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<document ID-DOI="10.11646/zootaxa.5244.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7663518" ID-ZooBank="82968DC8-49F4-4E1E-9DFA-EE71531EE121" approvalRequired="23" approvalRequired_for_taxonomicNames="13" approvalRequired_for_textStreams="4" approvalRequired_for_treatments="6" checkinTime="1677050933648" checkinUser="plazi" docAuthor="Danielopol, Dan L., Cabral, M. Cristina, Horne, David J., Namiotko, Tadeusz &amp; Lord, Alan R." docDate="2023" docId="03DA682B2E14FFB1A398FA7AFB513DA2" docLanguage="en" docName="zootaxa.5244.4.1.pdf" docOrigin="Zootaxa 5244 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microceratina andreui Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="327" masterDocId="FFE310532E0CFFABA30FFFC1FFCC3A7F" masterDocTitle="Reconciling diagnostic traits in living and fossil taxa: The taxonomy and evolution of the genus Microceratina (Crustacea, Ostracoda, Cytheruridae)" masterLastPageNumber="340" masterPageNumber="301" pageNumber="325" updateTime="1677051428792" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Reconciling diagnostic traits in living and fossil taxa: The taxonomy and evolution of the genus Microceratina (Crustacea, Ostracoda, Cytheruridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Danielopol, Dan L.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3968-5564</mods:nameIdentifier>
<mods:affiliation>Karl-Franzens University, Institute of Earth Sciences, Department of Geology and Palaeontology, Heinrichstrasse 26, A- 8010 Graz, Austria. dldanielopol @ icloud. com; https: // orcid. org / 0000 - 0003 - 3968 - 5564</mods:affiliation>
<mods:nameIdentifier type="email">dldanielopol@icloud.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cabral, M. Cristina</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8717-4043</mods:nameIdentifier>
<mods:affiliation>Instituto Dom Luiz (IDL), Faculdade de Ciências, Universidade de Lisboa, Campo Grande, C 1, 1749 - 016 Lisboa, Portugal &amp; Departamento de Geologia, Faculdade de Ciências, Universidade de Lisboa, Campo Grande, C 6, 1749 - 016 Lisboa, Portugal mccabral @ fc. ul. pt; https: // orcid. org / 0000 - 0001 - 8717 - 4043</mods:affiliation>
<mods:nameIdentifier type="email">mccabral@fc.ul.pt</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Horne, David J.</mods:namePart>
<mods:affiliation>School of Geography, Queen Mary University of London, Mile End Road, London E 1 4 NS, UK.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Namiotko, Tadeusz</mods:namePart>
<mods:affiliation>University of Gdansk, Faculty of Biology, Department of Evolutionary Genetics and Biosystematics, Laboratory of Biosystematics and Ecology of Aquatic Invertebrates, Wita Stwosza 59, 80 - 308 Gdańsk, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lord, Alan R.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0008-7746</mods:nameIdentifier>
<mods:affiliation>Senckenberg Forschungsinstitut Frankfurt, Senckenberganlage 25, D- 60325 Frankfurt-am-Main, Germany alan. lord @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 0008 - 7746</mods:affiliation>
<mods:nameIdentifier type="email">alan.lord@senckenberg.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5244</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>301</mods:start>
<mods:end>340</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.4.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5244.4.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7663518</mods:identifier>
<mods:identifier type="ZooBank">82968DC8-49F4-4E1E-9DFA-EE71531EE121</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DA682B2E14FFB1A398FA7AFB513DA2" httpUri="http://treatment.plazi.org/id/03DA682B2E14FFB1A398FA7AFB513DA2" lastPageId="26" lastPageNumber="327" pageId="24" pageNumber="325">
<subSubSection box="[151,725,1467,1494]" pageId="24" pageNumber="325" type="nomenclature">
<paragraph blockId="24.[151,725,1467,1529]" box="[151,725,1467,1494]" pageId="24" pageNumber="325">
<heading bold="true" box="[151,725,1467,1494]" fontSize="11" level="1" pageId="24" pageNumber="325" reason="1">
<emphasis bold="true" box="[151,725,1467,1494]" pageId="24" pageNumber="325">
<taxonomicName authority="Cabral &amp; Lord" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[151,624,1467,1494]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis bold="true" box="[151,426,1467,1494]" italics="true" pageId="24" pageNumber="325">Microceratina andreui</emphasis>
Cabral &amp; Lord
</taxonomicName>
<taxonomicNameLabel box="[631,725,1467,1493]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,259,1503,1529]" pageId="24" pageNumber="325" type="description">
<paragraph blockId="24.[151,725,1467,1529]" box="[151,259,1503,1529]" pageId="24" pageNumber="325">
<figureCitation box="[151,259,1503,1529]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="24" pageNumber="325">Figure 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="325" type="etymology">
