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<document id="ECBC79AC37C6A1448DFF669C0E37AAFC" ID-DOI="10.11646/phytotaxa.656.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13213072" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722614541709" checkinUser="felipe" docAuthor="Croat, Thomas B., Cedeño-Fonseca, Marco &amp; Ortiz, Orlando O." docDate="2024" docId="23768787FF894475DFC2FF72FF20F93A" docLanguage="en" docName="phytotaxa.156.1.1.pdf" docOrigin="Phytotaxa 656 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.656.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Monstera coclensis Croat, sp. nov." docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="DF4FFFFFFFB94445DF4AFFECFFF8FFD2" masterDocTitle="Revision of Monstera (Araceae: Monsteroideae) of Central America" masterLastPageNumber="197" masterPageNumber="1" pageNumber="49" updateTime="1726142891962" updateUser="guilherme">
<mods:mods id="5DEB9B0DD4A97952FAFB6ECB7201CB2A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="13B8374069E8AAF3B286867AEB174630">
<mods:title id="FBBFCB41B9F10AAF1683F7F1245D3197">Revision of Monstera (Araceae: Monsteroideae) of Central America</mods:title>
</mods:titleInfo>
<mods:name id="E49E9D690F7AB676BC315ED76DAB006E" type="personal">
<mods:role id="29FEBC4F6451F51169E0B7C8949FEEB2">
<mods:roleTerm id="FE677F970606FA0EC6A5DC204BBADCAC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6CA9B96F9CFB34EBB5DB79CCB2B9BA7B">Croat, Thomas B.</mods:namePart>
<mods:nameIdentifier id="72CC51D88F99683DBC625A0BA2FF8348" type="ORCID">0000-0002-1804-0532</mods:nameIdentifier>
<mods:affiliation id="329532417F96D4FBCBE88B382B218AC2">P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; email: thomas. croat @ mobot. org; https: // orcid. org / 0000 - 0002 - 1804 - 0532</mods:affiliation>
<mods:nameIdentifier id="D612FD31AB807D0C56C2AAEA34EECADA" type="email">thomas.croat@mobot.org</mods:nameIdentifier>
</mods:name>
<mods:name id="70CE0AEC52F9DFD29D0B1F14A970F2DA" type="personal">
<mods:role id="D8808962D8649A4FBA1485FB86D8344B">
<mods:roleTerm id="2A6A6ACC3EE404121769060E44462D50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74BF03C1CFE9391C90DD70128FD4DB61">Cedeño-Fonseca, Marco</mods:namePart>
<mods:nameIdentifier id="C5B5D6B727C1005BC378572AEB72348F" type="ORCID">0000-0003-0119-0427</mods:nameIdentifier>
<mods:affiliation id="B4211027A4CDA573B7365B94ED63E0BF">P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Strasse 6 - 8, D- 14195 Berlin, Germany. &amp; Herbario Luis Fournier Origgi (USJ), Centro de Investigación en Biodiversidad y Ecología Tropical, Universidad de Costa Rica, Apdo. 11501 - 2060, San José, Costa Rica. &amp; Jardín Botánico Lankester, Universidad de Costa Rica, Apartado 302 - 7050 Cartago, Costa Rica. &amp; email: marcovf. 09 @ gmail. com; https: // orcid. org / 0000 - 0003 - 0119 - 0427</mods:affiliation>
<mods:nameIdentifier id="4FCAA3D639672BF151D9E64AF1638F7B" type="email">marcovf.09@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="49C5D0F2EC8F3920C75E7CC3D7E59257" type="personal">
<mods:role id="0F8A100CE9E3B1188ACCA58708D3B6FA">
<mods:roleTerm id="85A01319DD964ACA185E0F634FC8D751">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08F98CFD7D424E1750524E5A4FBD397A">Ortiz, Orlando O.</mods:namePart>
<mods:nameIdentifier id="6071EB5A8AE44639FFDB0AAF3C81A469" type="ORCID">0000-0002-7805-0046</mods:nameIdentifier>
<mods:affiliation id="52000A9B5360B204A3F1EF9F40990162">P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Strasse 6 - 8, D- 14195 Berlin, Germany. &amp; P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; Herbario PMA, Universidad de Panamá, Estafeta Universitaria, Panama City, Republic of Panamá. &amp; P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; Coiba Scientific Station (COIBA AIP), Clayton, Panama City, Panama. &amp; P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A. &amp; email: ortizopma @ gmail. com; https: // orcid. org / 0000 - 0002 - 7805 - 0046 * Corresponding author. e-mail: marcovf. 09 @ gmail. com &amp; P. A. Schulze Curator, Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="A1903174A642A2D4680E00A676457DBB" type="email">ortizopma@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="59904E5388BCCA2DE1143D6BA2D46381">text</mods:typeOfResource>
