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<document ID-DOI="10.25221/fee.446.2" ID-ISSN="2713-2196" ID-Zenodo-Dep="7166738" approvalRequired="25" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="23" approvalRequired_for_treatments="1" checkinTime="1665256185880" checkinUser="felipe" docAuthor="Wang, F. L." docDate="2022" docId="2C16504AFFA8AA75FF1CFE84D117F99D" docLanguage="en" docName="FarEastEntomol.446.7-12.pdf" docOrigin="Far Eastern Entomologist 446" docSource="http://dx.doi.org/10.25221/fee.446.2" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Tocama huangjianbini Wang 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="D02F2832FFA9AA76FFFFFFBBD54FFFB8" masterDocTitle="A new species of the genus Tocama Reitter, 1902 (Coleoptera: Scarabaeidae, Melolonthinae) from China" masterLastPageNumber="12" masterPageNumber="7" pageNumber="8" updateTime="1665270736685" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of the genus Tocama Reitter, 1902 (Coleoptera: Scarabaeidae, Melolonthinae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, F. L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>446</mods:number>
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<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>12</mods:end>
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<mods:url>http://dx.doi.org/10.25221/fee.446.2</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.446.2</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7166738</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:2C16504AFFA8AA75FF1CFE84D117F99D" httpUri="http://treatment.plazi.org/id/2C16504AFFA8AA75FF1CFE84D117F99D" lastPageId="3" lastPageNumber="10" pageId="1" pageNumber="8">
<subSubSection pageId="1" pageNumber="8" type="nomenclature">
<paragraph blockId="1.[227,583,318,340]" box="[227,583,318,340]" pageId="1" pageNumber="8">
<heading box="[227,583,318,340]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="8" reason="2">
<emphasis bold="true" box="[227,583,318,340]" pageId="1" pageNumber="8">
<taxonomicName authority="Wang" authorityName="Wang" authorityYear="2022" box="[227,518,318,340]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="huangjianbini" status="sp. n.">
<emphasis bold="true" box="[227,454,318,340]" italics="true" pageId="1" pageNumber="8">Tocama huangjianbini</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel box="[529,583,319,340]" pageId="1" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[227,1082,351,373]" box="[227,1082,351,373]" pageId="1" pageNumber="8">
http://zoobank.org/NomenclaturalActs/
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</paragraph>
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<subSubSection box="[227,310,384,406]" pageId="1" pageNumber="8" type="description">
<paragraph blockId="1.[227,310,384,406]" box="[227,310,384,406]" pageId="1" pageNumber="8">
<figureCitation box="[227,310,384,406]" captionStart-0="Figs 14" captionStart-1="Figs 511" captionStartId-0="2.[265,306,1588,1610]" captionStartId-1="3.[265,306,1193,1215]" captionTargetBox-0="[266,1089,272,1534]" captionTargetBox-1="[266,1108,300,1127]" captionTargetId-0="figure-0@2.[227,1110,257,1551]" captionTargetId-1="figure-0@3.[227,1123,257,1159]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs 14. Tocama huangjianbini sp. n., habitus. 1, 2 holotype, male; 3, 4 paratype" captionText-1="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi-0="http://doi.org/10.5281/zenodo.7166740" figureDoi-1="http://doi.org/10.5281/zenodo.7166742" httpUri-0="https://zenodo.org/record/7166740/files/figure.png" httpUri-1="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Figs 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="8" type="materials_examined">
<paragraph blockId="1.[227,1128,439,1592]" box="[265,1127,439,461]" pageId="1" pageNumber="8">
<materialsCitation box="[265,1127,439,461]" collectionCode="TYPE, MATERIAL" country="China" location="Pintan Island" municipality="Fuzhou City" pageId="1" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="holotype">
<collectionCode box="[265,459,439,460]" pageId="1" pageNumber="8">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[469,558,439,461]" pageId="1" pageNumber="8">Holotype</typeStatus>
<specimenCount box="[584,604,439,460]" pageId="1" pageNumber="8" type="male"></specimenCount>
,
<emphasis bold="true" box="[616,679,439,461]" pageId="1" pageNumber="8">