<paragraph blockId="24.[151,1437,1575,1961]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,290,1575,1601]" pageId="24" pageNumber="325">Etymology:</emphasis>
named after Bernard Andreu (University Paul Sabatier, Toulouse,
<collectingCountry box="[1089,1167,1575,1601]" name="France" pageId="24" pageNumber="325">France</collectingCountry>
) who first noticed the similarity with
<emphasis box="[322,491,1612,1637]" italics="true" pageId="24" pageNumber="325">
<taxonomicName box="[322,486,1612,1637]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">Microceratina</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="325" type="materials_examined">
<paragraph blockId="24.[151,1437,1575,1961]" box="[151,1337,1647,1674]" pageId="24" pageNumber="325">
<materialsCitation box="[151,1337,1647,1674]" collectionCode="SMF" pageId="24" pageNumber="325" specimenCode="BM52" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[151,269,1647,1673]" pageId="24" pageNumber="325">
<typeStatus box="[151,264,1647,1673]" pageId="24" pageNumber="325">Holotype</typeStatus>
:
</emphasis>
1C,
<collectionCode box="[323,379,1647,1673]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="325">SMF</collectionCode>
Xe 23990, L =
<quantity box="[556,658,1647,1674]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="24" pageNumber="325" unit="mm" value="0.39">0.39 mm</quantity>
, H =
<quantity box="[719,821,1647,1674]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="325" unit="mm" value="0.19">0.19 mm</quantity>
, W =
<quantity box="[888,990,1647,1674]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="325" unit="mm" value="0.19">0.19 mm</quantity>
, sample
<specimenCode box="[1088,1160,1647,1674]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="24" pageNumber="325">BM52</specimenCode>
B (
<figureCitation box="[1193,1329,1647,1673]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="24" pageNumber="325">Figs 14A–F</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph blockId="24.[151,1437,1575,1961]" pageId="24" pageNumber="325">
<materialsCitation box="[151,1100,1683,1710]" collectionCode="SMF" pageId="24" pageNumber="325" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[151,280,1683,1709]" pageId="24" pageNumber="325">
<typeStatus box="[151,275,1683,1709]" pageId="24" pageNumber="325">Paratypes</typeStatus>
:
</emphasis>
2C,
<collectionCode box="[335,391,1683,1709]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="325">SMF</collectionCode>
Xe 23991, L =
<quantity box="[570,673,1683,1710]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="24" pageNumber="325" unit="mm" value="0.39">0.39 mm</quantity>
, H =
<quantity box="[736,838,1683,1710]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="325" unit="mm" value="0.19">0.19 mm</quantity>
, W =
<quantity box="[907,1009,1683,1710]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="mm" value="0.2">0.20 mm</quantity>
, sample
</materialsCitation>
<materialsCitation box="[1108,1328,1683,1710]" collectionCode="BM" pageId="24" pageNumber="325" specimenCode="BM52" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1108,1181,1683,1710]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="24" pageNumber="325">BM52</specimenCode>
B (
<figureCitation box="[1214,1319,1683,1709]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="24" pageNumber="325">Fig. 14Q</figureCitation>
)
</materialsCitation>
;
<collectionCode box="[1340,1396,1683,1709]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="325">SMF</collectionCode>
Xe 23992, L =
<quantity box="[281,383,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="24" pageNumber="325" unit="mm" value="0.38">0.38 mm</quantity>
, H =
<quantity box="[445,545,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="24" pageNumber="325" unit="mm" value="0.18">0.18 mm</quantity>
W =
<quantity box="[606,708,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="325" unit="mm" value="0.19">0.19 mm</quantity>
, sample
<materialsCitation box="[806,1035,1719,1746]" collectionCode="BM" pageId="24" pageNumber="325" specimenCode="BM57" specimenCount="1" typeStatus="paratype">
<specimenCode box="[806,877,1719,1746]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="24" pageNumber="325">BM57</specimenCode>
(
<figureCitation box="[893,1026,1719,1745]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="24" pageNumber="325">Figs 14P, R</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="24.[151,1437,1575,1961]" box="[151,1087,1755,1782]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,265,1755,1781]" pageId="24" pageNumber="325">Material:</emphasis>
8C, 3 Toarcian, 5 Aalenian (3 C figured:
<collectionCode box="[730,786,1755,1781]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="325">SMF</collectionCode>
Xe 23993, 23994, 23995).