<mods:relatedItem id="5FA8867B71F9AD4D0AD7A5A9F3864B07" type="host">
<mods:titleInfo id="8230A3C4B352954A9D1BBFB4D394DA65">
<mods:title id="5D4FD33BCB085981E2FBBD6D9338036C">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="B0E27415C4745154DE053CCCFFBA93B0">
<mods:date id="02886BB774E79193A4846A433CD9F58E">2024</mods:date>
<mods:detail id="B17E90D158A8191398C5FCB3282C345E" type="pubDate">
<mods:number id="F42544B10FC2B7344D471BB04F37B138">2024-06-27</mods:number>
</mods:detail>
<mods:detail id="915D3F66FE92466BA835C23F096178FA" type="volume">
<mods:number id="8A0CE85062474C1F1E8899619703D770">656</mods:number>
</mods:detail>
<mods:detail id="517A522D3DD22E4BADB4970E3F3496D2" type="issue">
<mods:number id="A56616D3DFE5F6D090C9E769461F55FC">1</mods:number>
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<mods:start id="A7AB405C94A3586712BA00DBA4B9D35F">1</mods:start>
<mods:end id="10C44BA375762C6F6FFAAA95F9BB84D7">197</mods:end>
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<mods:location id="8B9644CFDF6B68C4A44CA5B175255049">
<mods:url id="3259FB228C81BB70906D15E05519B749">http://dx.doi.org/10.11646/phytotaxa.656.1.1</mods:url>
</mods:location>
<mods:classification id="60FD26C3A3E64DD29EEAB91043764379">journal article</mods:classification>
<mods:identifier id="D0EA8194D81FB25322F79552AFE16082" type="DOI">10.11646/phytotaxa.656.1.1</mods:identifier>
<mods:identifier id="AC899E85DCCDE3103BD3ECD619EAB512" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="17B052DDDB16D27B29F6C9172F5B428F" type="Zenodo-Dep">13213072</mods:identifier>
</mods:mods>
<treatment id="23768787FF894475DFC2FF72FF20F93A" ID-DOI="http://doi.org/10.5281/zenodo.13365765" ID-Zenodo-Dep="13365765" LSID="urn:lsid:plazi:treatment:23768787FF894475DFC2FF72FF20F93A" httpUri="http://treatment.plazi.org/id/23768787FF894475DFC2FF72FF20F93A" lastPageNumber="49" pageId="48" pageNumber="49">
<subSubSection id="E3C5651AFF894475DFC2FF72FDBDFF6B" box="[136,581,158,185]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="AB603691FF894475DFC2FF72FDBDFF6B" blockId="48.[136,581,158,185]" box="[136,581,158,185]" pageId="48" pageNumber="49">
9.
<taxonomicName id="6CDF4D12FF894475DFF7FF73FE1BFF6A" authority="Croat" authorityName="Croat" box="[189,483,158,185]" class="Liliopsida" family="Araceae" genus="Monstera" kingdom="Plantae" order="Alismatales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="coclensis" status="sp. nov.">
<emphasis id="99ABEA83FF894475DFF7FF73FE63FF6B" bold="true" box="[189,411,158,185]" italics="true" pageId="48" pageNumber="49">Monstera coclensis</emphasis>
Croat
</taxonomicName>
,
<emphasis id="99ABEA83FF894475DEA4FF73FDBDFF6B" bold="true" box="[494,581,159,185]" italics="true" pageId="48" pageNumber="49">
<taxonomicNameLabel id="829857F8FF894475DEA4FF73FDBDFF6B" box="[494,581,159,185]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFC2FF06FC3DFEF1" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="AB603691FF894475DFC2FF06FC3DFEF1" blockId="48.[136,1452,234,291]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFC2FF06FC3DFEF1" italics="true" pageId="48" pageNumber="49">The species is characterized by its petioles sheathed to the geniculum, a persistent sheath, entire, narrowly ovate-elliptic, non-perforate, dark brown-drying blades with the spadix more than twice as long as the spathe.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFC2FEBAFDB4FE5D" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="AB603691FF894475DFC2FEBAFDB4FE5D" blockId="48.[136,1451,342,400]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFC2FEBAFF3AFEB9" bold="true" box="[136,194,342,363]" pageId="48" pageNumber="49">Type:</emphasis>
—
<materialsCitation id="1BB73CCCFF894475DF91FEBAFE48FE42" collectionCode="PMA" collectorName="Sytsma &amp; Andersson" country="Panama" elevation="1250" location="El Cope" municipality="Above El Potroso" pageId="48" pageNumber="49" specimenCode="MO-2908952" specimenCount="2" stateProvince="Cocle" typeStatus="holotype">
<collectingCountry id="D3C87601FF894475DF91FEBAFEBCFEB9" box="[219,324,342,363]" name="Panama" pageId="48" pageNumber="49">PANAMA</collectingCountry>
.