<collectingCountry box="[616,679,439,461]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
</emphasis>
:
<collectingRegion box="[692,845,439,461]" country="China" name="Fujian" pageId="1" pageNumber="8">Fujian Province</collectingRegion>
/
<collectingMunicipality box="[866,989,439,461]" pageId="1" pageNumber="8">Fuzhou City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2C16504AFFA8AA75FF1CFE84D117F99D:A160B787FFA8AA77FC17FE0CD128FE75" box="[1000,1127,439,461]" country="China" municipality="Fuzhou City" name="Pintan Island" pageId="1" pageNumber="8" stateProvince="Fujian">Pintan Island</location>
</materialsCitation>
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
/ Longfengtou Coast / alt. 0m,
<date box="[541,642,467,489]" pageId="1" pageNumber="8" value="2020-04-06">6.IV 2020</date>
/ Jian-Bin Huang leg. //
<materialsCitation collectionCode="MYNU" pageId="1" pageNumber="8" specimenCount="1" typeStatus="holotype">
<typeStatus box="[898,1025,467,488]" pageId="1" pageNumber="8">HOLOTYPE</typeStatus>
/
<taxonomicName authorityName="Wang" authorityYear="2022" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="huangjianbini">
<emphasis italics="true" pageId="1" pageNumber="8">Tocama huangjianbini</emphasis>
</taxonomicName>
/ FL-Wang det. 2021 (
<collectionCode box="[585,664,494,515]" pageId="1" pageNumber="8">MYNU</collectionCode>
)
</materialsCitation>
.
<materialsCitation box="[680,1113,494,516]" country="China" location="Fujian Province" pageId="1" pageNumber="8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<typeStatus box="[680,773,495,516]" pageId="1" pageNumber="8">Paratypes</typeStatus>
<specimenCount box="[799,829,494,515]" pageId="1" pageNumber="8" type="male">1♂</specimenCount>
,
<specimenCount box="[841,871,494,515]" pageId="1" pageNumber="8" type="female">1♀</specimenCount>
,
<emphasis bold="true" box="[884,947,494,516]" pageId="1" pageNumber="8">
<collectingCountry box="[884,947,494,516]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
</emphasis>
:
<collectingRegion box="[960,1113,494,516]" country="China" name="Fujian" pageId="1" pageNumber="8">Fujian Province</collectingRegion>
</materialsCitation>
/
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,1126,522,544]" pageId="1" pageNumber="8">
Fuzhou City, Pintan Island / Longfengtou Coast / alt. 0 m,
<date box="[796,894,522,544]" pageId="1" pageNumber="8" value="2020-04-06">6.IV 2020</date>
/ Jian-Bin Huang leg. //
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,1127,549,571]" pageId="1" pageNumber="8">
<materialsCitation box="[227,895,549,571]" collectionCode="MYNU" pageId="1" pageNumber="8" specimenCount="1" typeStatus="paratype">
<typeStatus box="[227,351,550,571]" pageId="1" pageNumber="8">PARATYPE</typeStatus>
/
<taxonomicName authorityName="Wang" authorityYear="2022" box="[370,589,549,571]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="huangjianbini">
<emphasis box="[370,589,549,571]" italics="true" pageId="1" pageNumber="8">Tocama huangjianbini</emphasis>
</taxonomicName>
/ FL-Wang det. 2021 (
<collectionCode box="[810,889,550,571]" pageId="1" pageNumber="8">MYNU</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[905,1127,549,571]" country="China" location="Fujian" pageId="1" pageNumber="8" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount box="[905,937,550,571]" pageId="1" pageNumber="8" type="male">2♂</specimenCount>
,
<specimenCount box="[947,977,550,571]" pageId="1" pageNumber="8" type="female">1♀</specimenCount>
,
<emphasis bold="true" box="[989,1052,549,571]" pageId="1" pageNumber="8">
<collectingCountry box="[989,1052,549,571]" name="China" pageId="1" pageNumber="8">China</collectingCountry>
</emphasis>
:
<collectingRegion box="[1065,1127,549,571]" country="China" name="Fujian" pageId="1" pageNumber="8">
<location LSID="urn:lsid:plazi:treatment:2C16504AFFA8AA75FF1CFE84D117F99D:A160B787FFA8AA77FBD6FD9ED128FD83" box="[1065,1127,549,571]" country="China" name="Fujian" pageId="1" pageNumber="8">Fujian</location>
</collectingRegion>
</materialsCitation>
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,1126,577,599]" pageId="1" pageNumber="8">
Province / Fuzhou City, Pintan Island / Longfengtou Coast / alt. 0 m,
<date box="[923,1022,577,599]" pageId="1" pageNumber="8" value="2020-04-06">6.IV 2020</date>
/ Jian-Bin
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,1021,604,626]" pageId="1" pageNumber="8">
<materialsCitation box="[227,1018,604,626]" collectionCode="CFLW" collectorName="Huang" pageId="1" pageNumber="8" specimenCount="1" typeStatus="paratype">