</paragraph>
<paragraph blockId="24.[151,1437,1575,1961]" box="[151,1242,1791,1818]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,313,1791,1817]" pageId="24" pageNumber="325">Type locality:</emphasis>
Boca da Mata, Alvaiázere,
<emphasis box="[623,635,1792,1817]" italics="true" pageId="24" pageNumber="325">c</emphasis>
.
<quantity box="[650,735,1791,1817]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="325" unit="km" value="170.0">170 km</quantity>
NE of
<collectingRegion box="[816,897,1791,1817]" country="Portugal" name="Lisboa" pageId="24" pageNumber="325">Lisbon</collectingRegion>
,
<collectingCountry box="[908,1007,1791,1817]" name="Portugal" pageId="24" pageNumber="325">Portugal</collectingCountry>
;
<geoCoordinate box="[1018,1127,1791,1818]" degrees="39" direction="north" minutes="48" orientation="latitude" pageId="24" pageNumber="325" precision="925" value="39.8">39º 48’ N</geoCoordinate>
;
<geoCoordinate box="[1139,1238,1791,1818]" degrees="8" direction="west" minutes="24" orientation="longitude" pageId="24" pageNumber="325" precision="925" value="-8.4">8º 24’ W</geoCoordinate>
.
</paragraph>
<paragraph blockId="24.[151,1437,1575,1961]" box="[151,1085,1827,1854]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,280,1827,1853]" pageId="24" pageNumber="325">Type level:</emphasis>
S. Gi„o Formation; Upper Toarcian; Speciosum Zone; sample
<collectionCode box="[987,1033,1827,1854]" country="United Kingdom" name="Bristol Museum" pageId="24" pageNumber="325">BM</collectionCode>
52B.
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="325" type="diagnosis">
<paragraph blockId="24.[151,1437,1575,1961]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,276,1863,1889]" pageId="24" pageNumber="325">Diagnosis:</emphasis>
A species of
<taxonomicName box="[429,590,1864,1889]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis box="[429,590,1864,1889]" italics="true" pageId="24" pageNumber="325">Microceratina</emphasis>
</taxonomicName>
with an inflated and elongated carapace; shallow dorso-median sulcus; reticulate ornamentation with polygonal cells, whose sola are covered by round fossae; two short and thin, slightly curved antero-ventral ridges.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7663552" ID-Zenodo-Dep="7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="25" pageNumber="326" startId="25.[151,250,1498,1522]" targetBox="[270,1317,181,1474]" targetPageId="25">
<paragraph blockId="25.[151,1437,1498,2026]" pageId="25" pageNumber="326">
<emphasis bold="true" box="[151,290,1498,1522]" pageId="25" pageNumber="326">FIGURE 14.</emphasis>
<taxonomicName authority="Cabral &amp; Lord" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[299,694,1498,1522]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[299,532,1498,1522]" italics="true" pageId="25" pageNumber="326">Microceratina andreui</emphasis>
Cabral &amp; Lord
</taxonomicName>
<emphasis bold="true" box="[704,787,1498,1522]" pageId="25" pageNumber="326">
<taxonomicNameLabel box="[704,787,1498,1522]" pageId="25" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus).
</paragraph>
</caption>
<subSubSection pageId="26" pageNumber="327" type="description">
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">
<emphasis bold="true" box="[151,299,150,176]" pageId="26" pageNumber="327">Description:</emphasis>
Carapace small, subrectangular in lateral outline, with valves of almost the same size. L &lt;
<quantity box="[1341,1432,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="327" unit="mm" value="0.5">0.5 mm</quantity>
, H/L% &lt;50% located in the anterior third of L (
<typeStatus box="[678,787,187,213]" pageId="26" pageNumber="327">Holotype</typeStatus>
,
<figureCitation box="[798,902,187,213]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14A</figureCitation>
). Greatest L above mid-H. Greatest W approximately at three-fifths of L from anterior, less than half of length. AMZ well-developed on both valves (
<figureCitation captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14A, B, D, T, U</figureCitation>
); it represents 2.5% of valve L (in
<figureCitation box="[626,728,259,285]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14A</figureCitation>
, with L =
<quantity box="[842,946,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="327" unit="mm" value="0.39">0.39 mm</quantity>
).