<collectingRegion id="691BF873FF894475DE04FEBAFE72FEBE" box="[334,394,342,364]" country="Panama" name="Cocle" pageId="48" pageNumber="49">Coclé</collectingRegion>
:
<collectorName id="062A5347FF894475DED9FEBAFDB9FEB9" box="[403,577,342,364]" pageId="48" pageNumber="49">
<collectingMunicipality id="4B04ACEBFF894475DED9FEBAFDB9FEB9" box="[403,577,342,364]" pageId="48" pageNumber="49">Above El Potroso</collectingMunicipality>
</collectorName>
sawmill at
<collectorName id="062A5347FF894475DDFCFEBAFC6CFEB9" box="[694,916,342,364]" pageId="48" pageNumber="49">Continental Divide, N.</collectorName>
of
<collectorName id="062A5347FF894475DCFDFEBAFBF1FEB9" box="[951,1033,342,364]" pageId="48" pageNumber="49">
<location id="AE00604AFF894475DCFDFEBAFBF1FEB9" LSID="urn:lsid:plazi:treatment:23768787FF894475DFC2FF72FF20F93A:AE00604AFF894475DCFDFEBAFBF1FEB9" box="[951,1033,342,364]" country="Panama" municipality="Above El Potroso" name="El Cope" pageId="48" pageNumber="49" stateProvince="Cocle">El Copé</location>
</collectorName>
,
<quantity id="6C279B74FF894475DB5FFEBAFB65FEB9" box="[1045,1181,342,363]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<elevation id="20F2D1A2FF894475DB5FFEBAFB65FEB9" box="[1045,1181,342,363]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</elevation>
</quantity>
,
<emphasis id="99ABEA83FF894475DBE3FEBAFA53FEB9" box="[1193,1451,342,363]" italics="true" pageId="48" pageNumber="49">
<collectorName id="062A5347FF894475DBE3FEBAFB15FEB9" box="[1193,1261,342,363]" pageId="48" pageNumber="49">Sytsma</collectorName>
&amp;
<collectorName id="062A5347FF894475DA44FEBAFA8BFEB9" box="[1294,1395,342,363]" pageId="48" pageNumber="49">Andersson</collectorName>
4529
</emphasis>
(
<typeStatus id="74648833FF894475DF8EFE96FEE2FE42" box="[196,282,378,400]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
<specimenCode id="FB799EEAFF894475DE55FE96FE52FE42" box="[287,426,378,400]" collectionCode="MO-" pageId="48" pageNumber="49">MO-2908952</specimenCode>
!,
</materialsCitation>
<materialsCitation id="1BB73CCCFF894475DEFCFE96FDB0FE5D" box="[438,584,378,400]" collectionCode="PMA" collectorName="Sytsma &amp; Andersson" country="Panama" elevation="1250" location="El Cope" municipality="Above El Potroso" pageId="48" pageNumber="49" specimenCode="MO-2908952" specimenCount="2" stateProvince="Cocle" typeStatus="holotype">
<typeStatus id="74648833FF894475DEFCFE96FE03FE42" box="[438,507,378,400]" pageId="48" pageNumber="49" type="isotype">isotype</typeStatus>
<collectionCode id="CDCEAE54FF894475DD4BFE96FDC6FE5D" box="[513,574,378,399]" country="Canada" name="Provincial Museum of Alberta" pageId="48" pageNumber="49">PMA</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFC2FE52FD22FB1E" pageId="48" pageNumber="49" type="description">
<paragraph id="AB603691FF894475DFC2FE52FD22FB1E" blockId="48.[136,1452,446,1768]" pageId="48" pageNumber="49">
Nomadic vine, appressed-climbing habit. SEEDLINGS: unknown. JUVENILE PLANTS: root climbers;
<emphasis id="99ABEA83FF894475DA64FE52FA89FE0A" bold="true" box="[1326,1393,446,472]" pageId="48" pageNumber="49">stems</emphasis>
dark green, minutely spotted, with yellow-brown deciduous epidermis, drying surface dark brown, matte, longitudinally striated;
<emphasis id="99ABEA83FF894475DFA3FDEAFE9FFDF2" bold="true" box="[233,359,518,544]" pageId="48" pageNumber="49">internodes</emphasis>
1–3(10) cm long,
<quantity id="6C279B74FF894475DD7CFDEAFD4BFDF3" box="[566,691,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="48" pageNumber="49" unit="cm" value="1.05" valueMax="1.5" valueMin="0.6">0.6–1.5 cm</quantity>
diam., 1.6–2.0 times longer than wide;
<emphasis id="99ABEA83FF894475DB3AFDEAFB44FDF2" bold="true" box="[1136,1212,518,544]" pageId="48" pageNumber="49">blades</emphasis>
dark green, minutely stained yellow-brown, drying dark brown, matte, longitudinally striated, cordate or rounded at base, faintly glaucous adaxially.ADULT PLANTS: root climbers;
<emphasis id="99ABEA83FF894475DD3EFDA2FD4FFDBA" bold="true" box="[628,695,590,616]" pageId="48" pageNumber="49">stems</emphasis>
drying dark brown, matte, ribbed;
<emphasis id="99ABEA83FF894475DB73FDA2FB4FFDBA" bold="true" box="[1081,1207,590,616]" pageId="48" pageNumber="49">internodes</emphasis>