<collectorName box="[227,291,605,626]" pageId="1" pageNumber="8">Huang</collectorName>
leg. //
<typeStatus box="[358,482,605,626]" pageId="1" pageNumber="8">PARATYPE</typeStatus>
/
<taxonomicName authorityName="Wang" authorityYear="2022" box="[500,719,604,626]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="huangjianbini">
<emphasis box="[500,719,604,626]" italics="true" pageId="1" pageNumber="8">Tocama huangjianbini</emphasis>
</taxonomicName>
/ FL-Wang det. 2021 (
<collectionCode box="[938,1010,605,626]" pageId="1" pageNumber="8">CFLW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="10" pageId="1" pageNumber="8" type="description">
<paragraph blockId="1.[227,1128,439,1592]" box="[265,1127,632,654]" pageId="1" pageNumber="8">
DESCRIPTION. MALE (
<typeStatus box="[519,609,632,654]" pageId="1" pageNumber="8">holotype</typeStatus>
). Habitus (
<figureCitation box="[717,808,632,654]" captionStart="Figs 14" captionStartId="2.[265,306,1588,1610]" captionTargetBox="[266,1089,272,1534]" captionTargetId="figure-0@2.[227,1110,257,1551]" captionTargetPageId="2" captionText="Figs 14. Tocama huangjianbini sp. n., habitus. 1, 2 holotype, male; 3, 4 paratype" figureDoi="http://doi.org/10.5281/zenodo.7166740" httpUri="https://zenodo.org/record/7166740/files/figure.png" pageId="1" pageNumber="8">Figs 1, 2</figureCitation>
). Length: 16.2 mm, width: 10.7
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,793,660,682]" pageId="1" pageNumber="8">
mm. Body elongate ovoid, rather convex in profile (
<figureCitation box="[727,780,660,682]" captionStart="Figs 511" captionStartId="3.[265,306,1193,1215]" captionTargetBox="[266,1108,300,1127]" captionTargetId="figure-0@3.[227,1123,257,1159]" captionTargetPageId="3" captionText="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi="http://doi.org/10.5281/zenodo.7166742" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Fig 5</figureCitation>
).
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">Color. Head, pronotum, scutellum, and abdomen dark brown to blackish brown, pronotum brown, antennal and legs dark reddish brown.</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">Head. Clypeus rectangular, bordered, anterior corners board rounded, anterior margin reflexed, incurved in the middle, surface with dense, coarse, setiferous punctures; fronto-</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,1127,798,820]" pageId="1" pageNumber="8">clypeal suture raised in the middle; frons densely setiferous punctate, coarsely in the middle,</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[227,851,825,847]" pageId="1" pageNumber="8">hair-like setae slightly long; antennal club longer than footstalks.</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">Pronotum wide trapezoid, surface densely punctate, densely covered with long hair-like setae; sides well crooked, anterior angles obtuse, apex rounded, posterior angles acute. Scutellum densely punctate, surface with long hair-like setae.</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">Elytra. Surface rugose with three developed costae between suture and humerus, sutural costa (as costa 1) and costae 2 complete and distinct, costa 3 weak; overall surface with setae which shorter than those on pronotum, setae on hind angles denser and shorter.</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
Propygidium with dense setae on sides, little on disc. Pygidium. apex rounded, lateral margins straight, slightly incurved before apex; surface densely with setae, also with long hairlike setae overall spread. Venter (
<figureCitation box="[551,606,1073,1095]" captionStart="Figs 14" captionStartId="2.[265,306,1588,1610]" captionTargetBox="[266,1089,272,1534]" captionTargetId="figure-0@2.[227,1110,257,1551]" captionTargetPageId="2" captionText="Figs 14. Tocama huangjianbini sp. n., habitus. 1, 2 holotype, male; 3, 4 paratype" figureDoi="http://doi.org/10.5281/zenodo.7166740" httpUri="https://zenodo.org/record/7166740/files/figure.png" pageId="1" pageNumber="8">Fig 2</figureCitation>
). Prosternal process hardly protruding. Metepisternum and metepimeron overall covered with long hair. Sides of abdominal sternites 15 weakly with disconnected maculation consisting of whitish scale-like setae; abdominal sternites with rather dense, short setae on sides, setae on the middle sparer, overall covered with another kind of long setae.