</paragraph>
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">
Lateral surface with a weak dorso-median sulcus, ending in an enlarged subcentral depression where the adductor muscle scars can be seen externally (
<figureCitation box="[710,818,331,357]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14C</figureCitation>
). Mid and postero-ventral zones inflated, forming a longitudinal expansion with a curved outline, which ends in a round protuberance (
<figureCitation box="[1097,1273,367,393]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14A, E, R</figureCitation>
). Eye tubercle present, represented by a subtle elongated swelling with an associated shallow postocular sulcus. In dorsal view (
<figureCitation box="[159,263,439,465]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14Q</figureCitation>
) the carapace is inflated, mostly posteriorly, with anterior and posterior ends pointed and dorso-median sulcus marked; in ventral view (
<figureCitation box="[518,617,475,501]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14P</figureCitation>
) it has a sagittate outline, i.e., anterior pointed and the posterior margin extends beyond the two ventro-lateral expansions resembling an arrowhead. Sexual dimorphism not evident.
</paragraph>
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">Dorsal margin straight, with obtuse cardinal angles. Ventral margin weakly sinuous, tapering towards the posterior, partially covered by the postero-ventral longitudinal expansion. Anterior margin rounded, infracurvate. Posterior margin rounded to slightly pointed, with extremity supracurvate, without forming a caudal process. Both anterior and posterior margins flattened.</paragraph>
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">
Reticulate ornamentation, with mostly pentagonal cells; in the flattened anterior and posterior zones the reticulation has a square mesh (
<figureCitation box="[502,702,727,753]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14A, B, E, F</figureCitation>
). The sola of the reticula are covered with fossae (
<emphasis box="[1263,1275,727,752]" italics="true" pageId="26" pageNumber="327">c</emphasis>
. 17 fossae per mesh), generally round and shallow. The median diameter for several fossae visible in
<figureCitation box="[1131,1307,763,789]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14L, M, O</figureCitation>
is 2.95 µm (r = 2.65–3.4 µm, N =10). Minute elongated grooves are visible on the floors of some fossae (
<figureCitation box="[1210,1314,799,825]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14N</figureCitation>
). In one of these latter fossae with a diameter of 2.74 µm, the size of one of the elongated grooves is 0.7 x 0.1 µm. We presume that it represents the preserved remains of a foveola (= LStPC) similar to those illustrated for
<taxonomicName baseAuthorityName="Barbeito-Gonzalez" baseAuthorityYear="1971" box="[1219,1436,871,897]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="pseudoamfibola">
<emphasis box="[1219,1436,871,897]" italics="true" pageId="26" pageNumber="327">M. pseudoamfibola</emphasis>
</taxonomicName>
(e.g.,
<figureCitation box="[216,337,907,933]" captionStart="FIGURE 8" captionStartId="18.[151,250,1718,1742]" captionTargetBox="[192,1396,309,1694]" captionTargetId="figure-96@18.[192,1396,309,1694]" captionTargetPageId="18" captionText="FIGURE 8. Microceratina pseudoamfibola (Barbeito-Gonzàlez, 1971), valve morphology: A–G—RV-1; H–K—LV-1; L–N— RV-2 (Calvi, Corsica, France); A–G—RV, external; A—general view; B—detail of A, posterior half of valve (light gray arrow indicates angular ventro-posterior ridge); C—detail of B, intrareticular solum with fossae and LStPC; D—detail of A, anterior half of valve; E, F—detail of fossae with LStPC (white arrows indicate aperture of tubuli with a loophole pore); G—detail of A (rotated 90º), intramural fossae within a reticular structure; H–K—LV; H—external, detail of the ventral side (white arrow points to anterior); I–K—internal; I–J—details of pores of LStPC; K—central area with adductor muscle scars (indicated by central white arrow) and apertures of two NPC (lateral white arrows); L–N—RV, internal; L—general view; M, N—detail of L, anterior (M) and posterior (N) hinge (black arrows indicate the positive terminal elements). (M. Peeters sample deposited at Royal Belgian Institute of Natural Sciences, Brussels (RBINS), Inv. Nr. 12703/0C2075)." figureDoi="http://doi.org/10.5281/zenodo.7663540" httpUri="https://zenodo.org/record/7663540/files/figure.png" pageId="26" pageNumber="327">Figs 8E, F</figureCitation>
). In the anterior zone, two short, thin and slightly curved ridges are present, the first one oblique, ending near the flattened anterior margin, the second one slightly longer, vertical, ending in the ventral margin (
<figureCitation captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Fig. 14B</figureCitation>
); in the Aalenian forms, these ridges are less clear.