with a ripped yellow-brown epidermis, shorter than wide,
<quantity id="6C279B74FF894475DD55FD9EFD92FD5F" box="[543,618,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="48" pageNumber="49" unit="cm" value="1.5">1.5 cm</quantity>
diam.;
<emphasis id="99ABEA83FF894475DDF7FD9EFCF4FD5E" bold="true" box="[701,780,626,652]" pageId="48" pageNumber="49">petiole</emphasis>
pale green, sometimes with pale dots, dark brown when dry, matte,
<quantity id="6C279B74FF894475DF9FFD7AFEBEFD63" box="[213,326,662,689]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="48" pageNumber="49" unit="cm" value="25.5" valueMax="30.0" valueMin="21.0">21–30 cm</quantity>
long, prominently sheathed to base of the geniculum or
<quantity id="6C279B74FF894475DC80FD7AFBFAFD63" box="[970,1026,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="cm" value="1.0">1 cm</quantity>
above the geniculum;
<emphasis id="99ABEA83FF894475DA4CFD7AFA54FD62" bold="true" box="[1286,1452,662,688]" pageId="48" pageNumber="49">petiole sheath</emphasis>
persistent; geniculum markedly sulcate;
<emphasis id="99ABEA83FF894475DD06FD56FD60FD06" bold="true" box="[588,664,698,724]" pageId="48" pageNumber="49">blades</emphasis>
narrowly ovate-elliptic, equilateral, one side
<quantity id="6C279B74FF894475DBDEFD56FB1FFD07" box="[1172,1255,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="48" pageNumber="49" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
wide, rounded to obtuse at base, coriaceous, short acuminate, drying dark-green to brown, matte or faintly glossy, (18)32–34 × (6)
<quantity id="6C279B74FF894475DA38FD32FF55FCCF" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="48" pageNumber="49" unit="cm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 cm</quantity>
, (2.1)2.3–2.9 times longer than wide;
<emphasis id="99ABEA83FF894475DD28FCEEFD4CFCCE" bold="true" box="[610,692,770,796]" pageId="48" pageNumber="49">midrib</emphasis>
ribbed adaxially, convex abaxially, drying light-brown or reddish-brown on both surfaces;
<emphasis id="99ABEA83FF894475DEDFFCCAFD76FC92" bold="true" box="[405,654,806,832]" pageId="48" pageNumber="49">primary lateral veins</emphasis>
7–10 per side, strongly sunken adaxially, prominent abaxially, departing midrib at 30–45°, drying narrowly convex and concolorous adaxially, narrowly rounded and slightly paler abaxially,
<emphasis id="99ABEA83FF894475DFC2FC82FEB2FC5A" bold="true" box="[136,330,878,904]" pageId="48" pageNumber="49">secondary veins</emphasis>
parallel and prominent, reticulate at the margin;
<emphasis id="99ABEA83FF894475DCD8FC82FBB1FC5A" bold="true" box="[914,1097,878,904]" pageId="48" pageNumber="49">collective veins</emphasis>
slightly visible;
<emphasis id="99ABEA83FF894475DA5BFC82FA53FC5A" bold="true" box="[1297,1451,878,904]" pageId="48" pageNumber="49">fenestrations</emphasis>
apparently absent;
<emphasis id="99ABEA83FF894475DE11FC7EFE45FC7E" bold="true" box="[347,445,914,940]" pageId="48" pageNumber="49">margins</emphasis>
entire. INFLORESCENCES on ascending stems;
<emphasis id="99ABEA83FF894475DCB8FC7EFBA5FC7E" bold="true" box="[1010,1117,914,940]" pageId="48" pageNumber="49">peduncle</emphasis>
smooth, pale green or brown, drying dark, matte to faintly glossy,
<quantity id="6C279B74FF894475DD6BFC5AFD68FC03" box="[545,656,950,977]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" metricValueMax="2.1" metricValueMin="1.3" pageId="48" pageNumber="49" unit="cm" value="17.0" valueMax="21.0" valueMin="13.0">13–21 cm</quantity>
long,
<quantity id="6C279B74FF894475DD93FC5AFCBCFC03" box="[729,836,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="48" pageNumber="49" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
diam.;
<emphasis id="99ABEA83FF894475DCD3FC5AFC1FFC02" bold="true" box="[921,999,950,976]" pageId="48" pageNumber="49">spathe</emphasis>
acuminate, white or greenish externally and white or creamy internally at anthesis, completely open in the apical and basal part, drying dark brownish-purple, 11–26 × 8.2–15.0 cm, up to
<quantity id="6C279B74FF894475DE9BFC12FDF2FBCB" box="[465,522,1022,1049]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="cm" value="4.0">4 cm</quantity>
longer than the spadix;
