515,542,564]" italics="true" pageId="20" pageNumber="289">Ammoecius dentatus</emphasis>
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, 1908: 504
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.
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</paragraph>
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<treatmentCitationGroup id="B8A411BAFFA32A53FE89FDFBA8459938" pageId="20" pageNumber="289">
<taxonomicName id="5FB44D17FFA32A53FE89FDFBA9749905" authorityName="SCHMIDT" authorityYear="1908" box="[317,628,570,592]" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="289" phylum="Arthropoda" rank="species" species="dentatus" subGenus="Ammoecius">
<emphasis id="AAC0EA86FFA32A53FE89FDFBA9749905" box="[317,628,570,592]" italics="true" pageId="20" pageNumber="289">Aphodius (Ammoecius) dentatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="19151085FFA32A53FD36FDFBA8499905" author="SCHMIDT, A." box="[642,841,570,592]" page="126" pageId="20" pageNumber="289" year="1913">
<bibRefCitation id="FC254B65FFA32A53FD36FDFBA8499905" author="SCHMIDT, A." box="[642,841,570,592]" pageId="20" pageNumber="289" pagination="117 - 178" refId="ref19967" refString="SCHMIDT, A. (1913) Erster Versuch einer Einteilung de exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv Naturges., Abt. A 79: 117 - 178." type="journal article" year="1913">
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1913: 126
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;
<treatmentCitation id="19151085FFA32A53FCE7FDFBAF0E9905" author="SCHMIDT, A." box="[851,1038,570,592]" page="70" pageId="20" pageNumber="289" year="1922">
<bibRefCitation id="FC254B65FFA32A53FCE7FDFBAF0E9905" author="SCHMIDT, A." box="[851,1038,570,592]" pageId="20" pageNumber="289" refId="ref20002" refString="SCHMIDT, A. (1922) Coleoptera Aphodiinae. (Das Tierreich). 45. W. de Gruyter, Berlin &amp; Leipzig, 614 pp." type="book" year="1922">
<smallCapsWord id="9EEDA048FFA32A53FCE7FDFBA8B0991A" baselines="586,587" box="[851,944,570,592]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schmidt" pageId="20" pageNumber="289">SCHMIDT</smallCapsWord>
1922: 70
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;
<treatmentCitation id="19151085FFA32A53FBACFDFBAFF29905" author="BALTHASAR, V." box="[1048,1266,570,592]" page="78" pageId="20" pageNumber="289" year="1964">
<bibRefCitation id="FC254B65FFA32A53FBACFDFBAFF29905" author="BALTHASAR, V." box="[1048,1266,570,592]" pageId="20" pageNumber="289" refId="ref18448" refString="BALTHASAR, V. (1964) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3, Aphodiidae. Verlag der Tschechoslowakischen Akademie der Wissenschaften, Praha, 652 pp." type="book" year="1964">
<smallCapsWord id="9EEDA048FFA32A53FBACFDFBAF93991A" baselines="586,587" box="[1048,1171,570,592]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Balthasar" pageId="20" pageNumber="289">BALTHASAR</smallCapsWord>
1964: 78
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="19151085FFA32A53FECAFD96A9339938" author="BARAUD, J." box="[382,563,599,621]" page="68" pageId="20" pageNumber="289" year="1971">
<bibRefCitation id="FC254B65FFA32A53FECAFD96A9339938" author="BARAUD, J." box="[382,563,599,621]" pageId="20" pageNumber="289" pagination="63 - 71" refId="ref18511" refString="BARAUD, J. (1971) Revision des Aphodius palearctiques du sous-genre Ammoecius Muls. Bull. Soc. Ent. France 76: 63 - 71." type="journal article" year="1971">
<smallCapsWord id="9EEDA048FFA32A53FECAFD96AAD59939" baselines="615,616" box="[382,469,599,621]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Baraud" pageId="20" pageNumber="289">BARAUD</smallCapsWord>
1971: 68
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;
<smallCapsWord id="9EEDA048FFA32A53FD89FD96A9B69939" baselines="615,616" box="[573,694,599,621]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Dellacasa" pageId="20" pageNumber="289">DELLACASA</smallCapsWord>
M. 1988: 119.
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<typeStatus id="470F8836FFA32A53FECAFD50AAAF99F2" box="[382,431,657,679]" pageId="20" pageNumber="289">Type</typeStatus>
locality: “
<collectingCountry id="E0A37604FFA32A53FDA2FD50A95E99F2" box="[534,606,657,679]" name="Morocco" pageId="20" pageNumber="289">Tanger</collectingCountry>
, Marokko”. [
<collectingCountry id="E0A37604FFA32A53FD6BFD50A83D99F2" box="[735,829,657,679]" name="Morocco" pageId="20" pageNumber="289">Morocco</collectingCountry>
.]