</paragraph>
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">
<materialsCitation collectionCode="NPC" pageId="26" pageNumber="327" specimenCount="1">
Normal pore canals (conuli
<typeStatus box="[510,558,1015,1041]" pageId="26" pageNumber="327">type</typeStatus>
A”) can be seen adjacent or on the tectum of muri. A
<collectionCode box="[1155,1209,1015,1041]" country="India" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="26" pageNumber="327">NPC</collectionCode>
placed on a conulus adjacent to a murus (
<figureCitation box="[385,514,1051,1077]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14G–I</figureCitation>
) has a diameter of 1 µm. Other NPCs are visible on the muri (
<figureCitation box="[1211,1412,1051,1077]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14J, K, U, V</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="327">
Internal features not seen; four slightly elongated adductor muscle scars, arranged in a vertical row, can be seen externally in the subcentral depression (
<figureCitation box="[597,736,1123,1149]" captionStart="FIGURE 14" captionStartId="25.[151,250,1498,1522]" captionTargetBox="[270,1317,181,1474]" captionTargetId="figure-19@25.[270,1317,181,1474]" captionTargetPageId="25" captionText="FIGURE 14. Microceratina andreui Cabral &amp; Lord sp. nov.: A–F—C, holotype, SMF Xe 23990; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; A—left view; B—detail of A, AMZ (white arrow) and ornamentation (antero-ventral ridges); C—detail of B, adductor muscle scars; D—detail of the AMZ from B; E—detail of A, posterior, round postero-ventral protuberance; F—detail of E, posterior extremity and reticulation; G–O—C, SMF Xe 23993; L = 0.41 mm, H = 0.20 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM161; G—right view; H—detail of G, anterior, NPC (white arrow indicates pore conulus, A”-type); I—detail of H, pore conulus; J— detail of G, reticulate cells with sola covered by fossae and intramural NPC, A”-type (white arrow); K—detail of J, pentagonal cells with fossae and intramural NPC (white arrow); L—detail of G, several fossae on sola; M—detail of L, fossae; N—detail of G, fossae; O—detail of G, fossae; P—C, ventral view, paratype, SMF Xe 23992; L = 0.38 mm, H = 0.18 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM57; Q—C, dorsal view, paratype, SMF Xe 23991; L = 0.39 mm, H = 0.19 mm; Boca da Mata section, Portugal; Upper Toarcian, Speciosum Zone; sample BM52B; R—C, right view, paratype, same as P; S—C, left view, SMF Xe 23994; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM169B; T–V—C, SMF Xe 23995; L = 0.40 mm, H = 0.19 mm; Boca da Mata section, Portugal; Lower Aalenian, Opalinum Zone; sample BM162T; T—right view (white arrow indicates the AMZ); U—detail of T; V—detail of U, intramural NPC, A”-type (pore conulus)." figureDoi="http://doi.org/10.5281/zenodo.7663552" httpUri="https://zenodo.org/record/7663552/files/figure.png" pageId="26" pageNumber="327">Figs 14B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="327" type="discussion">
<paragraph blockId="26.[151,1437,1195,1905]" pageId="26" pageNumber="327">
<emphasis bold="true" box="[151,269,1195,1221]" pageId="26" pageNumber="327">Remarks:</emphasis>
<taxonomicName authority="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord, 2023" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[277,531,1195,1221]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[277,531,1195,1221]" italics="true" pageId="26" pageNumber="327">Microceratina andreui</emphasis>
</taxonomicName>
<emphasis bold="true" box="[538,628,1195,1221]" pageId="26" pageNumber="327">
<taxonomicNameLabel box="[538,628,1195,1221]" pageId="26" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the oldest known species of a long-lasting evolutionary lineage ranging from the Early Jurassic to living species. However, we are at present unable to recognise other close relatives of Early Jurassic age and it may be that ancestral forms are of Triassic age.