<emphasis id="99ABEA83FF894475DC69FC12FC89FBCA" bold="true" box="[803,881,1022,1048]" pageId="48" pageNumber="49">spadix</emphasis>
white or creamy (both during development and a anthesis),
<quantity id="6C279B74FF894475DFB6FBCEFE83FBEF" box="[252,379,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" metricValueMax="9.5" metricValueMin="3.7" pageId="48" pageNumber="49" unit="cm" value="6.6" valueMax="9.5" valueMin="3.7">3.7–9.5 cm</quantity>
long,
<quantity id="6C279B74FF894475DE8CFBCEFDBDFBEF" box="[454,581,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="48" pageNumber="49" unit="cm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 cm</quantity>
diam., 2.1–4.6 times longer than wide;
<emphasis id="99ABEA83FF894475DB5BFBCEFAF9FBEE" bold="true" box="[1041,1281,1058,1084]" pageId="48" pageNumber="49">basal sterile flowers</emphasis>
<quantity id="6C279B74FF894475DA40FBCEFA91FBEF" box="[1290,1385,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="48" pageNumber="49" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long;
<emphasis id="99ABEA83FF894475DFC2FBAAFED7FBB2" bold="true" box="[136,303,1094,1120]" pageId="48" pageNumber="49">fertile flowers</emphasis>
<quantity id="6C279B74FF894475DE7DFBAAFE6EFBB3" box="[311,406,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="48" pageNumber="49" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long; ovary rectangular in longitudinal section, ribbed, 3–4 ×
<quantity id="6C279B74FF894475DB16FBAAFB1FFBB3" box="[1116,1255,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="48" pageNumber="49" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
; style rounded to irregularly angular, more or less truncate to markedly sunken at the medial part with raised margins and moderately irregular-lumpy, densely papillose, 3–5 ×
<quantity id="6C279B74FF894475DD16FB62FD28FB7B" box="[604,720,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="48" pageNumber="49" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
; stigma yellow-orange, moderately raised, margins slightly paler, narrow and erect;
<emphasis id="99ABEA83FF894475DE19FB5EFE5FFB1E" bold="true" box="[339,423,1202,1228]" pageId="48" pageNumber="49">berries</emphasis>
unknown;
<emphasis id="99ABEA83FF894475DD6FFB5EFD9BFB1E" bold="true" box="[549,611,1202,1228]" pageId="48" pageNumber="49">seeds</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFF7FB3AFD45FAEB" pageId="48" pageNumber="49" type="distribution">
<paragraph id="AB603691FF894475DFF7FB3AFC73FAC7" blockId="48.[136,1452,446,1768]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFF7FB3AFE0AFB22" bold="true" box="[189,498,1238,1264]" pageId="48" pageNumber="49">Distribution and ecology:</emphasis>
—
<taxonomicName id="6CDF4D12FF894475DD44FB3BFD12FB22" box="[526,746,1238,1264]" class="Liliopsida" family="Araceae" genus="Monstera" kingdom="Plantae" order="Alismatales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="coclensis">
<emphasis id="99ABEA83FF894475DD44FB3BFD12FB22" box="[526,746,1238,1264]" italics="true" pageId="48" pageNumber="49">Monstera coclensis</emphasis>
</taxonomicName>
is endemic to the
<collectingCountry id="D3C87601FF894475DC89FB3BFBE4FB23" box="[963,1052,1239,1265]" name="Panama" pageId="48" pageNumber="49">Panama</collectingCountry>
from the El Copé region of
<collectingRegion id="691BF873FF894475DA20FB3AFF28FAC6" country="Panama" name="Cocle" pageId="48" pageNumber="49">Coclé region</collectingRegion>
of
<collectingRegion id="691BF873FF894475DFBFFB16FE5BFAC6" box="[245,419,1274,1300]" country="Panama" name="Cocle" pageId="48" pageNumber="49">Coclé Province</collectingRegion>
, in
<emphasis id="99ABEA83FF894475DE86FB17FD3FFAC6" box="[460,711,1274,1300]" italics="true" pageId="48" pageNumber="49">Premontane wet forest</emphasis>
at
<quantity id="6C279B74FF894475DDA3FB16FC7FFAC7" box="[745,903,1274,1301]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</quantity>
.
</paragraph>
<paragraph id="AB603691FF894475DFF7FAF2FD45FAEB" blockId="48.[136,1452,446,1768]" box="[189,701,1310,1337]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFF7FAF2FEBBFAEA" bold="true" box="[189,323,1310,1336]" pageId="48" pageNumber="49">Phenology:</emphasis>
—Flowering in January and May.
