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<mods:title id="D2D5A4A7607B04A13AF036BA6412A9D3">Taxonomy of freshwater gobies of the genus Rhinogobius (Oxudercidae, Gobiiformes) from central Vietnam, with descriptions of two new species</mods:title>
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<mods:namePart id="3480A96D737E80E737C2F43C759037F3">Maeda, Ken</mods:namePart>
<mods:affiliation id="3121AE7149D98F88F79DEBD46465228F">Marine Eco-Evo-Devo Unit, Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 -</mods:affiliation>
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<mods:namePart id="E29B63EC475EC8C1E3E404AA005D8C0B">Kobayashi, Hirozumi</mods:namePart>
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<mods:affiliation id="D720FF8890537BD027E188807B8A85CE">Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903 - 0213, Japan. &amp; Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:namePart id="C6EE3C4D248906BC2FBB7272E52D1F74">Iida, Midori</mods:namePart>
<mods:affiliation id="FEC4C80FC9C9030DB6B31804B4890E0A">Marine Biological Station, Sado Island Center for Ecological Sustainability, Niigata University, 87 Tassha, Sado, Niigata 952 - 2135,</mods:affiliation>
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<mods:namePart id="915569FDC9C12FB9EB81F018CCB9352B">Tran, Hau Duc</mods:namePart>
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<mods:affiliation id="EF7757648A93BB0F65D0E6B494E75FCB">Institute of Natural Science, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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Key to the species of
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<emphasis id="CCEFEF3BFFF67D42A1C2FC94FDD8FCDC" bold="true" box="[413,568,798,825]" italics="true" pageId="30" pageNumber="537">Rhinogobius</emphasis>
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in
<collectingRegion id="3C5FFDCBFFF67D42A200FC94FD2DFCDD" box="[607,717,798,824]" country="Vietnam" name="Da Nang" pageId="30" pageNumber="537">Da Nang</collectingRegion>
City and Thua Thien Hue Province,
<collectingCountry id="868C73B9FFF67D42A4C6FC94FAE3FCDD" box="[1177,1283,798,824]" name="Vietnam" pageId="30" pageNumber="537">Vietnam</collectingCountry>
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1 Predorsal scales extending anteriorly to near posterior margin of eyes; cheek with several short transverse rows of cutaneous sensory papillae, and reddish-brown dots and lines forming diagonal stripes running upward from anterior to posterior...................................................................................................
<taxonomicName id="399B48AAFFF67D42A51CFC14FA7DFC56" ID-CoL="4S92Y" authorityName="Gill" authorityYear="1859" box="[1347,1437,926,947]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="similis">
<emphasis id="CCEFEF3BFFF67D42A51CFC14FA7DFC56" box="[1347,1437,926,947]" italics="true" pageId="30" pageNumber="537">R. similis</emphasis>
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<paragraph id="FE243329FFF67D42A0C8FC36FA7CFBEE" blockId="30.[151,1437,867,1563]" pageId="30" pageNumber="537">
<keyLead id="456A901CFFF67D42A0C8FC36FA7CFBEE" pageId="30" pageNumber="537">- Predorsal scales not reaching posterior margin of preopercle or ending shortly anterior to it (not reaching close to eyes); cheek without transverse row of cutaneous sensory papillae, and without diagonal stripes, or with diagonal stripes directing downward from anterior to posterior............................................................................... 2</keyLead>
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2 When mouth closed, anterior tip of lower jaw significantly protruding beyond upper jaw in larger male (&gt;
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SL); pectoral fin usually with 19 rays; orange (male), reddish-brown, or grey (female) diagonally downward stripes on cheek.....