</paragraph>
<paragraph blockId="26.[151,1437,1195,1905]" pageId="26" pageNumber="327">
<taxonomicName authority="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord, 2023" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[199,456,1303,1329]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[199,456,1303,1329]" italics="true" pageId="26" pageNumber="327">Microceratina andreui</emphasis>
</taxonomicName>
<emphasis bold="true" box="[466,559,1303,1329]" pageId="26" pageNumber="327">
<taxonomicNameLabel box="[466,559,1303,1329]" pageId="26" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all the other known fossil and living species of
<taxonomicName box="[1276,1437,1303,1328]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis box="[1276,1437,1303,1328]" italics="true" pageId="26" pageNumber="327">Microceratina</emphasis>
</taxonomicName>
by the presence of the two short antero-ventral ridges in surface ornamentation. Moreover, the presence of the ocular structures (tubercle and postocular sulcus), uncommon in the genus, also allows the easy distinction of
<taxonomicName authority="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord, 2023" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[185,312,1411,1437]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[185,312,1411,1437]" italics="true" pageId="26" pageNumber="327">M. andreui</emphasis>
</taxonomicName>
<emphasis bold="true" box="[322,415,1411,1437]" pageId="26" pageNumber="327">
<taxonomicNameLabel box="[322,415,1411,1437]" pageId="26" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from most other species of its genus. However, in some fossil species these structures, though very weak, also appear. In
<taxonomicName authority="Colin, Dejax &amp; Geze, 2005" authorityName="Colin, Dejax &amp; Geze" authorityYear="2005" box="[551,1066,1446,1473]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="bhannesensis">
<emphasis box="[551,739,1447,1473]" italics="true" pageId="26" pageNumber="327">M. bhannesensis</emphasis>
<bibRefCitation author="Colin, J-P. &amp; Andreu, B. &amp; Dejax, J. &amp; Geze, R." box="[749,1066,1446,1473]" pageId="26" pageNumber="327" pagination="15 - 24" refId="ref27880" refString="Colin, J-P., Andreu, B., Dejax, J. &amp; Geze, R. (2005) Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa presence tethysienne des le Jurassique superieur. Revue de micropaleontologie, 48, 15 - 24. https: // doi. org / 10.1016 / j. revmic / 2004.12.0058" type="journal article" year="2005">Colin, Dejax &amp; Gèze, 2005</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation author="Colin, J-P. &amp; Andreu, B. &amp; Dejax, J. &amp; Geze, R." box="[1116,1321,1446,1473]" pageId="26" pageNumber="327" pagination="15 - 24" refId="ref27880" refString="Colin, J-P., Andreu, B., Dejax, J. &amp; Geze, R. (2005) Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa presence tethysienne des le Jurassique superieur. Revue de micropaleontologie, 48, 15 - 24. https: // doi. org / 10.1016 / j. revmic / 2004.12.0058" type="journal article" year="2005">
Colin
<emphasis box="[1188,1240,1447,1473]" italics="true" pageId="26" pageNumber="327">et al</emphasis>
., 2005
</bibRefCitation>
) from the Kimmeridgian (Late Jurassic) of
<collectingCountry box="[533,634,1483,1509]" name="Lebanon" pageId="26" pageNumber="327">Lebanon</collectingCountry>
, a very shallow postocular sulcus seems to exist (
<figureCitation box="[1210,1281,1483,1509]" captionStart="FIGURE 9" captionStartId="19.[151,250,957,981]" captionTargetBox="[569,1018,181,931]" captionTargetId="figure-19@19.[569,1018,181,933]" captionTargetPageId="19" captionText="FIGURE 9. General shape of Microceratina morphotypes:A—Microceratina bhannesensis Colin, Dejax &amp; Gèze, 2005, C right view (holotype); B, C—Microceratina azazoulensis Andreu &amp; Colin, 2005, silhouettes of (B) C, right side and (C) RV, lateral view, standardised for length (bounding box method); B—Tarhzoute, Morocco, Colin et al. (2005, pl. 2, fig. 4); C—Sergipe- Alagoas Basin, Brazil, valve nr. ULVG-12791 also figured in Vázquez-García et al. (2021, fig. 4L)." figureDoi="http://doi.org/10.5281/zenodo.7663542" httpUri="https://zenodo.org/record/7663542/files/figure.png" pageId="26" pageNumber="327">Fig. 9</figureCitation>
); this species is close to
<taxonomicName authority="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord, 2023" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[289,419,1519,1545]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[289,419,1519,1545]" italics="true" pageId="26" pageNumber="327">M. andreui</emphasis>
</taxonomicName>