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFF7FAAEFB6FFA8E" box="[189,1175,1346,1373]" pageId="48" pageNumber="49" type="etymology">
<paragraph id="AB603691FF894475DFF7FAAEFB6FFA8E" blockId="48.[136,1452,446,1768]" box="[189,1175,1346,1373]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFF7FAAEFEB0FA8E" bold="true" box="[189,328,1346,1372]" pageId="48" pageNumber="49">Etymology:</emphasis>
—The species named for the Province of
<collectingRegion id="691BF873FF894475DC50FAAEFCA3FA8E" box="[794,859,1346,1372]" country="Panama" name="Cocle" pageId="48" pageNumber="49">Coclé</collectingRegion>
where it was first collected.
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFF7FA8AFF07F9C2" pageId="48" pageNumber="49" type="discussion">
<paragraph id="AB603691FF894475DFF7FA8AFF07F9C2" blockId="48.[136,1452,446,1768]" pageId="48" pageNumber="49">
<emphasis id="99ABEA83FF894475DFF7FA8AFEBDFA52" bold="true" box="[189,325,1382,1408]" pageId="48" pageNumber="49">Discussion:</emphasis>
—The species is a member of sect.
<taxonomicName id="6CDF4D12FF894475DD83FA8BFCCCFA52" box="[713,820,1383,1408]" class="Liliopsida" family="Araceae" genus="Monstera" kingdom="Plantae" order="Alismatales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="genus">
<emphasis id="99ABEA83FF894475DD83FA8BFCCCFA52" box="[713,820,1383,1408]" italics="true" pageId="48" pageNumber="49">Monstera</emphasis>
</taxonomicName>
, it is probably most easily confused with the morphotype of
<taxonomicName id="6CDF4D12FF894475DFE2FA67FE8CFA76" authorityName="Engler" authorityYear="1905" box="[168,372,1418,1444]" class="Liliopsida" family="Araceae" genus="Monstera" kingdom="Plantae" order="Alismatales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="epipremnoides">
<emphasis id="99ABEA83FF894475DFE2FA67FE8CFA76" box="[168,372,1418,1444]" italics="true" pageId="48" pageNumber="49">M. epipremnoides</emphasis>
</taxonomicName>
from southwestern
<collectingCountry id="D3C87601FF894475DD13FA66FD21FA76" box="[601,729,1418,1444]" name="Costa Rica" pageId="48" pageNumber="49">Costa Rica</collectingCountry>
. That species differs in having more ovate blades, ca 2.1 times longer than broad, a broader petiole sheath, the stigmas subrounded and barely more than
<quantity id="6C279B74FF894475DBCDFA42FB31FA1B" box="[1159,1225,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
long and especially by having the spathe only slightly longer than the spadix at anthesis (vs. over 2.5 times longer than wide in
<taxonomicName id="6CDF4D12FF894475DAC7FA3FFF17F9C2" class="Liliopsida" family="Araceae" genus="Monstera" kingdom="Plantae" order="Alismatales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="coclensis">
<emphasis id="99ABEA83FF894475DAC7FA3FFF17F9C2" italics="true" pageId="48" pageNumber="49">M. coclensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E3C5651AFF894475DFF7F9F6FF20F93A" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="AB603691FF894475DFF7F9F6FF20F93A" blockId="48.[136,1452,446,1768]" pageId="48" pageNumber="49">
<materialsCitation id="1BB73CCCFF894475DFF7F9F6FE76F9AE" collectingDate="1981-05-13" collectionCode="MO" collectorName="K. J. Sytsma &amp; L. Andersson" country="Panama" elevation="1250" latitude="8.676667" location="Continental Divide" longLatPrecision="21" longitude="-80.61" municipality="Above El Potroso" pageId="48" pageNumber="49" specimenCount="2" stateProvince="Cocle">
<emphasis id="99ABEA83FF894475DFF7F9F6FD21F9E6" bold="true" box="[189,729,1562,1588]" pageId="48" pageNumber="49">
Additional specimens examined (
<typeStatus id="74648833FF894475DD05F9F6FD34F9E6" box="[591,716,1562,1588]" pageId="48" pageNumber="49" type="paratype">paratypes</typeStatus>
):
</emphasis>
—
<collectingCountry id="D3C87601FF894475DDBCF9F6FC88F9E6" box="[758,880,1562,1588]" name="Panama" pageId="48" pageNumber="49">PANAMA</collectingCountry>
.