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" box="[265,1127,1211,1233]" pageId="1" pageNumber="8">Legs. Protibia slender, surface with short setae, tridentate with terminal tooth rather pro-</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">longed, middle tooth acute, basal tooth weakly developed; mesotibia and metatibia cylindrical, surface with short setae, inner surface with a row of strong setae, apex gradually widening, both with two unequal length spurs; surface of tarsal with dense, short setae.</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
Parameres (
<figureCitation box="[340,426,1321,1343]" captionStart="Figs 511" captionStartId="3.[265,306,1193,1215]" captionTargetBox="[266,1108,300,1127]" captionTargetId="figure-0@3.[227,1123,257,1159]" captionTargetPageId="3" captionText="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi="http://doi.org/10.5281/zenodo.7166742" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Figs 69</figureCitation>
) asymmetrical, apex of parameres without setae; in lateral view, apex of parameres rounded.
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
FEMALE. Similar with male in general appearance (compare
<figureCitation box="[867,952,1377,1399]" captionStart="Figs 14" captionStartId="2.[265,306,1588,1610]" captionTargetBox="[266,1089,272,1534]" captionTargetId="figure-0@2.[227,1110,257,1551]" captionTargetPageId="2" captionText="Figs 14. Tocama huangjianbini sp. n., habitus. 1, 2 holotype, male; 3, 4 paratype" figureDoi="http://doi.org/10.5281/zenodo.7166740" httpUri="https://zenodo.org/record/7166740/files/figure.png" pageId="1" pageNumber="8">Figs 14</figureCitation>
) but differs in the following characters: 1) body generally wider; 2) antennal shorter than footstalks; 3) protibia wider, hind tooth distinctly; 4) pygidium shorter (
<figureCitation box="[703,812,1432,1454]" captionStart="Figs 511" captionStartId="3.[265,306,1193,1215]" captionTargetBox="[266,1108,300,1127]" captionTargetId="figure-0@3.[227,1123,257,1159]" captionTargetPageId="3" captionText="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi="http://doi.org/10.5281/zenodo.7166742" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Figs 10, 11</figureCitation>
).