<emphasis bold="true" box="[432,528,1519,1545]" pageId="26" pageNumber="327">
<taxonomicNameLabel box="[432,528,1519,1545]" pageId="26" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the posterior extremity is located more ventrally, and the anterior and posterior margins denticulate. In
<taxonomicName authority="Andreu &amp; Colin, 2005" authorityName="Andreu &amp; Colin" authorityYear="2005" box="[535,989,1554,1581]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="azazoulensis">
<emphasis box="[535,718,1555,1581]" italics="true" pageId="26" pageNumber="327">M. azazoulensis</emphasis>
Andreu &amp;
<bibRefCitation author="Colin, J-P. &amp; Andreu, B. &amp; Dejax, J. &amp; Geze, R." box="[853,989,1554,1581]" pageId="26" pageNumber="327" pagination="15 - 24" refId="ref27880" refString="Colin, J-P., Andreu, B., Dejax, J. &amp; Geze, R. (2005) Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa presence tethysienne des le Jurassique superieur. Revue de micropaleontologie, 48, 15 - 24. https: // doi. org / 10.1016 / j. revmic / 2004.12.0058" type="journal article" year="2005">Colin, 2005</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation author="Colin, J-P. &amp; Andreu, B. &amp; Dejax, J. &amp; Geze, R." box="[1041,1251,1554,1581]" pageId="26" pageNumber="327" pagination="15 - 24" refId="ref27880" refString="Colin, J-P., Andreu, B., Dejax, J. &amp; Geze, R. (2005) Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa presence tethysienne des le Jurassique superieur. Revue de micropaleontologie, 48, 15 - 24. https: // doi. org / 10.1016 / j. revmic / 2004.12.0058" type="journal article" year="2005">
Colin
<emphasis box="[1114,1167,1555,1581]" italics="true" pageId="26" pageNumber="327">et al</emphasis>
., 2005
</bibRefCitation>
), from the midlate Cenomanian (Upper Cretaceous) of
<collectingCountry box="[624,728,1591,1617]" name="Morocco" pageId="26" pageNumber="327">Morocco</collectingCountry>
, a weak postocular sulcus can be seen, although the authors say that there is no ocular tubercle (
<bibRefCitation author="Colin, J-P. &amp; Andreu, B. &amp; Dejax, J. &amp; Geze, R." box="[575,779,1626,1653]" pageId="26" pageNumber="327" pagination="15 - 24" refId="ref27880" refString="Colin, J-P., Andreu, B., Dejax, J. &amp; Geze, R. (2005) Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa presence tethysienne des le Jurassique superieur. Revue de micropaleontologie, 48, 15 - 24. https: // doi. org / 10.1016 / j. revmic / 2004.12.0058" type="journal article" year="2005">
Colin
<emphasis box="[650,710,1627,1653]" italics="true" pageId="26" pageNumber="327">et al.</emphasis>
2005
</bibRefCitation>
, p. 21—“absence de tubercule oculaire”); this species is much more quadrate than
<taxonomicName authority="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord, 2023" authorityName="Danielopol &amp; Cabral &amp; Horne &amp; Namiotko &amp; Lord" authorityYear="2023" box="[450,575,1663,1689]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="andreui" status="sp. nov.">
<emphasis box="[450,575,1663,1689]" italics="true" pageId="26" pageNumber="327">M. andreui</emphasis>
</taxonomicName>
<emphasis bold="true" box="[584,676,1663,1689]" pageId="26" pageNumber="327">
<taxonomicNameLabel box="[584,676,1663,1689]" pageId="26" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with a round posterior margin and a less developed ventro-lateral expansion; the late Albian-early Cenomanian Brazilian specimens of
<taxonomicName authorityName="Andreu &amp; Colin" authorityYear="2005" box="[922,1100,1699,1725]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="azazoulensis">
<emphasis box="[922,1100,1699,1725]" italics="true" pageId="26" pageNumber="327">M. azazoulensis</emphasis>
</taxonomicName>
, figured in
<bibRefCitation author="Vazquez-Garcia, B. &amp; Ceolin, D. &amp; Fauth, G. &amp; Borghi, L. &amp; Valle, B. &amp; Netto, A. M. R." pageId="26" pageNumber="327" pagination="1 - 24" refId="ref30395" refString="Vazquez-Garcia, B., Ceolin, D., Fauth, G., Borghi, L., Valle, B. &amp; Netto, A. M. R. (2021) Ostracods from the late Albian - early Cenomanian of the Sergipe - Alagoas Basin, Brazil: New taxonomic and biostratigraphic inferences. Journal of South American Earth Sciences, 108, 1 - 24. https: // doi. org / 10.1016 / j. jsames. 2021.103169" type="journal article" year="2021">
Vázquez-García
<emphasis italics="true" pageId="26" pageNumber="327">et al</emphasis>
. (2021)
</bibRefCitation>
, almost certainly belong to another species (see Discussion below); they are longer than the Portuguese species, with a less marked dorso-median sulcus and ventro-lateral expansion, and without ocular structures.