<emphasis id="99ABEA83FF894475DC37F9F5FC31F9E6" bold="true" box="[893,969,1561,1588]" pageId="48" pageNumber="49">
<collectingRegion id="691BF873FF894475DC37F9F5FC3CF9E6" box="[893,964,1561,1588]" country="Panama" name="Cocle" pageId="48" pageNumber="49">Coclé</collectingRegion>
:
</emphasis>
<collectorName id="062A5347FF894475DC9BF9F6FB67F9E7" box="[977,1183,1562,1589]" pageId="48" pageNumber="49">
<collectingMunicipality id="4B04ACEBFF894475DC9BF9F6FB67F9E7" box="[977,1183,1562,1589]" pageId="48" pageNumber="49">Above El Potroso</collectingMunicipality>
</collectorName>
sawmill at
<location id="AE00604AFF894475DA63F9F6FF2FF98A" LSID="urn:lsid:plazi:treatment:23768787FF894475DFC2FF72FF20F93A:AE00604AFF894475DA63F9F6FF2FF98A" country="Panama" latitude="8.676667" longLatPrecision="21" longitude="-80.61" municipality="Above El Potroso" name="Continental Divide" pageId="48" pageNumber="49" stateProvince="Cocle">Continental Divide</location>
,
<locationDeviation id="BAD22FB3FF894475DFAEF9D3FE46F98B" box="[228,446,1598,1625]" location="El Cope. Low" pageId="48" pageNumber="49">N of El Cope. Low</locationDeviation>
cloud forest,
<geoCoordinate id="CEEB5056FF894475DD16F9D2FD1DF98A" box="[604,741,1598,1624]" degrees="08" direction="north" minutes="40" orientation="latitude" pageId="48" pageNumber="49" precision="15" seconds="36" value="8.676667">08°40’36”N</geoCoordinate>
<geoCoordinate id="CEEB5056FF894475DDA7F9D2FC73F98A" box="[749,907,1598,1624]" degrees="080" direction="west" minutes="36" orientation="longitude" pageId="48" pageNumber="49" precision="15" seconds="36" value="-80.61">080°36’36”W</geoCoordinate>
,
<quantity id="6C279B74FF894475DCDDF9D2FBCEF98B" box="[919,1078,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<elevation id="20F2D1A2FF894475DCDDF9D2FBCEF98B" box="[919,1078,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</elevation>
</quantity>
,
<date id="DF611051FF894475DB09F9D2FB24F98A" box="[1091,1244,1598,1625]" pageId="48" pageNumber="49" value="1981-05-13">
<collectingDate id="CF25E9B9FF894475DB09F9D2FB24F98A" box="[1091,1244,1598,1625]" pageId="48" pageNumber="49" value="1981-05-13">13 May 1981</collectingDate>
</date>
,
<emphasis id="99ABEA83FF894475DBA0F9D3FEC4F9AE" italics="true" pageId="48" pageNumber="49">
<collectorName id="062A5347FF894475DBA0F9D3FA96F98B" box="[1258,1390,1599,1625]" pageId="48" pageNumber="49">K.J. Sytsma</collectorName>
&amp;
<collectorName id="062A5347FF894475DADFF9D3FF06F9AE" pageId="48" pageNumber="49">L. Andersson</collectorName>
4585
</emphasis>
(
<collectionCode id="CDCEAE54FF894475DE01F98EFE79F9AE" box="[331,385,1634,1660]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="48" pageNumber="49" type="Herbarium">MO</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="1BB73CCCFF894475DEDDF98EFBAFF972" collectingDate="1981-05-13" collectionCode="MO" collectorName="K. J. Sytsma &amp; L. Andersson" country="Panama" elevation="1250" latitude="8.676667" location="Continental Divide" longLatPrecision="21" longitude="-80.61" municipality="Above El Potroso" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Cocle">
<collectorName id="062A5347FF894475DEDDF98EFDA4F9AF" box="[407,604,1634,1661]" pageId="48" pageNumber="49">
<collectingMunicipality id="4B04ACEBFF894475DEDDF98EFDA4F9AF" box="[407,604,1634,1661]" pageId="48" pageNumber="49">Above El Potroso</collectingMunicipality>
</collectorName>
sawmill at
<location id="AE00604AFF894475DD90F98EFC49F9AE" LSID="urn:lsid:plazi:treatment:23768787FF894475DFC2FF72FF20F93A:AE00604AFF894475DD90F98EFC49F9AE" box="[730,945,1634,1660]" country="Panama" latitude="8.676667" longLatPrecision="21" longitude="-80.61" municipality="Above El Potroso" name="Continental Divide" pageId="48" pageNumber="49" stateProvince="Cocle">Continental Divide</location>
,
<locationDeviation id="BAD22FB3FF894475DCF1F98FFB74F9AF" box="[955,1164,1634,1661]" location="El Cope. Low" pageId="48" pageNumber="49">N of El Cope. Low</locationDeviation>
cloud forest,
<geoCoordinate id="CEEB5056FF894475DA68F98EFA53F9AE" box="[1314,1451,1634,1660]" degrees="08" direction="north" minutes="40" orientation="latitude" pageId="48" pageNumber="49" precision="15" seconds="36" value="8.676667">08°40’36”N</geoCoordinate>