<taxonomicName id="399B48AAFFF67D42A50AFBBBFA7CFBA3" authorityName="Maeda &amp; Kobayashi &amp; Iida &amp; Tran" authorityYear="2024" box="[1365,1436,1073,1094]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="rong">
<emphasis id="CCEFEF3BFFF67D42A50AFBBBFA7CFBA3" box="[1365,1436,1073,1094]" italics="true" pageId="30" pageNumber="537">R. rong</emphasis>
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- Positions of anterior tips of lower and upper jaws approximately same, or upper jaw protrudes more anteriorly; pectoral fin with 18 or fewer rays (sometimes
<quantity id="39639ECCFFF67D42A1A9FBE1FDC7FB64" box="[502,551,1131,1153]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="30" pageNumber="537" unit="in" value="19.0">19 in</quantity>
<taxonomicName id="399B48AAFFF67D42A273FBE6FD6DFB65" ID-CoL="4S8ZL" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[556,653,1131,1153]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="leavelli">
<emphasis id="CCEFEF3BFFF67D42A273FBE6FD6DFB65" box="[556,653,1131,1153]" italics="true" pageId="30" pageNumber="537">R. leavelli</emphasis>
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); cheek with three parallel, diagonally downward, reddish-brown or dusky-brown stripes (male of
<taxonomicName id="399B48AAFFF67D42A122FB03FE28FB7B" authorityName="Maeda &amp; Kobayashi &amp; Iida &amp; Tran" authorityYear="2024" box="[381,456,1161,1182]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="nami">
<emphasis id="CCEFEF3BFFF67D42A122FB03FE28FB7B" box="[381,456,1161,1182]" italics="true" pageId="30" pageNumber="537">R. nami</emphasis>
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), or without diagonal stripes (other species)............................................. 3
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3 Dark-brown vertical band at caudal-fin base often combined with larger dark-brown lateral blotch on caudal peduncle; opercular membrane, interopercle, and lower part of subopercle of male grey with many orange stripes running along margin of preopercle and opercle; first dorsal fin of male with 13 distinct black spots on proximal parts (not in middle) of membranes between anterior 24 spines; pectoral fin with 1619 rays..............................................
<taxonomicName id="399B48AAFFF67D42A564FB74FA7DFAF6" ID-CoL="4S8ZL" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[1339,1437,1278,1299]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="leavelli">
<emphasis id="CCEFEF3BFFF67D42A564FB74FA7DFAF6" box="[1339,1437,1278,1299]" italics="true" pageId="30" pageNumber="537">R. leavelli</emphasis>
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<keyLead id="456A901CFFF67D42A0C8FA96FA7CFA6D" pageId="30" pageNumber="537">- Black, vertical rectangular or elliptical spot at mid-base of caudal fin; opercular membrane, interopercle, and lower part of subopercle of male light bluish-grey with many red or orange spots; first dorsal fin of male with series of black or brown markings on middle parts of membranes between first to fifth spines (anterior one or two markings usually clear); pectoral fin usually with 1416 rays................................................................................ 4</keyLead>
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4 Five, longitudinal, reddish-brown or dusky-brown stripes (dotted lines) running on lateral and dorsal sides from trunk to caudal peduncle; three parallel, diagonally downward, reddish-brown or dusky-brown stripes on ventral part of cheek in male; number of vertebrae usually 26............................................................................
<taxonomicName id="399B48AAFFF67D42A50DFA46FA7CFA04" authorityName="Maeda &amp; Kobayashi &amp; Iida &amp; Tran" authorityYear="2024" box="[1362,1436,1484,1505]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="nami">
<emphasis id="CCEFEF3BFFF67D42A50DFA46FA7CFA04" box="[1362,1436,1484,1505]" italics="true" pageId="30" pageNumber="537">R. nami</emphasis>
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<paragraph id="FE243329FFF67D42A0C8FA63FA7CF9FE" blockId="30.[151,1437,867,1563]" pageId="30" pageNumber="537">
<keyLead id="456A901CFFF67D42A0C8FA63FA7CF9FE" pageId="30" pageNumber="537">
- Lateral and dorsal sides of body without longitudinal stripes; cheek without diagonal stripes; number of vertebrae usually 27............................................................................................
<taxonomicName id="399B48AAFFF67D42A542F98CFA77F9FE" ID-CoL="4S934" authorityName="Chen, Kottelat &amp; Miller" authorityYear="1999" box="[1309,1431,1542,1563]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="537" phylum="Chordata" rank="species" species="taenigena">
<emphasis id="CCEFEF3BFFF67D42A542F98CFA77F9FE" box="[1309,1431,1542,1563]" italics="true" pageId="30" pageNumber="537">R. taenigena</emphasis>
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.
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