</paragraph>
<paragraph blockId="26.[151,1437,1195,1905]" pageId="26" pageNumber="327">
<taxonomicName authority="Herrig, 1991" authorityName="Herrig" authorityYear="1991" box="[199,634,1807,1833]" class="Ostracoda" family="Cytheruridae" genus="Microceratina" kingdom="Animalia" order="Podocopida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="aequabilis">
<emphasis box="[199,483,1807,1833]" italics="true" pageId="26" pageNumber="327">Microceratina aequabilis</emphasis>
<materialsCitation box="[489,634,1807,1833]" country="Germany" county="Portuguese" location="Maastrichtian" municipality="Upper Cretaceous" pageId="26" pageNumber="327" specimenCount="1" typeStatus="holotype">
<bibRefCitation author="Herrig, E." box="[489,634,1807,1833]" pageId="26" pageNumber="327" pagination="99 - 115" refId="ref28369" refString="Herrig, E. (1991) Neue Ostrakoden aus verkieselten kalksteinen der hoeheren Oberkreide (Ober-Maastricht) der Danisch- Polnischen Furche / Ostsee, 2. Geologisches Jahrbuch, A, 128, 99 - 115." type="journal article" year="1991">Herrig, 1991</bibRefCitation>
</materialsCitation>
</taxonomicName>
, from the
<location LSID="urn:lsid:plazi:treatment:03DA682B2E14FFB1A398FA7AFB513DA2:8EAC8FE62E16FFB1A1E5F8CEFC483D56" box="[746,900,1807,1833]" country="Germany" county="Portuguese" municipality="Upper Cretaceous" name="Maastrichtian" pageId="26" pageNumber="327">Maastrichtian</location>
(
<collectingMunicipality box="[914,1118,1807,1833]" pageId="26" pageNumber="327">Upper Cretaceous</collectingMunicipality>
) of
<collectingCountry box="[1162,1267,1807,1833]" name="Germany" pageId="26" pageNumber="327">Germany</collectingCountry>
, is very similar to the
<collectingCounty box="[225,349,1843,1869]" pageId="26" pageNumber="327">Portuguese</collectingCounty>
species, but is larger (L =
<quantity box="[662,767,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="26" pageNumber="327" unit="mm" value="0.44">0.44 mm</quantity>
, H =
<quantity box="[835,939,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="26" pageNumber="327" unit="mm" value="0.21">0.21 mm</quantity>
,
<typeStatus box="[953,1062,1843,1869]" pageId="26" pageNumber="327">Holotype</typeStatus>
), with a less projecting posterior margin.
</paragraph>
<paragraph blockId="26.[151,1436,1950,2013]" pageId="26" pageNumber="327">
<emphasis bold="true" box="[151,664,1950,1977]" pageId="26" pageNumber="327">Occurrence and stratigraphic distribution:</emphasis>
A rare species found in the Upper Toarcian, Lower Jurassic (S. Gi„o Formation) and Lower Aalenian, Middle Jurassic (Póvoa da Lomba Formation) of
<collectingCountry box="[1078,1177,1987,2013]" name="Portugal" pageId="26" pageNumber="327">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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