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<materialsCitation id="28DC3CC9FFA32A53FECAFD6FA8AE9991" box="[382,942,686,708]" location="Naturhistoriska Riksmuseet" pageId="20" pageNumber="289" specimenCount="1" stateProvince="Stockholm" typeStatus="holotype">
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depository:
<location id="9D6B604FFFA32A53FD9EFD6FA8399991" LSID="urn:lsid:plazi:treatment:101D8782FFA32A53FDF4FE76AFD59E2B:9D6B604FFFA32A53FD9EFD6FA8399991" box="[554,825,686,708]" name="Naturhistoriska Riksmuseet" pageId="20" pageNumber="289" stateProvince="Stockholm">Naturhistoriska Riksmuseet</location>
,
<collectingRegion id="5A70F876FFA32A53FCF7FD6FA8AE9991" box="[835,942,686,708]" country="Sweden" name="Stockholm" pageId="20" pageNumber="289">Stockholm</collectingRegion>
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.
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<emphasis id="AAC0EA86FFA32A53FECAFD29AAEF99A8" box="[382,495,744,765]" italics="true" pageId="20" pageNumber="289">Description</emphasis>
Length 4.05.0 mm. Short, oval, strongly convex, moderately shiny, glabrous (
<figureCitation id="008F2A11FFA32A53FEF0FCC5AA85984F" box="[324,389,772,794]" captionStart="Figs 4952" captionStartId="21.[317,360,1692,1714]" captionTargetBox="[367,1215,440,1647]" captionTargetId="figure-16@21.[362,1224,422,1663]" captionTargetPageId="21" captionText="Figs 4952. Habitus of: 49 = Ammoecius muchei (PETROVITZ, 1962) (male, length 6.0 mm, Turkey: Vil. Kastamonu, Kastamonu env.), 50 = A. satanas (CARPANETO, 1976) (male, length 7.0 mm, Turkey: Vil. Antalya, 40 km S of Akseki, 3 km S Fersin), 51 = A. felscheanus REITTER, 1904 (male, length 3.5 mm, Algeria: Oran prov., Oued-Imbert), 52 = A. dentatus SCHMIDT, 1908 (male, length" figureDoi="http://doi.org/10.5281/zenodo.12587438" httpUri="https://zenodo.org/record/12587438/files/figure.png" pageId="20" pageNumber="289">Fig 52</figureCitation>
). Black, clypeal margin widely brown-reddish; legs brown-reddish with tarsi paler; antennal club testaceous. Head with cupuliform epistoma distinctly, finely, subregularly punctured on disc; laterally mixed with few large punctures; punctation fine and regular in front of short but strongly raised anterior transverse carina; clypeus widely sinuate at middle, sharply denticulate at sides, very thinly bordered, border feebly upturned; genae angulose, elongately ciliate, protruding more than eyes; frontal suture finely impressed; frons finely and subregularly punctured; epipharynx:
<figureCitation id="008F2A11FFA32A53FB1FFC55AFF298FF" box="[1195,1266,916,938]" captionStart="Figs 6980" captionStartId="28.[317,360,1745,1767]" captionTargetBox="[393,1195,437,1707]" captionTargetId="figure-18@28.[388,1198,425,1717]" captionTargetPageId="28" captionText="Figs 6980. Epipharynx (69, 73, 77), apex of corypha (lateral view) (70, 74, 78) and aedeagus (dorsal and lateral view) (7172, 7576, 7980) of: 6972 = Ammoecius amplicollis (PEYERIMHOFF, 1939)," figureDoi="http://doi.org/10.5281/zenodo.12587446" httpUri="https://zenodo.org/record/12587446/files/figure.png" pageId="20" pageNumber="289">Fig. 77</figureCitation>
; apex of corypha:
<figureCitation id="008F2A11FFA32A53FE55FC70A9279892" box="[481,551,945,967]" captionStart="Figs 6980" captionStartId="28.[317,360,1745,1767]" captionTargetBox="[393,1195,437,1707]" captionTargetId="figure-18@28.[388,1198,425,1717]" captionTargetPageId="28" captionText="Figs 6980. Epipharynx (69, 73, 77), apex of corypha (lateral view) (70, 74, 78) and aedeagus (dorsal and lateral view) (7172, 7576, 7980) of: 6972 = Ammoecius amplicollis (PEYERIMHOFF, 1939)," figureDoi="http://doi.org/10.5281/zenodo.12587446" httpUri="https://zenodo.org/record/12587446/files/figure.png" pageId="20" pageNumber="289">Fig. 78</figureCitation>
. Pronotum moderately transverse, strongly convex, doubly punctured; large punctures sparser on disc and absent on anterior quarter, mixed with fine punctures regularly scattered throughout; anterior margin finely bordered; sides feebly rounded, distinctly bordered; hind angles obtusely rounded; base not bisinuate, rather thickly bordered. Scutellum regularly triangular, irregularly punctured on basal half. Elytra strongly convex, feebly broadened backward; striae almost fine, superficially punctured, feebly crenulate; interstices very slightly convex with extremely fine sparse punctures. Hind tibiae with superior apical spur longer than first tarsal segment; latter shorter than following three combined. Male: pronotum with large punctures relatively less densely arranged;inferiorapicalspurofmiddletibiaeveryshort,obtuselyroundedapically;metasternalplate distinctly concave; aedeagus:
<figureCitation id="008F2A11FFA32A53FDD4FB75A9D19F9F" box="[608,721,1204,1226]" captionStart="Figs 6980" captionStartId="28.[317,360,1745,1767]" captionTargetBox="[393,1195,437,1707]" captionTargetId="figure-18@28.[388,1198,425,1717]" captionTargetPageId="28" captionText="Figs 6980. Epipharynx (69, 73, 77), apex of corypha (lateral view) (70, 74, 78) and aedeagus (dorsal and lateral view) (7172, 7576, 7980) of: 6972 = Ammoecius amplicollis (PEYERIMHOFF, 1939)," figureDoi="http://doi.org/10.5281/zenodo.12587446" httpUri="https://zenodo.org/record/12587446/files/figure.png" pageId="20" pageNumber="289">Figs 7980</figureCitation>
. Female: pronotum with large punctures relatively more densely arranged; inferior apical spur of hind tibiae elongate and regularly acuminate apically; metasternal plate flat.
</paragraph>
<paragraph id="980B3694FFA32A53FECAFAE6AF7C9E68" blockId="20.[382,1148,1319,1341]" box="[382,1148,1319,1341]" pageId="20" pageNumber="289">
Distribution:
<collectingCountry id="E0A37604FFA32A53FDB4FAE6A9519E68" box="[512,593,1319,1341]" name="Portugal" pageId="20" pageNumber="289">Portugal</collectingCountry>
(Algarve),
<collectingCountry id="E0A37604FFA32A53FD70FAE6A9FB9E68" box="[708,763,1319,1341]" name="Spain" pageId="20" pageNumber="289">Spain</collectingCountry>
(
<collectingRegion id="5A70F876FFA32A53FCBDFAE6A86F9E68" box="[777,879,1319,1341]" country="Spain" name="Andalucia" pageId="20" pageNumber="289">Andalusia</collectingRegion>
),
<collectingCountry id="E0A37604FFA32A53FC35FAE6A8D89E68" box="[897,984,1319,1341]" name="Morocco" pageId="20" pageNumber="289">Morocco</collectingCountry>
(Atlantic coast).
</paragraph>
<paragraph id="980B3694FFA32A53FECAFAA5AFD59E2B" blockId="20.[382,1237,1380,1406]" box="[382,1237,1380,1406]" pageId="20" pageNumber="289">
<emphasis id="AAC0EA86FFA32A53FECAFAA5AAE59E2B" box="[382,485,1380,1406]" italics="true" pageId="20" pageNumber="289">Remarks</emphasis>
Coastal sands seem to be the exclusive habitat of this species.
</paragraph>
</subSubSection>
</treatment>
</document>