<geoCoordinate id="CEEB5056FF894475DFC2F96AFEDDF972" box="[136,293,1670,1696]" degrees="080" direction="west" minutes="36" orientation="longitude" pageId="48" pageNumber="49" precision="15" seconds="36" value="-80.61">080°36’36”W</geoCoordinate>
,
<quantity id="6C279B74FF894475DE78F96AFE28F973" box="[306,464,1670,1697]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<elevation id="20F2D1A2FF894475DE78F96AFE28F973" box="[306,464,1670,1697]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="48" pageNumber="49" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</elevation>
</quantity>
,
<date id="DF611051FF894475DE97F96AFD8FF972" box="[477,631,1670,1697]" pageId="48" pageNumber="49" value="1981-05-13">
<collectingDate id="CF25E9B9FF894475DE97F96AFD8FF972" box="[477,631,1670,1697]" pageId="48" pageNumber="49" value="1981-05-13">13 May 1981</collectingDate>
</date>
,
<emphasis id="99ABEA83FF894475DDCEF96BFBFBF972" box="[644,1027,1670,1697]" italics="true" pageId="48" pageNumber="49">
<collectorName id="062A5347FF894475DDCEF96BFCF0F973" box="[644,776,1671,1697]" pageId="48" pageNumber="49">K.J. Sytsma</collectorName>
&amp;
<collectorName id="062A5347FF894475DC64F96BFC3BF972" box="[814,963,1670,1696]" pageId="48" pageNumber="49">L. Andersson</collectorName>
4529
</emphasis>
(
<collectionCode id="CDCEAE54FF894475DB5EF96AFBB2F972" box="[1044,1098,1670,1696]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="48" pageNumber="49" type="Herbarium">MO</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="1BB73CCCFF894475DB2EF96AFF2CF93A" collectingDate="1987-07-14" collectionCode="MO" collectorName="Thomas B. Croat" country="Panama" elevation="900" latitude="8.616667" location="Cerro Gaital" longLatPrecision="1301" longitude="-80.13333" municipality="Vicinity of La Mesa" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Cocle">
<collectingMunicipality id="4B04ACEBFF894475DB2EF96AFAB2F973" box="[1124,1354,1670,1697]" pageId="48" pageNumber="49">Vicinity of La Mesa</collectingMunicipality>
,
<locationDeviation id="BAD22FB3FF894475DA1DF96BFEC8F916" location="El Valle de Anton" pageId="48" pageNumber="49">N of El Valle de Antón</locationDeviation>
, along N slope of
<location id="AE00604AFF894475DEB3F946FD7EF916" LSID="urn:lsid:plazi:treatment:23768787FF894475DFC2FF72FF20F93A:AE00604AFF894475DEB3F946FD7EF916" box="[505,646,1706,1732]" country="Panama" latitude="8.616667" longLatPrecision="1301" longitude="-80.13333" municipality="Vicinity of La Mesa" name="Cerro Gaital" pageId="48" pageNumber="49" stateProvince="Cocle">Cerro Gaital</location>
,
<geoCoordinate id="CEEB5056FF894475DDDBF946FD0AF916" box="[657,754,1706,1732]" degrees="08" direction="north" minutes="37" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="8.616667">08°37’N</geoCoordinate>
<geoCoordinate id="CEEB5056FF894475DDB3F946FC95F916" box="[761,877,1706,1732]" degrees="080" direction="west" minutes="08" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-80.13333">080°08’W</geoCoordinate>
,
<quantity id="6C279B74FF894475DC32F946FC01F917" box="[888,1017,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="48" pageNumber="49" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">
<elevation id="20F2D1A2FF894475DC32F946FC01F917" box="[888,1017,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="48" pageNumber="49" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">850–950 m</elevation>
</quantity>
,
<date id="DF611051FF894475DB4EF946FB6FF916" box="[1028,1175,1706,1732]" pageId="48" pageNumber="49" value="1987-07-14">
<collectingDate id="CF25E9B9FF894475DB4EF946FB6FF916" box="[1028,1175,1706,1732]" pageId="48" pageNumber="49" value="1987-07-14">14 July 1987</collectingDate>
</date>
,
<emphasis id="99ABEA83FF894475DBEBF946FA53F916" box="[1185,1451,1706,1733]" italics="true" pageId="48" pageNumber="49">
<collectorName id="062A5347FF894475DBEBF946FAA7F917" box="[1185,1375,1706,1733]" pageId="48" pageNumber="49">Thomas B. Croat</collectorName>
67294
</emphasis>
(
<collectionCode id="CDCEAE54FF894475DFDBF922FF30F93A" box="[145,200,1742,1768]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="48" pageNumber="49" type="Herbarium">MO</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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