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
DIFFERENTIAL DIAGNOSIS. The new species can be distinguished from congeners by apex of parameres of male genitalia rounded and without setae (
<figureCitation box="[877,966,1487,1509]" captionStart="Figs 511" captionStartId="3.[265,306,1193,1215]" captionTargetBox="[266,1108,300,1127]" captionTargetId="figure-0@3.[227,1123,257,1159]" captionTargetPageId="3" captionText="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi="http://doi.org/10.5281/zenodo.7166742" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Figs 69</figureCitation>
). New species is most similar to
<taxonomicName authority="(Yu, Kobayashi &amp; Chu, 1998)" baseAuthorityName="Yu, Kobayashi &amp; Chu" baseAuthorityYear="1998" box="[379,809,1515,1537]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="8" phylum="Arthropoda" rank="species" species="formosana">
<emphasis box="[379,508,1515,1536]" italics="true" pageId="1" pageNumber="8">T. formosana</emphasis>
(Yu, Kobayashi &amp; Chu, 1998)
</taxonomicName>
in appearance, but protibia slen-
</paragraph>
<paragraph blockId="1.[227,1128,439,1592]" pageId="1" pageNumber="8">
derer (compare
<figureCitation box="[385,444,1542,1564]" captionStart="Figs 14" captionStartId="2.[265,306,1588,1610]" captionTargetBox="[266,1089,272,1534]" captionTargetId="figure-0@2.[227,1110,257,1551]" captionTargetPageId="2" captionText="Figs 14. Tocama huangjianbini sp. n., habitus. 1, 2 holotype, male; 3, 4 paratype" figureDoi="http://doi.org/10.5281/zenodo.7166740" httpUri="https://zenodo.org/record/7166740/files/figure.png" pageId="1" pageNumber="8">Fig. 1</figureCitation>
with Wang
<emphasis box="[570,616,1542,1564]" italics="true" pageId="1" pageNumber="8">et al</emphasis>
., 2015: fig. 8) and elytra not incurved distinctly in lateral view (compare
<figureCitation box="[442,498,1570,1592]" captionStart="Figs 511" captionStartId="3.[265,306,1193,1215]" captionTargetBox="[266,1108,300,1127]" captionTargetId="figure-0@3.[227,1123,257,1159]" captionTargetPageId="3" captionText="Figs 511. Tocama huangjianbini sp. n. 5 left lateral view of holotype; 69 male" figureDoi="http://doi.org/10.5281/zenodo.7166742" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="1" pageNumber="8">Fig. 5</figureCitation>
with Wang
<emphasis box="[616,659,1570,1591]" italics="true" pageId="1" pageNumber="8">et al</emphasis>
., 2015: fig. 24).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166740" ID-Zenodo-Dep="7166740" box="[265,1087,1588,1610]" httpUri="https://zenodo.org/record/7166740/files/figure.png" pageId="2" pageNumber="9" startId="2.[265,306,1588,1610]" targetBox="[266,1089,272,1534]" targetPageId="2">
<paragraph blockId="2.[227,1087,1588,1637]" box="[265,1087,1588,1610]" pageId="2" pageNumber="9">
Figs 14.
<taxonomicName authority="Wang, 2022" authorityName="Wang" authorityYear="2022" box="[360,579,1588,1609]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="9" phylum="Arthropoda" rank="species" species="huangjianbini" status="sp. n.">
<emphasis box="[360,579,1588,1609]" italics="true" pageId="2" pageNumber="9">Tocama huangjianbini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[584,637,1588,1609]" pageId="2" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
, habitus. 1, 2 holotype, male; 3, 4 paratype
</paragraph>
</caption>
<paragraph blockId="2.[227,1087,1588,1637]" box="[227,675,1615,1637]" pageId="2" pageNumber="9">female. 1, 3 dorsal view; 2, 4 ventral view.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166742" ID-Zenodo-Dep="7166742" box="[265,1127,1193,1215]" httpUri="https://zenodo.org/record/7166742/files/figure.png" pageId="3" pageNumber="10" startId="3.[265,306,1193,1215]" targetBox="[266,1108,300,1127]" targetPageId="3">
<paragraph blockId="3.[227,1127,1193,1270]" box="[265,1127,1193,1215]" pageId="3" pageNumber="10">
Figs 511.
<taxonomicName authority="Wang, 2022" authorityName="Wang" authorityYear="2022" box="[379,601,1193,1215]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="species" species="huangjianbini" status="sp. n.">
<emphasis box="[379,601,1193,1215]" italics="true" pageId="3" pageNumber="10">Tocama huangjianbini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[610,665,1193,1214]" pageId="3" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
5 left lateral view of holotype; 69 male
</paragraph>
</caption>
<paragraph blockId="3.[227,1127,1193,1270]" pageId="3" pageNumber="10">
genitalia of
<typeStatus box="[342,430,1220,1242]" pageId="3" pageNumber="10">holotype</typeStatus>
: 6 dorsal view, 7 ventral view, 8 lateral view from right, 9 lateral view from left; 10 pygidium of
<typeStatus box="[549,636,1248,1270]" pageId="3" pageNumber="10">holotype</typeStatus>
; 11 pygidium of female
<typeStatus box="[889,975,1249,1270]" pageId="3" pageNumber="10">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="10" type="discussion">
<paragraph blockId="3.[227,1127,1331,1573]" pageId="3" pageNumber="10">
REMARKS. Wang
<emphasis box="[471,520,1331,1353]" italics="true" pageId="3" pageNumber="10">et al</emphasis>
. (2015) described genus
<taxonomicName authorityName="REITTER" authorityYear="1902" box="[791,868,1332,1353]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis box="[791,868,1332,1353]" italics="true" pageId="3" pageNumber="10">Tocama</emphasis>
</taxonomicName>
as: “possessing strongly asymmetrical parameres of the male genitalia with a bundle of curled, robust setae at the apex of the parameres”. So far,
<taxonomicName authority="Wang, 2022" authorityName="Wang" authorityYear="2022" box="[549,770,1386,1408]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="species" species="huangjianbini" status="sp. n.">
<emphasis box="[549,770,1386,1408]" italics="true" pageId="3" pageNumber="10">Tocama huangjianbini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[779,834,1386,1407]" pageId="3" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
is unique species of
<taxonomicName authorityName="REITTER" authorityYear="1902" box="[1049,1126,1387,1408]" class="Insecta" family="Melolonthidae" genus="Tocama" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis box="[1049,1126,1387,1408]" italics="true" pageId="3" pageNumber="10">Tocama</emphasis>
</taxonomicName>
with apex of parameres without setae.
</paragraph>
<paragraph blockId="3.[227,1127,1331,1573]" box="[265,1127,1441,1463]" pageId="3" pageNumber="10">HABITAT. The habitat of the new species is a seaside village of Pintan Island</paragraph>
<paragraph blockId="3.[227,1127,1331,1573]" pageId="3" pageNumber="10">
(130 kilometers northwest of
<collectingCountry box="[511,652,1469,1491]" name="Taiwan Island" pageId="3" pageNumber="10">Taiwan Island</collectingCountry>
), Fujian as shown in
<figureCitation box="[860,972,1469,1491]" captionStart="Figs 1215" captionStartId="4.[265,306,949,971]" captionTargetBox="[232,1113,265,927]" captionTargetId="figure-0@4.[227,1117,257,933]" captionTargetPageId="4" captionText="Figs 1215. Type locality of Tocama huangjianbini sp. n. (Pintan Island, Longfengtou Coast). Photos by Jian-Bin Huang." figureDoi="http://doi.org/10.5281/zenodo.7166744" httpUri="https://zenodo.org/record/7166744/files/figure.png" pageId="3" pageNumber="10">Figs 1215</figureCitation>
. The specimens examined were collected around light at night.
</paragraph>
</subSubSection>
<subSubSection box="[265,1099,1524,1546]" pageId="3" pageNumber="10" type="distribution">
<paragraph blockId="3.[227,1127,1331,1573]" box="[265,1099,1524,1546]" pageId="3" pageNumber="10">
<materialsCitation box="[265,1096,1524,1546]" country="China" location="South" pageId="3" pageNumber="10" specimenCount="1" stateProvince="Fujian">
DISTRIBUTION. Known only from the
<typeStatus box="[658,699,1524,1545]" pageId="3" pageNumber="10">type</typeStatus>
locality in
<collectingRegion box="[808,963,1524,1546]" country="China" name="Fujian" pageId="3" pageNumber="10">Fujian Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:2C16504AFFA8AA75FF1CFE84D117F99D:A160B787FFAAAA75FC32FA4FD14AF9B2" box="[973,1029,1524,1546]" country="China" name="South" pageId="3" pageNumber="10" stateProvince="Fujian">South</location>
<collectingCountry box="[1035,1096,1524,1546]" name="China" pageId="3" pageNumber="10">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[265,1112,1551,1573]" pageId="3" pageNumber="10" type="etymology">
<paragraph blockId="3.[227,1127,1331,1573]" box="[265,1112,1551,1573]" pageId="3" pageNumber="10">
ETYMOLOGY. The new species is named after Jian-Bin Huang, who collected all
<typeStatus box="[1056,1109,1552,1573]" pageId="3" pageNumber="10">types</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>